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<document id="D34E29C1111F28FC07C6AEB6C533EF6F" ID-CLB-Dataset="39184" ID-DOI="10.11646/zootaxa.4105.5.2" ID-GBIF-Dataset="60ca4dbd-002b-4ca6-999d-9174f9b02324" ID-ISSN="1175-5326" ID-Zenodo-Dep="255425" ID-ZooBank="52401DF9-2901-4AF6-BA17-696D39280B3C" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1461736713249" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2016" docId="E338D675350D2175FF05FE3275A56D47" docLanguage="en" docName="zootaxa.4105.5.2.pdf.imf" docOrigin="Zootaxa 4105 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dorisiana cachla, n. comb" docType="treatment" docVersion="22" lastPageNumber="449" masterDocId="1F01AE0D351E2166FF92FF9876486E39" masterDocTitle="The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species" masterLastPageNumber="454" masterPageNumber="430" pageNumber="449" updateTime="1698625263794" updateUser="plazi">
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<mods:title id="B319E9212956E62D80A7567D8434BEBA">The cicada genus Guyalna Boulard &amp; Martinelli, 1996 (Hemiptera: Cicadidae: Cicadinae: Fidicinini): generic description, twelve new combinations, and a key to species</mods:title>
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<mods:namePart id="ED3D5B122FEFC14A1CB267E2BC53932A">Sanborn, Allen F.</mods:namePart>
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<mods:date id="60F3608AFC09E9AF6E294491163C7B6C">2016</mods:date>
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<treatment id="E338D675350D2175FF05FE3275A56D47" ID-DOI="http://doi.org/10.5281/zenodo.4951988" ID-GBIF-Taxon="119651811" ID-Zenodo-Dep="4951988" LSID="urn:lsid:plazi:treatment:E338D675350D2175FF05FE3275A56D47" httpUri="http://treatment.plazi.org/id/E338D675350D2175FF05FE3275A56D47" lastPageNumber="449" pageId="19" pageNumber="449">
<subSubSection id="238B34E8350D2175FF05FE3274D06FFD" box="[151,664,426,452]" pageId="19" pageNumber="449" type="nomenclature">
<paragraph id="6B2E6763350D2175FF05FE3274D06FFD" blockId="19.[151,664,426,486]" box="[151,664,426,452]" pageId="19" pageNumber="449">
<heading id="3066D00F350D2175FF05FE3274D06FFD" bold="true" box="[151,664,426,452]" fontSize="11" level="1" pageId="19" pageNumber="449" reason="1">
<emphasis id="59E5BB71350D2175FF05FE3274D06FFD" bold="true" box="[151,664,426,452]" pageId="19" pageNumber="449">
<taxonomicName id="AC911CE0350D2175FF05FE32746D6FFD" ID-CoL="37DNW" authority="(Distant, 1899)" baseAuthorityName="Distant" baseAuthorityYear="1899" box="[151,549,426,452]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="species" species="cachla" status="comb. nov.">
<emphasis id="59E5BB71350D2175FF05FE32772F6FFD" bold="true" box="[151,359,426,452]" italics="true" pageId="19" pageNumber="449">Dorisiana cachla</emphasis>
(
<bibRefCitation id="0F001A92350D2175FEE4FE3274556FFD" author="Distant, W. L." box="[374,541,426,452]" firstAuthor="Distant" journalOrPublisher="Annals and Magazine of Natural History" pageId="19" pageNumber="449" pagination="112 - 113" part="4" refId="ref12934" refString="Distant, W. L. (1899) Some apparently undescribed Neotropical Homoptera. Annals and Magazine of Natural History, Series 7, 4, 112 - 113. http: // dx. doi. org / 10.1080 / 00222939908678169" title="Some apparently undescribed Neotropical Homoptera" type="journal article" year="1899">Distant, 1899</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="42D6060A350D2175FDBFFE3274DB6FFD" box="[557,659,426,452]" pageId="19" pageNumber="449">n. comb</taxonomicNameLabel>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="238B34E8350D2175FF05FE5576B16FDF" box="[151,249,461,486]" pageId="19" pageNumber="449" type="description">
<paragraph id="6B2E6763350D2175FF05FE5576B16FDF" blockId="19.[151,664,426,486]" box="[151,249,461,486]" pageId="19" pageNumber="449">
(
<figureCitation id="F3AA7BE6350D2175FF0DFE5576B96FDF" box="[159,241,461,486]" captionStart="FIGURE 26" captionStartId="19.[151,250,1500,1522]" captionTargetBox="[151,1436,949,1478]" captionTargetId="figure@19.[151,1436,949,1478]" captionTargetPageId="19" captionText="FIGURE 26. Dorisiana cachla (Distant, 1899) n. comb. A, male, Costa Rica (AFSC). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255451/files/figure.png" pageId="19" pageNumber="449">Fig. 26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="238B34E8350D2175FF05FD96749B6C7A" pageId="19" pageNumber="449" type="reference_group">
<paragraph id="6B2E6763350D2175FF05FD9677B06C1D" blockId="19.[151,723,526,579]" box="[151,504,526,548]" pageId="19" pageNumber="449">
<treatmentCitationGroup id="4B81404D350D2175FF05FD9677B06C1D" box="[151,504,526,548]" pageId="19" pageNumber="449">
<taxonomicName id="AC911CE0350D2175FF05FD9677BC6C1D" ID-CoL="6HYYY" authority="Distant 1899" authorityName="Distant" authorityPageNumber="112" authorityYear="1899" box="[151,500,526,548]" class="Insecta" family="Cicadidae" genus="Fidicina" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="species" species="cachla">
<emphasis id="59E5BB71350D2175FF05FD96777F6C1D" box="[151,311,526,548]" italics="true" pageId="19" pageNumber="449">Fidicina cachla</emphasis>
<treatmentCitation id="EA304172350D2175FEAFFD9677BC6C1D" author="Distant" box="[317,500,526,548]" page="112" pageId="19" pageNumber="449" year="1899">
<bibRefCitation id="0F001A92350D2175FEAFFD9677BC6C1D" author="Distant, W. L." box="[317,500,526,548]" firstAuthor="Distant" journalOrPublisher="Annals and Magazine of Natural History" page="112" pageId="19" pageNumber="449" pagination="112 - 113" part="4" refId="ref12934" refString="Distant, W. L. (1899) Some apparently undescribed Neotropical Homoptera. Annals and Magazine of Natural History, Series 7, 4, 112 - 113. http: // dx. doi. org / 10.1080 / 00222939908678169" title="Some apparently undescribed Neotropical Homoptera" type="journal article" year="1899">Distant 1899: 112</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B2E6763350D2175FF05FDB5749B6C7A" blockId="19.[151,723,526,579]" box="[151,723,557,579]" pageId="19" pageNumber="449">
<treatmentCitationGroup id="4B81404D350D2175FF05FDB5749B6C7A" box="[151,723,557,579]" pageId="19" pageNumber="449">
<taxonomicName id="AC911CE0350D2175FF05FDB577286C7A" ID-CoL="8HXLR" authority="(Distant, 1899)" baseAuthorityName="Distant" baseAuthorityYear="1899" box="[151,352,557,579]" class="Insecta" family="Cicadidae" genus="Fidicinoides" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="species" species="cachla">
<emphasis id="59E5BB71350D2175FF05FDB577286C7A" box="[151,352,557,579]" italics="true" pageId="19" pageNumber="449">Fidicinoides cachla</emphasis>
</taxonomicName>
<treatmentCitation id="EA304172350D2175FEF7FDB574876C7A" ID-DOI="http://doi.org/10.5281/zenodo.8189676" ID-Zenodo-Dep="8189676" author="Sanborn" box="[357,719,557,579]" httpUri="http://treatment.plazi.org/id/03846100FFBB4F32FCB7FA7CFA54FA54" page="10" pageId="19" pageNumber="449" year="2008">
<bibRefCitation id="0F001A92350D2175FEF7FDB574876C7A" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[357,719,557,579]" firstAuthor="Sanborn" journalOrPublisher="Zootaxa" page="10" pageId="19" pageNumber="449" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">Sanborn, Moore &amp; Young 2008: 10</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="238B34E8350D2175FF05FDF174A66CBB" box="[151,750,617,642]" pageId="19" pageNumber="449" type="materials_examined">
<paragraph id="6B2E6763350D2175FF05FDF174A66CBB" blockId="19.[151,1436,617,894]" box="[151,750,617,642]" pageId="19" pageNumber="449">
<materialsCitation id="DBF96D3E350D2175FF05FDF174A66CBB" ID-GBIF-Occurrence="4115800311" box="[151,750,617,642]" country="Costa Rica" location="Rosaria de Desamparados" pageId="19" pageNumber="449" specimenCount="1" typeStatus="type">
<emphasis id="59E5BB71350D2175FF05FDF1777D6CBB" bold="true" box="[151,309,617,642]" pageId="19" pageNumber="449">
<typeStatus id="B42AD9C1350D2175FF05FDF1769A6CBB" box="[151,210,617,642]" pageId="19" pageNumber="449">Type</typeStatus>
locality.
</emphasis>
<location id="6E4E31B8350D2175FEAEFDF1742A6CBB" LSID="urn:lsid:plazi:treatment:E338D675350D2175FF05FE3275A56D47:6E4E31B8350D2175FEAEFDF1742A6CBB" box="[316,610,617,642]" country="Costa Rica" name="Rosaria de Desamparados" pageId="19" pageNumber="449">Rosaria de Desamparados</location>
,
<collectingCountry id="138627F3350D2175FDFFFDF274A36CBB" box="[621,747,617,642]" name="Costa Rica" pageId="19" pageNumber="449">Costa Rica</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="238B34E8350D2175FF55FD1574A76D0C" pageId="19" pageNumber="449" type="discussion">
<paragraph id="6B2E6763350D2175FF55FD1574A76D0C" blockId="19.[151,1436,617,894]" pageId="19" pageNumber="449">
<emphasis id="59E5BB71350D2175FF55FD1577736C9F" bold="true" box="[199,315,653,678]" pageId="19" pageNumber="449">Remarks.</emphasis>
The species was assigned to
<taxonomicName id="AC911CE0350D2175FD04FD1575696C9F" box="[662,801,653,678]" class="Insecta" family="Cicadidae" genus="Fidicinoides" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="genus">Fidicinoides</taxonomicName>
based on the shape of the cruciform elevation and the relationship between the posterior mesonotum and the metanotum (
<bibRefCitation id="0F001A92350D2175FC0BFD2972C96CF3" author="Sanborn, A. F. &amp; Moore, T. E. &amp; Young, A. M." box="[921,1153,689,714]" etAl="et al." firstAuthor="Sanborn" journalOrPublisher="Zootaxa" pageId="19" pageNumber="449" pagination="1 - 20" part="1846" refId="ref14538" refString="Sanborn, A. F., Moore, T. E. &amp; Young, A. M. (2008) Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica. Zootaxa, 1846, 1 - 20." title="Two new cicada species from Costa Rica (Hemiptera: Cicadomorpha: Cicadidae) with a key to Fidicinoides in Costa Rica" type="journal article" year="2008">
Sanborn
<emphasis id="59E5BB71350D2175FB93FD2A72746CF3" box="[1025,1084,689,714]" italics="true" pageId="19" pageNumber="449">et al.</emphasis>
2008
</bibRefCitation>
). Further examination of the species shows it possesses three-part tarsi and the triangular timbal cover apex characteristic of
<taxonomicName id="AC911CE0350D2175FB7CFD4E73286CD4" box="[1262,1376,726,749]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB71350D2175FB7CFD4E73286CD4" box="[1262,1376,726,749]" italics="true" pageId="19" pageNumber="449">Dorisiana</emphasis>
</taxonomicName>
(
<figureCitation id="F3AA7BE6350D2175FAFCFD4D76FF6D2B" captionStart="FIGURE 26" captionStartId="19.[151,250,1500,1522]" captionTargetBox="[151,1436,949,1478]" captionTargetId="figure@19.[151,1436,949,1478]" captionTargetPageId="19" captionText="FIGURE 26. Dorisiana cachla (Distant, 1899) n. comb. A, male, Costa Rica (AFSC). Scale bar: 2 cm; B, timbal cover. Scale bar: 2 mm." httpUri="https://zenodo.org/record/255451/files/figure.png" pageId="19" pageNumber="449">Fig. 26</figureCitation>
). As a result the species is herby reassigned to
<taxonomicName id="AC911CE0350D2175FD55FD6275716D28" box="[711,825,762,785]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="genus">
<emphasis id="59E5BB71350D2175FD55FD6275716D28" box="[711,825,762,785]" italics="true" pageId="19" pageNumber="449">Dorisiana</emphasis>
</taxonomicName>
to become
<taxonomicName id="AC911CE0350D2175FC2EFD6273676D2B" authority="Walker, 1850" authorityName="Walker" authorityYear="1850" box="[956,1327,761,786]" class="Insecta" family="Cicadidae" genus="Dorisiana" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="449" phylum="Arthropoda" rank="species" species="cachla" status="comb. nov.">
<emphasis id="59E5BB71350D2175FC2EFD6272C86D2B" box="[956,1152,761,786]" italics="true" pageId="19" pageNumber="449">Dorisiana cachla</emphasis>
(
<bibRefCitation id="0F001A92350D2175FB1CFD61736E6D2B" author="Walker, F." bookContentInfo="260 pp." box="[1166,1318,761,786]" firstAuthor="Walker" journalOrPublisher="British Museum Trustees, London" pageId="19" pageNumber="449" refId="ref14974" refString="Walker, F. (1850) List of the Specimens of Homopterous Insects in the Collection of the British Museum. British Museum Trustees, London, 260 pp." title="List of the Specimens of Homopterous Insects in the Collection of the British Museum" type="book" year="1850">Walker, 1850</bibRefCitation>
)
</taxonomicName>
<emphasis id="59E5BB71350D2175FAA4FD6173D46D2B" bold="true" box="[1334,1436,761,786]" pageId="19" pageNumber="449">
<taxonomicNameLabel id="42D6060A350D2175FAA4FD6173DF6D2B" box="[1334,1431,761,786]" pageId="19" pageNumber="449">n. comb</taxonomicNameLabel>
.
</emphasis>
Body length
<quantity id="AC69CA86350D2175FEB5FC8677EB6D0C" box="[295,419,798,822]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="19" pageNumber="449" unit="mm" value="31.0" valueMax="33.0" valueMin="29.0">29–33 mm</quantity>
, fore wing length
<quantity id="AC69CA86350D2175FDFCFC8674A26D0C" box="[622,746,798,822]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="19" pageNumber="449" unit="mm" value="41.0" valueMax="44.0" valueMin="38.0">38–44 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="238B34E8350D2175FF55FCD975A56D47" pageId="19" pageNumber="449" type="distribution">
<paragraph id="6B2E6763350D2175FF55FCD975A56D47" blockId="19.[151,1436,617,894]" pageId="19" pageNumber="449">
<emphasis id="59E5BB71350D2175FF55FCD977286D63" bold="true" box="[199,352,833,858]" pageId="19" pageNumber="449">Distribution.</emphasis>
The species has been reported previously from
<collectingCountry id="138627F3350D2175FC19FCDA72426D63" box="[907,1034,833,858]" name="Costa Rica" pageId="19" pageNumber="449">Costa Rica</collectingCountry>
,
<collectingCountry id="138627F3350D2175FB85FCD972CC6D63" box="[1047,1156,833,858]" name="Honduras" pageId="19" pageNumber="449">Honduras</collectingCountry>
and
<collectingCountry id="138627F3350D2175FB2DFCD9737A6D63" box="[1215,1330,833,858]" name="Nicaragua" pageId="19" pageNumber="449">Nicaragua</collectingCountry>
(
<bibRefCitation id="0F001A92350D2175FAD1FCD9769A6D47" author="Metcalf, Z. P." firstAuthor="Metcalf" journalOrPublisher="North Carolina State College Contribution" pageId="19" pageNumber="449" pagination="1 - 585" part="1502" refId="ref13609" refString="Metcalf, Z. P. (1963) General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae. North Carolina State College Contribution, 1502, i - vii, 1 - 585." title="General catalogue of the Homoptera, Fascicle VIII. Cicadoidea. Part 1. Cicadidae. Section I. Tibiceninae" type="journal article" year="1963">Metcalf 1963</bibRefCitation>
;
<bibRefCitation id="0F001A92350D2175FF4FFCFD77356D47" author="Sanborn, A. F." box="[221,381,869,894]" firstAuthor="Sanborn" journalOrPublisher="Florida Entomologist" pageId="19" pageNumber="449" pagination="75 - 79" part="89" refId="ref13838" refString="Sanborn, A. F. (2006) New records for the cicada fauna from four Central American countries (Hemiptera: Cicadoidea: Cicadidae). Florida Entomologist, 89, 75 - 79. http: // dx. doi. org / 10.1653 / 0015 - 4040 (2006) 89 [75: NRFTCF] 2.0. CO; 2" title="New records for the cicada fauna from four Central American countries (Hemiptera: Cicadoidea: Cicadidae)" type="journal article" year="2006">Sanborn 2006</bibRefCitation>
;
<bibRefCitation id="0F001A92350D2175FE1BFCFE778C6D47" author="Sanborn, A. F." bookContentInfo="1001 pp." box="[393,452,870,894]" firstAuthor="Sanborn" journalOrPublisher="Elsevier / Academic Press, San Diego" pageId="19" pageNumber="449" refId="ref14140" refString="Sanborn, A. F. (2013) Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet. Elsevier / Academic Press, San Diego, 1001 pp." title="Catalogue of the Cicadoidea (Hemiptera: Cicadoidea). With contributions to the bibliography by Martin H. Villet" type="book" year="2013">2013</bibRefCitation>
;
<bibRefCitation id="0F001A92350D2175FE5DFCFE74426D47" author="Sanborn, A. F." box="[463,522,870,894]" firstAuthor="Sanborn" journalOrPublisher="Check List" pageId="19" pageNumber="449" pagination="246 - 252" part="10" refId="ref14181" refString="Sanborn, A. F. (2014) Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Costa Rica including new records for fourteen species. Check List, 10, 246 - 252. http: // dx. doi. org / 10.15560 / 10.2.246" title="Checklist of the cicadas (Insecta: Hemiptera: Cicadidae) of Costa Rica including new records for fourteen species" type="journal article" year="2014">2014</bibRefCitation>
;
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<bibRefCitation id="0F001A92350D2175FCB0FCFE75A96D47" author="Maes, J. - M. &amp; Moulds, M. &amp; Sanborn, A. F." box="[802,993,869,894]" etAl="et al." firstAuthor="Maes" journalOrPublisher="Revista Nicaraguense Entomologia" pageId="19" pageNumber="449" pagination="1 - 138" part="72" refId="ref13472" refString="Maes, J. - M., Moulds, M. &amp; Sanborn, A. F. (2012) Cicadidae (Homoptera) de Nicaragua: catalogo ilustrado, incluyendo especies exoticas del Museo Entomologico de Leon. Revista Nicaraguense Entomologia, 72 (Suplemento 2), 1 - 138." title="Cicadidae (Homoptera) de Nicaragua: catalogo ilustrado, incluyendo especies exoticas del Museo Entomologico de Leon" type="journal article" year="2012">
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).
</paragraph>
</subSubSection>
</treatment>
</document>
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