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<document id="9326C686B6BFC1FC84247801E6B5CDC9" ID-CLB-Dataset="39184" ID-DOI="10.11646/zootaxa.4105.5.2" ID-GBIF-Dataset="60ca4dbd-002b-4ca6-999d-9174f9b02324" ID-ISSN="1175-5326" ID-Zenodo-Dep="255425" ID-ZooBank="52401DF9-2901-4AF6-BA17-696D39280B3C" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1461736713249" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2016" docId="E338D6753516216EFF05FED473D26CDD" docLanguage="en" docName="zootaxa.4105.5.2.pdf.imf" docOrigin="Zootaxa 4105 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guyalna coffea, n. comb" docType="treatment" docVersion="20" lastPageNumber="438" masterDocId="1F01AE0D351E2166FF92FF9876486E39" masterDocTitle="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" masterLastPageNumber="454" masterPageNumber="430" pageNumber="438" updateTime="1698625263794" updateUser="plazi">
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<mods:title id="C0DCD3960356E8F61CCD60AB461FDFAD">The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species</mods:title>
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<mods:namePart id="F42076207190ECBA8DB5884D0E6F284E">Sanborn, Allen F.</mods:namePart>
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<mods:date id="250CC9EA87CA8C123780F981CDCBF833">2016</mods:date>
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<mods:identifier id="C8E35F78E97DB9E6F1EED6BF4BD82AC7" type="DOI">10.11646/zootaxa.4105.5.2</mods:identifier>
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<treatment id="E338D6753516216EFF05FED473D26CDD" ID-DOI="http://doi.org/10.5281/zenodo.3511629" ID-GBIF-Taxon="119651792" ID-Zenodo-Dep="3511629" LSID="urn:lsid:plazi:treatment:E338D6753516216EFF05FED473D26CDD" httpUri="http://treatment.plazi.org/id/E338D6753516216EFF05FED473D26CDD" lastPageNumber="438" pageId="8" pageNumber="438">
<subSubSection id="238B34E83516216EFF05FED475226F5F" box="[151,874,332,358]" pageId="8" pageNumber="438" type="nomenclature">
<paragraph id="6B2E67633516216EFF05FED475226F5F" blockId="8.[151,874,332,391]" box="[151,874,332,358]" pageId="8" pageNumber="438">
<heading id="3066D00F3516216EFF05FED475226F5F" bold="true" box="[151,874,332,358]" fontSize="11" level="1" pageId="8" pageNumber="438" reason="1">
<emphasis id="59E5BB713516216EFF05FED475226F5F" bold="true" box="[151,874,332,358]" pageId="8" pageNumber="438">
<taxonomicName id="AC911CE03516216EFF05FED474B06F5F" ID-CoL="3HN6T" authority="(Sanborn, Moore &amp; Young, 2008)" baseAuthorityName="Sanborn, Moore &amp; Young" baseAuthorityYear="2008" box="[151,760,332,358]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="438" phylum="Arthropoda" rank="species" species="coffea" status="comb. nov.">
<emphasis id="59E5BB713516216EFF05FED477186F5F" bold="true" box="[151,336,332,358]" italics="true" pageId="8" pageNumber="438">Guyalna coffea</emphasis>
(
<bibRefCitation id="0F001A923516216EFEF2FED474B86F5F" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[352,752,332,358]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="8" pageNumber="438" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">Sanborn, Moore &amp; Young, 2008</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="42D6060A3516216EFC92FED4752D6F5F" box="[768,869,332,358]" pageId="8" pageNumber="438">n. comb</taxonomicNameLabel>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="238B34E83516216EFF05FEF676A36FBE" box="[151,235,366,391]" pageId="8" pageNumber="438" type="description">
<paragraph id="6B2E67633516216EFF05FEF676A36FBE" blockId="8.[151,874,332,391]" box="[151,235,366,391]" pageId="8" pageNumber="438">
(
<figureCitation id="F3AA7BE63516216EFF0DFEF676AA6FBE" box="[159,226,366,391]" captionStart="FIGURE 8" captionStartId="8.[151,250,1375,1397]" captionTargetBox="[151,1435,796,1353]" captionTargetId="figure@8.[151,1436,796,1354]" captionTargetPageId="8" captionText="FIGURE 8. Guyalna coffea (Sanborn, Moore &amp; Young, 2008) n. comb. A, paratype male, Costa Rica (AFSC). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255433/files/figure.png" pageId="8" pageNumber="438">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="238B34E83516216EFF05FE2E74886FD2" pageId="8" pageNumber="438" type="reference_group">
<paragraph id="6B2E67633516216EFF05FE2E743D6FF5" blockId="8.[151,704,438,491]" box="[151,629,438,460]" pageId="8" pageNumber="438">
<treatmentCitationGroup id="4B81404D3516216EFF05FE2E743D6FF5" box="[151,629,438,460]" pageId="8" pageNumber="438">
<taxonomicName id="AC911CE03516216EFF05FE2E77046FF5" ID-CoL="8HGGV" authority="(Sanborn, Moore &amp; Young, 2008)" baseAuthorityName="Sanborn, Moore &amp; Young" baseAuthorityYear="2008" box="[151,332,438,460]" class="Insecta" family="Cicadidae" genus="Fidicina" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="438" phylum="Arthropoda" rank="genus" status="comb. nov.">
<emphasis id="59E5BB713516216EFF05FE2E77046FF5" box="[151,332,438,460]" italics="true" pageId="8" pageNumber="438">Fidicina “coffea”</emphasis>
</taxonomicName>
nom. nud.
<treatmentCitation id="EA3041723516216EFE2DFE2F74396FF5" author="Young" box="[447,625,438,460]" page="270" pageId="8" pageNumber="438" year="1977">
<bibRefCitation id="0F001A923516216EFE2DFE2F74396FF5" author="Young, A. M." box="[447,625,438,460]" firstAuthor="Young" journalOrPublisher="Revista de Biologica Tropical" page="270" pageId="8" pageNumber="438" pagination="267 - 279" part="24" refId="ref15038" refString="Young, A. M. (1977) Notes on the faunistic complexity of cicadas (Homoptera: Cicadidae) in Northern Costa Rica. Revista de Biologica Tropical, 24, 267 - 279." title="Notes on the faunistic complexity of cicadas (Homoptera: Cicadidae) in Northern Costa Rica" type="journal article" year="1977">Young 1977: 270</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B2E67633516216EFF05FE4D74886FD2" blockId="8.[151,704,438,491]" box="[151,704,469,491]" pageId="8" pageNumber="438">
<treatmentCitationGroup id="4B81404D3516216EFF05FE4D74886FD2" box="[151,704,469,491]" pageId="8" pageNumber="438">
<taxonomicName id="AC911CE03516216EFF05FE4D74F36FD2" ID-CoL="6HYSB" authority="Sanborn, Moore &amp; Young 2008" authorityName="Sanborn, Moore &amp; Young" authorityPageNumber="3" authorityYear="2008" box="[151,699,469,491]" class="Insecta" family="Cicadidae" genus="Fidicinoides" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="438" phylum="Arthropoda" rank="species" species="coffea">
<emphasis id="59E5BB713516216EFF05FE4D77126FD2" box="[151,346,469,491]" italics="true" pageId="8" pageNumber="438">Fidicinoides coffea</emphasis>
<treatmentCitation id="EA3041723516216EFEF2FE4D74F36FD2" ID-DOI="http://doi.org/10.5281/zenodo.3499864" ID-Zenodo-Dep="3499864" author="Sanborn" box="[352,699,469,491]" httpUri="http://treatment.plazi.org/id/03846100FFA94F2BFCE5FE6DFB62F962" page="3" pageId="8" pageNumber="438" year="2008">
<bibRefCitation id="0F001A923516216EFEF2FE4D74F36FD2" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[352,699,469,491]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" page="3" pageId="8" pageNumber="438" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">Sanborn, Moore &amp; Young 2008: 3</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="238B34E83516216EFF05FD8F756D6C09" box="[151,805,535,560]" pageId="8" pageNumber="438" type="materials_examined">
<paragraph id="6B2E67633516216EFF05FD8F756D6C09" blockId="8.[151,1436,535,741]" box="[151,805,535,560]" pageId="8" pageNumber="438">
<materialsCitation id="DBF96D3E3516216EFF05FD8F756D6C09" ID-GBIF-Occurrence="4115800319" box="[151,805,535,560]" country="Costa Rica" location="San Ramon" pageId="8" pageNumber="438" specimenCount="1" stateProvince="Alajuela Province" typeStatus="type">
<emphasis id="59E5BB713516216EFF05FD8F777D6C09" bold="true" box="[151,309,535,560]" pageId="8" pageNumber="438">
<typeStatus id="B42AD9C13516216EFF05FD8F769A6C09" box="[151,210,535,560]" pageId="8" pageNumber="438">Type</typeStatus>
locality.
</emphasis>
<location id="6E4E31B83516216EFEAEFD8077896C16" LSID="urn:lsid:plazi:treatment:E338D6753516216EFF05FED473D26CDD:6E4E31B83516216EFEAEFD8077896C16" box="[316,449,536,560]" country="Costa Rica" name="San Ramon" pageId="8" pageNumber="438" stateProvince="Alajuela Province">San Ramon</location>
,
<collectingRegion id="A955A9813516216EFE5EFD8F74D06C09" box="[460,664,535,560]" country="Costa Rica" name="Alajuela" pageId="8" pageNumber="438">Alajuela Province</collectingRegion>
,
<collectingCountry id="138627F33516216EFD31FD8075696C09" box="[675,801,535,560]" name="Costa Rica" pageId="8" pageNumber="438">Costa Rica</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="238B34E83516216EFF55FDA477576CF9" pageId="8" pageNumber="438" type="discussion">
<paragraph id="6B2E67633516216EFF55FDA477576CF9" blockId="8.[151,1436,535,741]" pageId="8" pageNumber="438">
<emphasis id="59E5BB713516216EFF55FDA477736C6C" bold="true" box="[199,315,572,597]" pageId="8" pageNumber="438">Remarks.</emphasis>
Re-examination of the species shows it possesses three-part tarsi and the curved timbal cover apex characteristic of
<taxonomicName id="AC911CE03516216EFEC3FDC777F96C41" box="[337,433,607,632]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB713516216EFEC3FDC777F96C41" box="[337,433,607,632]" italics="true" pageId="8" pageNumber="438">Guyalna</emphasis>
</taxonomicName>
(
<figureCitation id="F3AA7BE63516216EFE52FDC7744C6C41" box="[448,516,607,632]" captionStart="FIGURE 8" captionStartId="8.[151,250,1375,1397]" captionTargetBox="[151,1435,796,1353]" captionTargetId="figure@8.[151,1436,796,1354]" captionTargetPageId="8" captionText="FIGURE 8. Guyalna coffea (Sanborn, Moore &amp; Young, 2008) n. comb. A, paratype male, Costa Rica (AFSC). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255433/files/figure.png" pageId="8" pageNumber="438">Fig. 8</figureCitation>
). As a result the species is herby reassigned to
<taxonomicName id="AC911CE03516216EFB9DFDC772276C41" box="[1039,1135,607,632]" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="438" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB713516216EFB9DFDC772276C41" box="[1039,1135,607,632]" italics="true" pageId="8" pageNumber="438">Guyalna</emphasis>
</taxonomicName>
to become
<taxonomicName id="AC911CE03516216EFB62FDC7745A6CA5" authority="Sanborn, Moore &amp; Young, 2008" authorityName="Sanborn, Moore &amp; Young" authorityYear="2008" class="Insecta" family="Cicadidae" genus="Guyalna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="438" phylum="Arthropoda" rank="species" species="coffea" status="comb. nov.">
<emphasis id="59E5BB713516216EFB62FDC773D46C41" box="[1264,1436,607,632]" italics="true" pageId="8" pageNumber="438">Guyalna coffea</emphasis>
(
<bibRefCitation id="0F001A923516216EFF0CFD1C74436CA5" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[158,523,644,669]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="8" pageNumber="438" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">Sanborn, Moore &amp; Young, 2008</bibRefCitation>
)
</taxonomicName>
<emphasis id="59E5BB713516216EFD8BFD1C74376CA4" bold="true" box="[537,639,644,669]" pageId="8" pageNumber="438">
<taxonomicNameLabel id="42D6060A3516216EFD8BFD1C74326CA4" box="[537,634,644,669]" pageId="8" pageNumber="438">n. comb</taxonomicNameLabel>
.
</emphasis>
Body length
<quantity id="AC69CA863516216EFC87FD1C75F26CA5" box="[789,954,644,669]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" metricValueMax="2.66" metricValueMin="2.08" pageId="8" pageNumber="438" unit="mm" value="23.7" valueMax="26.6" valueMin="20.8">20.8–26.6 mm</quantity>
, fore wing length
<quantity id="AC69CA863516216EFB17FD1C73606CA5" box="[1157,1320,644,669]" metricMagnitude="-2" metricUnit="m" metricValue="3.505" metricValueMax="3.76" metricValueMin="3.25" pageId="8" pageNumber="438" unit="mm" value="35.05" valueMax="37.6" valueMin="32.5">32.5–37.6 mm</quantity>
(
<bibRefCitation id="0F001A923516216EFAA4FD1C775B6CF9" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." journalOrPublisher="Zootaxa" pageId="8" pageNumber="438" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">
Sanborn,
<emphasis id="59E5BB713516216EFF05FD3176876CF9" box="[151,207,679,704]" italics="true" pageId="8" pageNumber="438">et al.</emphasis>
2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="238B34E83516216EFF55FD5473D26CDD" box="[199,1434,716,741]" pageId="8" pageNumber="438" type="distribution">
<paragraph id="6B2E67633516216EFF55FD5473D26CDD" blockId="8.[151,1436,535,741]" box="[199,1434,716,741]" pageId="8" pageNumber="438">
<emphasis id="59E5BB713516216EFF55FD5477286CDC" bold="true" box="[199,352,716,741]" pageId="8" pageNumber="438">Distribution.</emphasis>
The species has been reported previously from
<collectingCountry id="138627F33516216EFCEAFD5475BA6CDC" box="[888,1010,716,741]" name="Costa Rica" pageId="8" pageNumber="438">Costa Rica</collectingCountry>
(
<bibRefCitation id="0F001A923516216EFB93FD5472AA6CDD" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[1025,1250,716,741]" etAl="et al." firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="8" pageNumber="438" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">
Sanborn
<emphasis id="59E5BB713516216EFBF5FD5572E86CDC" box="[1127,1184,716,741]" italics="true" pageId="8" pageNumber="438">et al.</emphasis>
2008
</bibRefCitation>
;
<bibRefCitation id="0F001A923516216EFB7FFD5473C66CDD" author="Sanborn, A. F." box="[1261,1422,716,741]" firstAuthor="Sanborn" journalOrPublisher="Check List" pageId="8" pageNumber="438" pagination="246 - 252" part="10" refId="ref14181" refString="Sanborn, A. F. (2014) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Costa Rica including new records for fourteen species. Check List, 10, 246 - 252. http: // dx. doi. org / 10.15560 / 10.2.246" title="Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Costa Rica including new records for fourteen species" type="journal article" year="2014">Sanborn 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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