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<document id="F2724639296D52EA9DFBFF158AB6CF8C" ID-CLB-Dataset="7806" ID-DOI="10.5281/zenodo.4503596" ID-GBIF-Dataset="9a3c7406-d4a1-4480-af4b-413daed3c500" ID-ISSN="0374-1036" ID-RefBankID="FF8F4870FFB95708A713FFC9743EFFF5" ID-Zenodo-Dep="4503596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612461472076" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2008" docId="E76D87CEC936FFA26797F3ECFC001310" docLanguage="en" docName="ActaEntMusNatPra.48.1.51-65.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus madianus Bernhauer 1936" docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="1B54FFB6C93EFFA8663FF129FF501064" masterDocTitle="Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="65" masterPageNumber="51" pageNumber="59" updateTime="1698927111911" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="112617D56FA65A6AE2C3308145BA4862" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="11356C5D220C81A52257B1A293075055">
<mods:title id="7830073CC48D88FD4A5FEBA6C75083C7">Revision of Afrotropical species of the Philonthus peripateticus species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
</mods:titleInfo>
<mods:name id="F60BEC5F35E637C0962F257A27422E30" type="personal">
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<mods:namePart id="E9AC8B8CFA46584606ED979D9BB574A6">Hromádka, Lubomír</mods:namePart>
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<mods:title id="F89B00C695F9B49DD175411B5994FDB0">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="0056D216F9ACA024C80167799A952543">
<mods:date id="B3141215BD5BF00DB7DC45AA0F5DC83C">2008</mods:date>
<mods:detail id="4361691E86A4B24DBE2308F8ECD42FAF" type="pubDate">
<mods:number id="69D9472C2E53AACA3E712FC0E9D257C9">2008-08-15</mods:number>
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<mods:number id="DCA0DA040C140DEA934159BF39AF8BE0">48</mods:number>
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<mods:detail id="86E5FD2C36A40F85FADF150C8AD2950F" type="issue">
<mods:number id="5F9AE604CBE05F7CB399D56F0FE84064">1</mods:number>
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<mods:identifier id="18E0075E7C477C85F4B635432E3B05AA" type="CLB-Dataset">7806</mods:identifier>
<mods:identifier id="1692C66455ACA401F9DF2B26946AEB21" type="DOI">10.5281/zenodo.4503596</mods:identifier>
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<treatment id="E76D87CEC936FFA26797F3ECFC001310" ID-DOI="http://doi.org/10.5281/zenodo.4503501" ID-GBIF-Taxon="180302219" ID-Zenodo-Dep="4503501" LSID="urn:lsid:plazi:treatment:E76D87CEC936FFA26797F3ECFC001310" httpUri="http://treatment.plazi.org/id/E76D87CEC936FFA26797F3ECFC001310" lastPageId="10" lastPageNumber="61" pageId="8" pageNumber="59">
<subSubSection id="27DE6553C936FFA06797F3ECFC0612BA" box="[424,854,709,734]" pageId="8" pageNumber="59" type="nomenclature">
<paragraph id="6F7B36D8C936FFA06797F3ECFC0612BA" blockId="8.[424,854,709,734]" box="[424,854,709,734]" pageId="8" pageNumber="59">
<heading id="343381B4C936FFA06797F3ECFC0612BA" bold="true" box="[424,854,709,734]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="59" reason="3">
<taxonomicName id="A8C44D5BC936FFA06797F3ECFC0612BA" ID-CoL="779WW" authority="Bernhauer, 1936" authorityName="Bernhauer" authorityYear="1936" box="[424,854,709,734]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="madianus">
<emphasis id="5DB0EACAC936FFA06797F3ECFC0612BA" bold="true" box="[424,854,709,734]" pageId="8" pageNumber="59">
<emphasis id="5DB0EACAC936FFA06797F3ECFDC412BA" bold="true" box="[424,660,709,734]" italics="true" pageId="8" pageNumber="59">Philonthus madianus</emphasis>
<bibRefCitation id="0B554B29C936FFA064A4F3EFFC0612BA" author="BERNHAUER M." box="[667,854,710,734]" pageId="8" pageNumber="59" pagination="261 - 267" refId="ref6059" refString="BERNHAUER M. 1936: Neue Staphylinidae aus Angola. Novitates Zoologicae 39: 261 - 267." type="journal article" year="1936">Bernhauer, 1936</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C936FFA06478F3CDFDE61293" box="[583,694,740,759]" pageId="8" pageNumber="59" type="description">
<paragraph id="6F7B36D8C936FFA06478F3CDFDE61293" blockId="8.[151,694,740,794]" box="[583,694,740,759]" pageId="8" pageNumber="59">
(
<figureCitation id="F7FF2A5DC936FFA06472F3CDFDE01293" box="[589,688,740,759]" captionStart="Figs" captionStartId="9.[113,152,1524,1543]" captionTargetBox="[121,1082,220,1478]" captionTargetId="figure-63@9.[118,1084,219,1479]" captionTargetPageId="9" captionText="Figs. 20-28. 20-24 – Philonthus lamtoensis Levasseur, 1967; 25-28 – P. madianus Bernhauer, 1936. 20-22, 25- 27 – aedeagus: 20, 25 ventral view, 21, 26 – dorsal view, 22, 27 – lateral view; 23, 28 – undersite of paramere, with sensory peg setae; 24 – apical portion of sternite VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4503610" httpUri="https://zenodo.org/record/4503610/files/figure.png" pageId="8" pageNumber="59">Figs. 25-28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C936FFA066A8F221FDD0137E" box="[151,640,775,794]" pageId="8" pageNumber="59" type="reference_group">
<paragraph id="6F7B36D8C936FFA066A8F221FDD0137E" blockId="8.[151,694,740,794]" box="[151,640,775,794]" pageId="8" pageNumber="59">
<treatmentCitationGroup id="4FD411F6C936FFA066A8F221FDD0137E" box="[151,640,775,794]" pageId="8" pageNumber="59">
<treatmentCitation id="EE6510C9C936FFA066A8F221FD2D137E" author="BERNHAUER M." box="[151,637,775,794]" page="263" pageId="8" pageNumber="59" year="1936">
<taxonomicName id="A8C44D5BC936FFA066A8F221FD2D137E" ID-CoL="779WW" authority="Bernhauer, 1936: 263" authorityName="Bernhauer" authorityPageNumber="263" authorityYear="1936" box="[151,637,775,794]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="madianus" subGenus="Philonthus">
<emphasis id="5DB0EACAC936FFA066A8F221FFA3137E" box="[151,243,776,794]" italics="true" pageId="8" pageNumber="59">Philonthus</emphasis>
(
<emphasis id="5DB0EACAC936FFA0673FF221FE0C137E" box="[256,348,776,794]" italics="true" pageId="8" pageNumber="59">Philonthus</emphasis>
)
<emphasis id="5DB0EACAC936FFA06756F221FEEC137E" box="[361,444,776,794]" italics="true" pageId="8" pageNumber="59">madianus</emphasis>
<bibRefCitation id="0B554B29C936FFA067FEF22EFD2D137E" author="BERNHAUER M." box="[449,637,775,794]" pageId="8" pageNumber="59" pagination="261 - 267" refId="ref6059" refString="BERNHAUER M. 1936: Neue Staphylinidae aus Angola. Novitates Zoologicae 39: 261 - 267." type="journal article" year="1936">Bernhauer, 1936: 263</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C936FFA066A8F218FC2013CA" pageId="8" pageNumber="59" type="materials_examined">
<paragraph id="6F7B36D8C936FFA066A8F218FEF41320" blockId="8.[151,1127,816,942]" box="[151,420,816,836]" pageId="8" pageNumber="59">
<emphasis id="5DB0EACAC936FFA066A8F218FE401320" bold="true" box="[151,272,816,836]" pageId="8" pageNumber="59">
<typeStatus id="B07F887AC936FFA066A8F218FF941320" box="[151,196,817,836]" pageId="8" pageNumber="59">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="17D37648C936FFA0672BF218FE051320" box="[276,341,817,836]" name="Angola" pageId="8" pageNumber="59">Angola</collectingCountry>
, Ebanga.
</paragraph>
<paragraph id="6F7B36D8C936FFA066A8F262FC2013CA" blockId="8.[151,1127,816,942]" pageId="8" pageNumber="59">
<emphasis id="5DB0EACAC936FFA066A8F262FE29133A" bold="true" box="[151,377,842,862]" pageId="8" pageNumber="59">Type material examined.</emphasis>
<materialsCitation id="DFAC3C85C936FFA06741F265FB58131D" ID-GBIF-Occurrence="3047799310" collectionCode="FMNH" collectorName="M. Bernhauer" pageId="8" pageNumber="59" specimenCount="1" typeStatus="holotype">
<typeStatus id="B07F887AC936FFA06741F265FE821339" box="[382,466,844,863]" pageId="8" pageNumber="59" type="holotype">HOLOTYPE</typeStatus>
: ♁, ‘
<collectorName id="C231530EC936FFA067C4F262FD10133A" box="[507,576,843,862]" pageId="8" pageNumber="59">Angola</collectorName>
,
<collectorName id="C231530EC936FFA06477F262FD85133A" box="[584,725,843,862]" pageId="8" pageNumber="59">Miss. Sc. Suisse</collectorName>
, 1932-1933 /
<collectorName id="C231530EC936FFA06574F262FCDB133A" box="[843,907,843,862]" pageId="8" pageNumber="59">Ebanga</collectorName>
nov., /
<collectorName id="C231530EC936FFA065F5F262FB41133A" box="[970,1041,843,862]" pageId="8" pageNumber="59">Chicago</collectorName>
NHMus.,
<collectorName id="C231530EC936FFA066A8F24FFE26131D" box="[151,374,870,889]" pageId="8" pageNumber="59">M. Bernhauer Collection.</collectorName>
//
<taxonomicName id="A8C44D5BC936FFA067B1F24FFD11131D" authority="Bernhauer" authorityName="Bernhauer" authorityYear="1936" box="[398,577,870,889]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="madianus">madianus Bernhauer</taxonomicName>
,
<collectorName id="C231530EC936FFA06474F24FFDD5131D" box="[587,645,870,889]" pageId="8" pageNumber="59">Typus.</collectorName>
[ochre oblong label, handwritten]’ (
<collectionCode id="09D5AE1DC936FFA06580F24FFB52131D" box="[959,1026,870,889]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="59" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<typeStatus id="B07F887AC936FFA0622EF24FFB30131C" box="[1041,1120,870,889]" pageId="8" pageNumber="59" type="paratype">PARATYPE</typeStatus>
:
<specimenCount id="79C2FD51C936FFA066A8F254FFF713F0" box="[151,167,893,916]" pageId="8" pageNumber="59" type="female">♀</specimenCount>
, ‘
<collectingCountry id="17D37648C936FFA0668AF2A8FE4713F0" box="[181,279,897,916]" name="Uganda" pageId="8" pageNumber="59">UGANDA</collectingCountry>
, Madi,
<date id="1B7A1018C936FFA06766F2A8FEC413F0" box="[345,404,897,916]" pageId="8" pageNumber="59" value="1927-05">v.1927</date>
, / G. D. H. Carpenter, / Brit. Mus. Don. Marshall, / Chicago NHMus. M. Bernhauer Collection, //
<taxonomicName id="A8C44D5BC936FFA06733F2B5FEA113CA" authority="Bernh. Cotypus." authorityName="Bernh. Cotypus." box="[268,497,923,942]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="madianus">
<emphasis id="5DB0EACAC936FFA06733F2B5FE0F13CA" box="[268,351,924,942]" italics="true" pageId="8" pageNumber="59">madianus</emphasis>
Bernh. Cotypus.
</taxonomicName>
[ochre oblong label, handwritten]’ (
<collectionCode id="09D5AE1DC936FFA0651CF2B2FC3613CA" box="[803,870,923,942]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="59" type="Museum">FMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C936FFA266A8F2EFFE4D12F7" lastPageId="10" lastPageNumber="61" pageId="8" pageNumber="59" type="description">
<paragraph id="6F7B36D8C936FFA066A8F2EFFEAC139A" blockId="8.[151,1127,966,1631]" pageId="8" pageNumber="59">
<emphasis id="5DB0EACAC936FFA066A8F2EFFE6D13BA" bold="true" box="[151,317,966,990]" pageId="8" pageNumber="59">Redescription.</emphasis>
Body length
<quantity id="A83C9B3DC936FFA067EDF2EEFD3813BA" box="[466,616,967,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.205" metricValueMax="1.22" metricValueMin="1.19" pageId="8" pageNumber="59" unit="mm" value="12.05" valueMax="12.2" valueMin="11.9">11.9-12.2 mm</quantity>
(with stretched abdomen), length of fore body (to the end of elytra)
<quantity id="A83C9B3DC936FFA06746F2CFFEA8139A" box="[377,504,998,1022]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="5.5" metricValueMin="5.2" pageId="8" pageNumber="59" unit="mm" value="5.35" valueMax="5.5" valueMin="5.2">5.2-5.5 mm</quantity>
.
</paragraph>
<paragraph id="6F7B36D8C936FFA0668AF52EFDAD145B" blockId="8.[151,1127,966,1631]" pageId="8" pageNumber="59">Colouration. Black, only clypeus along its anterior margin and antennal sockets narrowly yellow-brown; base of antennomere 2 and femora brown.</paragraph>
<paragraph id="6F7B36D8C936FFA0668AF56EFD2F149B" blockId="8.[151,1127,966,1631]" pageId="8" pageNumber="59">
Head distinctly transverse, wider than long (ratio 39:
<quantity id="A83C9B3DC936FFA064CDF56EFC78143B" box="[754,808,1095,1119]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="8" pageNumber="59" unit="in" value="36.0">36 in</quantity>
male, 39:
<quantity id="A83C9B3DC936FFA065A2F56EFC83143B" box="[925,979,1095,1119]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="8" pageNumber="59" unit="in" value="33.0">33 in</quantity>
female), with well marked posterior angles, subparallel-sided, only medially slightly concave. Eyes almost as long as temples (ratio 14: 15). Four coarse punctueres between eyes, distance between medial interocular punctures about four times as large as distance between medial and lateral interocular punctures. Temporal area with many punctures of different size and with one long black bristle. Surface without microsculpture.
</paragraph>
<paragraph id="6F7B36D8C936FFA0668AF42EFD6B151B" blockId="8.[151,1127,966,1631]" pageId="8" pageNumber="59">Antennae long, slightly widened towards apex, reaching posterior fifth of pronotum when reclined. Antennomere 1 distinctly shorter than antennomeres 2-3 combined, antennomere 2 slightly shorter than antennomere 3, antennomeres 4-10 of equal length, antennomere 11 somewhat longer than antennomere 10.</paragraph>
<paragraph id="6F7B36D8C936FFA0668AF4AEFB4215BB" blockId="8.[151,1127,966,1631]" pageId="8" pageNumber="59">Pronotum as long as wide, subparallel-sided, anterior angles almost rectangular, with very apex obtusely rounded, posterior angles markedly rounded. Each dorsal row with two puntures, far away from each other; sublateral rows missing. Surface without microsculpture.</paragraph>
<paragraph id="6F7B36D8C936FFA0668AF4CEFE67167B" blockId="8.[151,1127,966,1631]" pageId="8" pageNumber="59">Scutellum coarsely and densely punctate, punctures somewhat smaller than those of elytra. Setation black.</paragraph>
<paragraph id="6F7B36D8C936FFA2668AF70EFE761170" blockId="8.[151,1127,966,1631]" lastBlockId="10.[151,1127,219,884]" lastPageId="10" lastPageNumber="61" pageId="8" pageNumber="59">Elytra distinctly wider than long (ratio 61: 55), parallel-sided, vaguely widened posteriad, posterior margin relatively deeply cut out. Punctation coarse and dense, punctures somewhat larger than eye-facets, interspaces smaller than punctures. Surface without microsculpture, setation grey.</paragraph>
<caption id="3BBB6650C937FFA1664EF4DDFDE21658" ID-DOI="http://doi.org/10.5281/zenodo.4503610" ID-Zenodo-Dep="4503610" httpUri="https://zenodo.org/record/4503610/files/figure.png" pageId="9" pageNumber="60" startId="9.[113,152,1524,1543]" targetBox="[121,1082,220,1478]" targetPageId="9">
<paragraph id="6F7B36D8C937FFA1664EF4DDFDE21658" blockId="9.[113,1089,1524,1597]" pageId="9" pageNumber="60">
Figs. 20-28. 20-24 –
<taxonomicName id="A8C44D5BC937FFA16713F4DCFDD21663" authority="Levasseur, 1967" authorityName="Levasseur" authorityYear="1967" box="[300,642,1524,1543]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="lamtoensis">
<emphasis id="5DB0EACAC937FFA16713F4DCFEBB1663" box="[300,491,1525,1543]" italics="true" pageId="9" pageNumber="60">Philonthus lamtoensis</emphasis>
Levasseur, 1967
</taxonomicName>
; 25-28 –
<taxonomicName id="A8C44D5BC937FFA164E7F4DCFC8B1663" authority="Bernhauer, 1936" authorityName="Bernhauer" authorityYear="1936" box="[728,987,1524,1543]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="madianus">
<emphasis id="5DB0EACAC937FFA164E7F4DCFC121663" box="[728,834,1525,1543]" italics="true" pageId="9" pageNumber="60">P. madianus</emphasis>
<bibRefCitation id="0B554B29C937FFA16575F4DDFC8B1663" author="BERNHAUER M." box="[842,987,1524,1543]" pageId="9" pageNumber="60" pagination="261 - 267" refId="ref6059" refString="BERNHAUER M. 1936: Neue Staphylinidae aus Angola. Novitates Zoologicae 39: 261 - 267." type="journal article" year="1936">Bernhauer, 1936</bibRefCitation>
</taxonomicName>
. 20-22, 25- 27 – aedeagus: 20, 25 ventral view, 21, 26 – dorsal view, 22, 27 – lateral view; 23, 28 – undersite of paramere, with sensory peg setae; 24 – apical portion of sternite VIII, ventral view.
</paragraph>
</caption>
<paragraph id="6F7B36D8C934FFA2668AF035FC771130" blockId="10.[151,1127,219,884]" pageId="10" pageNumber="61">Legs. Metatibia as long as metatarsus. Metatarsomere 1 almost as long as metatarsomeres 2-4 combined, metatarsomere 5 longer than metatarsomere 2.</paragraph>
<paragraph id="6F7B36D8C934FFA2668AF072FDE311B7" blockId="10.[151,1127,219,884]" pageId="10" pageNumber="61">Abdomen from visible tergite V slightly narrowed towards both base and apex. Elevated area between two basal lines on first three visible tergites very densely and coarsely punctate. Punctation of all tergites dense and slightly finer than that on elytra. Surface without microsculpture. Setation similar to those on elytra.</paragraph>
<paragraph id="6F7B36D8C934FFA2668AF0F2FE7A1257" blockId="10.[151,1127,219,884]" pageId="10" pageNumber="61">
Male. Protarsomeres 1-3 strongly dilated, each densely covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones, heart-shaped. Aedeagus (
<figureCitation id="F7FF2A5DC934FFA266A0F332FE4D1257" box="[159,285,539,563]" captionStart="Figs" captionStartId="9.[113,152,1524,1543]" captionTargetBox="[121,1082,220,1478]" captionTargetId="figure-63@9.[118,1084,219,1479]" captionTargetPageId="9" captionText="Figs. 20-28. 20-24 – Philonthus lamtoensis Levasseur, 1967; 25-28 – P. madianus Bernhauer, 1936. 20-22, 25- 27 – aedeagus: 20, 25 ventral view, 21, 26 – dorsal view, 22, 27 – lateral view; 23, 28 – undersite of paramere, with sensory peg setae; 24 – apical portion of sternite VIII, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4503610" httpUri="https://zenodo.org/record/4503610/files/figure.png" pageId="10" pageNumber="61">Figs. 25-28</figureCitation>
).
</paragraph>
<paragraph id="6F7B36D8C934FFA2668AF312FE4D12F7" blockId="10.[151,1127,219,884]" pageId="10" pageNumber="61">Female. Protarsomeres 1-3 moderately dilated, slightly sub-bilobed, each covered with modified pale setae ventrally, protarsomere 4 scarcely dilated, distinctly narrower than preceding ones.</paragraph>
</subSubSection>
<subSubSection id="27DE6553C934FFA266A8F3B2FB0012B7" pageId="10" pageNumber="61" type="diagnosis">
<paragraph id="6F7B36D8C934FFA266A8F3B2FB0012B7" blockId="10.[151,1127,219,884]" pageId="10" pageNumber="61">
<emphasis id="5DB0EACAC934FFA266A8F3B2FEDC12D7" bold="true" box="[151,396,667,691]" pageId="10" pageNumber="61">Differential diagnosis.</emphasis>
<taxonomicName id="A8C44D5BC934FFA267AFF3B4FD3E12D0" authorityName="Bernhauer" authorityYear="1936" box="[400,622,669,692]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="madianus">
<emphasis id="5DB0EACAC934FFA267AFF3B4FD3E12D0" box="[400,622,669,692]" italics="true" pageId="10" pageNumber="61">Philonthus madianus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="A8C44D5BC934FFA2651CF3B4FC9412D0" authorityName="Tottenham" authorityYear="1949" box="[803,964,669,692]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="peripateticus">
<emphasis id="5DB0EACAC934FFA2651CF3B4FC9412D0" box="[803,964,669,692]" italics="true" pageId="10" pageNumber="61">P. peripateticus</emphasis>
</taxonomicName>
but it differs by the slenderer and longer antennae, wider head and by the different shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C934FFA266A8F3F5FEDC1337" pageId="10" pageNumber="61" type="discussion">
<paragraph id="6F7B36D8C934FFA266A8F3F5FEDE1357" blockId="10.[151,1127,219,884]" pageId="10" pageNumber="61">
<emphasis id="5DB0EACAC934FFA266A8F3F5FE561290" bold="true" box="[151,262,732,756]" pageId="10" pageNumber="61">Remarks.</emphasis>
It is not necessary to designate a
<typeStatus id="B07F887AC934FFA26461F3F2FD901297" box="[606,704,731,755]" pageId="10" pageNumber="61" type="lectotype">lectotype</typeStatus>
in this case, because evidently Berhauer clearly separated between ‘Typus’ and ‘Cotypus’, which translates to
<typeStatus id="B07F887AC934FFA26545F3D2FC871377" box="[890,983,763,787]" pageId="10" pageNumber="61" type="holotype">holotype</typeStatus>
and
<typeStatus id="B07F887AC934FFA26234F3D5FB371370" box="[1035,1127,764,788]" pageId="10" pageNumber="61" type="paratype">paratype</typeStatus>
of the modern concept.
</paragraph>
<paragraph id="6F7B36D8C934FFA266A8F212FEDC1337" blockId="10.[151,1127,219,884]" box="[151,396,827,851]" pageId="10" pageNumber="61">
<emphasis id="5DB0EACAC934FFA266A8F212FE451337" bold="true" box="[151,277,827,851]" pageId="10" pageNumber="61">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="27DE6553C934FFA266A8F272FC001310" box="[151,848,859,884]" pageId="10" pageNumber="61" type="distribution">
<paragraph id="6F7B36D8C934FFA266A8F272FC001310" blockId="10.[151,1127,219,884]" box="[151,848,859,884]" pageId="10" pageNumber="61">
<emphasis id="5DB0EACAC934FFA266A8F272FE7A1317" bold="true" box="[151,298,859,883]" pageId="10" pageNumber="61">Distribution.</emphasis>
<collectingCountry id="17D37648C934FFA26711F272FE2F1317" box="[302,383,859,883]" name="Angola" pageId="10" pageNumber="61">Angola</collectingCountry>
,
<collectingCountry id="17D37648C934FFA267B5F272FE8D1317" box="[394,477,859,883]" name="Uganda" pageId="10" pageNumber="61">Uganda</collectingCountry>
(
<bibRefCitation id="0B554B29C934FFA267D3F275FDF31317" author="BERNHAUER M." box="[492,675,859,884]" pageId="10" pageNumber="61" pagination="261 - 267" refId="ref6059" refString="BERNHAUER M. 1936: Neue Staphylinidae aus Angola. Novitates Zoologicae 39: 261 - 267." type="journal article" year="1936">BERNHAUER 1936</bibRefCitation>
,
<bibRefCitation id="0B554B29C934FFA26492F275FC151310" author="HERMAN L. H." box="[685,837,860,884]" pageId="10" pageNumber="61" pagination="2441 - 3020" refId="ref6077" refString="HERMAN L. H. 2001: Catalog of the Staphylinidae (Insecta, Coleoptera); 1758 to the end of the second millennium. V. Staphylinine group (Part 2): Diochini, Maorothiini, Othiini, Platyprosopini, Staphilinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History 265: 2441 - 3020." type="journal article" year="2001">HERMAN 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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