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<mods:title id="F0D818F59BD016B3463532027EF03BE8">Recent thecideide brachiopods (Thecideida, Thecideoidea) from northern Sulawesi (Indonesia) with discovery of a new Thecidellina species (Thecidellinidae)</mods:title>
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<treatment id="C4568788BB646132FF50F3E312C6FA85" ID-DOI="http://doi.org/10.5281/zenodo.5986336" ID-GBIF-Taxon="154507805" ID-Zenodo-Dep="5986336" LSID="urn:lsid:plazi:treatment:C4568788BB646132FF50F3E312C6FA85" httpUri="http://treatment.plazi.org/id/C4568788BB646132FF50F3E312C6FA85" lastPageId="12" lastPageNumber="493" pageId="11" pageNumber="492">
<subSubSection id="04E56515BB646135FF50F3E3132AFB0C" box="[151,783,1196,1221]" pageId="11" pageNumber="492" type="nomenclature">
<paragraph id="4C40369EBB646135FF50F3E3132AFB0C" blockId="11.[151,783,1196,1221]" box="[151,783,1196,1221]" pageId="11" pageNumber="492">
<heading id="170881F2BB646135FF50F3E3132AFB0C" bold="true" box="[151,783,1196,1221]" fontSize="10" level="2" pageId="11" pageNumber="492" reason="2">
<emphasis id="7E8BEA8CBB646135FF50F3E3132AFB0C" bold="true" box="[151,783,1196,1221]" pageId="11" pageNumber="492">
Comparison with
<emphasis id="7E8BEA8CBB646135FEAAF3E31279FB0C" bold="true" box="[365,604,1196,1221]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FEAAF3E311DBFB0C" ID-CoL="87KZF" authorityName="Thomson" authorityYear="1915" box="[365,510,1196,1221]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
(
<bibRefCitation id="286E4B6FBB646135FDACF3E31323FB0C" author="Hedley, C." box="[619,774,1196,1221]" pageId="11" pageNumber="492" pagination="491 - 510" refId="ref14930" refString="Hedley, C. (1899) The Mollusca of Funafuti. Part II. Pelecypoda and Brachiopoda. Memoirs of the Australian Museum, 3, 491 - 510. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1899.504" type="journal article" year="1899">Hedley, 1899</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="04E56515BB646132FF37F3AF1164FEF6" lastPageId="12" lastPageNumber="493" pageId="11" pageNumber="492" type="description">
<paragraph id="4C40369EBB646135FF37F3AF1313FB3F" blockId="11.[159,1014,1248,1683]" box="[240,822,1248,1270]" pageId="11" pageNumber="492">
<bibRefCitation id="286E4B6FBB646135FF37F3AF1107FB3F" author="Hedley, C." box="[240,290,1248,1270]" pageId="11" pageNumber="492" pagination="491 - 510" refId="ref14930" refString="Hedley, C. (1899) The Mollusca of Funafuti. Part II. Pelecypoda and Brachiopoda. Memoirs of the Australian Museum, 3, 491 - 510. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1899.504" type="journal article" year="1899">1899</bibRefCitation>
<emphasis id="7E8BEA8CBB646135FE96F3AF1225FB3F" box="[337,512,1248,1270]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F3AF118BFB3F" authorityName="Defrance" authorityYear="1822" box="[337,430,1248,1270]" class="Rhynchonellata" family="Thecideidae" genus="Thecidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidea</taxonomicName>
maxilla
</emphasis>
: Hedley, pp. 508510, fig. 57.
</paragraph>
<paragraph id="4C40369EBB646135FF37F246128BFAD6" blockId="11.[159,1014,1248,1683]" box="[240,686,1289,1311]" pageId="11" pageNumber="492">
1920
<emphasis id="7E8BEA8CBB646135FE96F2461207FAD6" box="[337,546,1289,1311]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F24611F5FAD6" authorityName="Thomson" authorityYear="1915" box="[337,464,1289,1311]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Dall, p. 283.
</paragraph>
<paragraph id="4C40369EBB646135FF37F27D12C7FA81" blockId="11.[159,1014,1248,1683]" box="[240,738,1330,1352]" pageId="11" pageNumber="492">
1927
<emphasis id="7E8BEA8CBB646135FE96F27D1207FA81" box="[337,546,1330,1352]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F27D11F5FA81" authorityName="Thomson" authorityYear="1915" box="[337,464,1330,1352]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Thomson, p. 140.
</paragraph>
<paragraph id="4C40369EBB646135FF37F21313E6FABB" blockId="11.[159,1014,1248,1683]" box="[240,963,1372,1394]" pageId="11" pageNumber="492">
1954
<emphasis id="7E8BEA8CBB646135FE96F2131207FABB" box="[337,546,1372,1394]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F21311F5FABB" authorityName="Thomson" authorityYear="1915" box="[337,464,1372,1394]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Cooper, pp. 317318, pl. 81, figs. 110.
</paragraph>
<paragraph id="4C40369EBB646135FF37F2CA12E2FA52" blockId="11.[159,1014,1248,1683]" box="[240,711,1413,1435]" pageId="11" pageNumber="492">
1985
<emphasis id="7E8BEA8CBB646135FE96F2CA1207FA52" box="[337,546,1413,1435]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F2CA11F5FA52" authorityName="Thomson" authorityYear="1915" box="[337,464,1413,1435]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Zezina, p. 208.
</paragraph>
<paragraph id="4C40369EBB646135FF58F2FF134BFA0D" blockId="11.[159,1014,1248,1683]" box="[159,878,1454,1477]" pageId="11" pageNumber="492">
<emphasis id="7E8BEA8CBB646135FF58F2FF10E0FA0C" box="[159,197,1456,1477]" italics="true" pageId="11" pageNumber="492">non</emphasis>
1987
<emphasis id="7E8BEA8CBB646135FE96F2E11207FA0D" box="[337,546,1454,1476]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F2E111F5FA0D" authorityName="Thomson" authorityYear="1915" box="[337,464,1454,1476]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: DHondt, p. 41, pl. 4, figs. 15.
</paragraph>
<paragraph id="4C40369EBB646135FF58F29713A1FA27" blockId="11.[159,1014,1248,1683]" box="[159,900,1496,1518]" pageId="11" pageNumber="492">
? 1997
<emphasis id="7E8BEA8CBB646135FE96F2971207FA27" box="[337,546,1496,1518]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F29711F5FA27" authorityName="Thomson" authorityYear="1915" box="[337,464,1496,1518]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Laurin, pp. 453454, fig. 40AB.
</paragraph>
<paragraph id="4C40369EBB646135FF58F14D13D3F9DE" blockId="11.[159,1014,1248,1683]" box="[159,1014,1537,1559]" pageId="11" pageNumber="492">
<emphasis id="7E8BEA8CBB646135FF58F14D109CF9DE" box="[159,185,1538,1559]" italics="true" pageId="11" pageNumber="492">pp</emphasis>
2007
<emphasis id="7E8BEA8CBB646135FE96F14E1207F9DE" box="[337,546,1537,1559]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F14E11F5F9DE" authorityName="Thomson" authorityYear="1915" box="[337,464,1537,1559]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Bitner, p. 498, fig. 3AE, 3GH,
<emphasis id="7E8BEA8CBB646135FC46F14D1382F9DE" box="[897,935,1538,1559]" italics="true" pageId="11" pageNumber="492">non</emphasis>
fig.
<specimenCount id="5AF9FD17BB646135FC11F14E13D3F9DE" box="[982,1014,1537,1559]" pageId="11" pageNumber="492" type="generic">3F.</specimenCount>
</paragraph>
<paragraph id="4C40369EBB646135FF58F164137BF989" blockId="11.[159,1014,1248,1683]" box="[159,862,1578,1600]" pageId="11" pageNumber="492">
<emphasis id="7E8BEA8CBB646135FF58F16410E0F989" box="[159,197,1579,1600]" italics="true" pageId="11" pageNumber="492">non</emphasis>
2009
<emphasis id="7E8BEA8CBB646135FE96F1651207F989" box="[337,546,1578,1600]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F16511F5F989" authorityName="Thomson" authorityYear="1915" box="[337,464,1578,1600]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Bitner, pp. 1718, fig. 12AJ.
</paragraph>
<paragraph id="4C40369EBB646135FF58F11A13C0F9A3" blockId="11.[159,1014,1248,1683]" box="[159,997,1620,1642]" pageId="11" pageNumber="492">
<emphasis id="7E8BEA8CBB646135FF58F11A109CF9A3" box="[159,185,1621,1642]" italics="true" pageId="11" pageNumber="492">pp</emphasis>
2014
<emphasis id="7E8BEA8CBB646135FE96F11B1207F9A3" box="[337,546,1620,1642]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F11B11F5F9A3" authorityName="Thomson" authorityYear="1915" box="[337,464,1620,1642]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Bitner, p. 259, fig. 11FI (
<emphasis id="7E8BEA8CBB646135FCFEF11A137AF9A3" box="[825,863,1621,1642]" italics="true" pageId="11" pageNumber="492">non</emphasis>
fig. 11DE).
</paragraph>
<paragraph id="4C40369EBB646135FF58F1311301F95A" blockId="11.[159,1014,1248,1683]" box="[159,804,1661,1683]" pageId="11" pageNumber="492">
<emphasis id="7E8BEA8CBB646135FF58F13110E0F95A" box="[159,197,1662,1683]" italics="true" pageId="11" pageNumber="492">non</emphasis>
2015
<emphasis id="7E8BEA8CBB646135FE96F1321207F95A" box="[337,546,1661,1683]" italics="true" pageId="11" pageNumber="492">
<taxonomicName id="8BFF4D1DBB646135FE96F13211F5F95A" authorityName="Thomson" authorityYear="1915" box="[337,464,1661,1683]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="genus">Thecidellina</taxonomicName>
maxilla
</emphasis>
: Bitner, p. 42, fig. 5AE.
</paragraph>
<paragraph id="4C40369EBB646135FF00F1931174F8F2" blockId="11.[151,1437,1754,2031]" pageId="11" pageNumber="492">
<emphasis id="7E8BEA8CBB646135FF00F193111FF93A" box="[199,314,1754,1779]" italics="true" pageId="11" pageNumber="492">T. maxilla</emphasis>
is a large species with horse-shoe shaped dorsal valve with a peripheral ridge smooth to finely granulate.
<taxonomicName id="8BFF4D1DBB646135FECBF1B01180F8DE" authority="Simon &amp; Lüter &amp; Logan &amp; Mottequin, 2018" authorityName="Simon &amp; Lüter &amp; Logan &amp; Mottequin" authorityYear="2018" box="[268,421,1790,1815]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="species" species="mawaliana" status="sp. nov.">
<emphasis id="7E8BEA8CBB646135FECBF1B01180F8DE" box="[268,421,1790,1815]" italics="true" pageId="11" pageNumber="492">T. mawaliana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65B857F7BB646135FE6CF1B11221F8DE" box="[427,516,1790,1815]" pageId="11" pageNumber="492" rank="species">
<emphasis id="7E8BEA8CBB646135FE6CF1B11221F8DE" bold="true" box="[427,516,1790,1815]" pageId="11" pageNumber="492">sp. nov.</emphasis>
</taxonomicNameLabel>
is a rather small species drop-like shaped with a peripheral ridge ornamented with strong tubercles.
</paragraph>
<paragraph id="4C40369EBB646135FF00F00814B2F802" blockId="11.[151,1437,1754,2031]" pageId="11" pageNumber="492">
In
<emphasis id="7E8BEA8CBB646135FF21F0081172F896" box="[230,343,1862,1887]" italics="true" pageId="11" pageNumber="492">T. maxilla</emphasis>
the brachial bridge is relatively narrower (
<bibRefCitation id="286E4B6FBB646135FCFBF009140FF896" author="Hedley, C." box="[828,1066,1862,1887]" pageId="11" pageNumber="492" pagination="491 - 510" refId="ref14930" refString="Hedley, C. (1899) The Mollusca of Funafuti. Part II. Pelecypoda and Brachiopoda. Memoirs of the Australian Museum, 3, 491 - 510. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1899.504" type="journal article" year="1899">Hedley 1899, fig. 57</bibRefCitation>
) and the median septum is long, with lateral flanges developed along the inner sides of the brachial cavities (
<bibRefCitation id="286E4B6FBB646135FC30F02514A8F84A" author="Hedley, C." box="[1015,1165,1898,1923]" pageId="11" pageNumber="492" pagination="491 - 510" refId="ref14930" refString="Hedley, C. (1899) The Mollusca of Funafuti. Part II. Pelecypoda and Brachiopoda. Memoirs of the Australian Museum, 3, 491 - 510. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1899.504" type="journal article" year="1899">Hedley 1899</bibRefCitation>
, fig. 57, p. 509;
<bibRefCitation id="286E4B6FBB646135FA8DF02410F7F86E" author="Cooper, G. A." pageId="11" pageNumber="492" pagination="315 - 318" refId="ref14248" refString="Cooper, G. A. (1954) Recent brachiopods in Bikini and nearby atolls, Marshall Islands. United States Geological Survey Professional Paper, 260 G, 315 - 318. https: // doi. org / 10.3133 / pp 260 g" type="journal article" year="1954">Cooper 1954</bibRefCitation>
, pl. 81, fig. 10). The brachial cavities are more elongate. The single massive spicules of the intrabrachial ridge which are not completely covering the brachial cavities are rather regular in shape and size.
</paragraph>
<paragraph id="4C40369EBB646132FF00F0981196FF1D" blockId="11.[151,1437,1754,2031]" lastBlockId="12.[151,1437,151,1356]" lastPageId="12" lastPageNumber="493" pageId="11" pageNumber="492">
In
<taxonomicName id="8BFF4D1DBB646135FF21F09811A5F826" authority="Simon &amp; Lüter &amp; Logan &amp; Mottequin, 2018" authorityName="Simon &amp; Lüter &amp; Logan &amp; Mottequin" authorityYear="2018" box="[230,384,2006,2031]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="11" pageNumber="492" phylum="Brachiopoda" rank="species" species="mawaliana" status="sp. nov.">
<emphasis id="7E8BEA8CBB646135FF21F09811A5F826" box="[230,384,2006,2031]" italics="true" pageId="11" pageNumber="492">T. mawaliana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65B857F7BB646135FE40F09911C4F826" box="[391,481,2006,2031]" pageId="11" pageNumber="492" rank="species">
<emphasis id="7E8BEA8CBB646135FE40F09911C4F826" bold="true" box="[391,481,2006,2031]" pageId="11" pageNumber="492">sp. nov.</emphasis>
</taxonomicNameLabel>
the brachial bridge is wider and the median septum is devoid of lateral flanges. The brachial cavities are oval. The spicules of the brachial cavities, forming complete canopies are thin, irregular in shape and fused together.
</paragraph>
<paragraph id="4C40369EBB636132FF00F7901164FEF6" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
The ventral valve floor in
<emphasis id="7E8BEA8CBB636132FE31F7AF1242FF31" box="[502,615,223,248]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
is not roughly tuberculate (nearly smooth), the planodeltidium is larger and quite concave dorsally (
<bibRefCitation id="286E4B6FBB636132FE10F64C1311FED5" author="Cooper, G. A." box="[471,820,259,284]" pageId="12" pageNumber="493" pagination="315 - 318" refId="ref14248" refString="Cooper, G. A. (1954) Recent brachiopods in Bikini and nearby atolls, Marshall Islands. United States Geological Survey Professional Paper, 260 G, 315 - 318. https: // doi. org / 10.3133 / pp 260 g" type="journal article" year="1954">Cooper 1954, pl. 81, figs. 56</bibRefCitation>
). The hemispondylium in
<emphasis id="7E8BEA8CBB636132FB98F64B14EBFED5" box="[1119,1230,259,284]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
is not attached to the valve floor.
</paragraph>
</subSubSection>
<subSubSection id="04E56515BB636132FF00F605144EFD25" pageId="12" pageNumber="493" type="discussion">
<paragraph id="4C40369EBB636132FF00F6051353FE07" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
<emphasis id="7E8BEA8CBB636132FF00F605111EFEAA" bold="true" box="[199,315,330,355]" pageId="12" pageNumber="493">Remarks.</emphasis>
<emphasis id="7E8BEA8CBB636132FE83F60311CBFEAA" box="[324,494,330,355]" italics="true" pageId="12" pageNumber="493">Non T. maxilla</emphasis>
in DHondt (1987). In this paper, this species is cited without description but the illustrated specimens are very distinct from the
<typeStatus id="9344883CBB636132FD68F62012FAFE4E" box="[687,735,367,391]" pageId="12" pageNumber="493">type</typeStatus>
of
<emphasis id="7E8BEA8CBB636132FCC2F6201350FE4E" box="[773,885,366,391]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
. Their median septum is extremely wide and the spicules, differently shaped, form complete canopies on the brachial cavities. This is possibly a new species and this material must be restudied to give accurate taxonomical data.
</paragraph>
<paragraph id="4C40369EBB636132FF00F695111FFDF3" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
?
<emphasis id="7E8BEA8CBB636132FF1DF695116CFE3B" box="[218,329,473,498]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
in
<bibRefCitation id="286E4B6FBB636132FEABF6961229FE3B" author="Laurin, B." box="[364,524,473,498]" pageId="12" pageNumber="493" pagination="411 - 471" refId="ref15260" refString="Laurin, B. (1997) Brachiopodes recoltes dans les eaux de la Nouvelle-Caledonie et des iles Loyaute, Matthew et Chesterfield. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, volume 18. Memoires du Museum national d'Histoire Naturelle, 176, 411 - 471." type="journal article" year="1997">Laurin (1997)</bibRefCitation>
. This identification remains unclear as no detailed description is provided and the illustrations in this paper are not informative. This material must be revisited to have accurate data for determination.
</paragraph>
<paragraph id="4C40369EBB636132FF00F509144EFD25" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
<emphasis id="7E8BEA8CBB636132FF00F509114AFD94" box="[199,367,580,605]" italics="true" pageId="12" pageNumber="493">Non T. maxilla</emphasis>
in
<bibRefCitation id="286E4B6FBB636132FE51F50B12C0FD94" author="Bitner, M. A." box="[406,741,580,605]" pageId="12" pageNumber="493" pagination="419 - 461" refId="ref14031" refString="Bitner, M. A. (2007) Recent brachiopods from the Austral Islands, French Polynesia, South-Central Pacific. Zoosystema, 29 (3), 419 - 461." type="journal article" year="2007">Bitner (2007, p. 498, fig. 3F)</bibRefCitation>
. This specimen is probably a representative of
<taxonomicName id="8BFF4D1DBB636132FAC7F5091116FD48" authority="Cooper, 1954" authorityName=": Cooper" authorityYear="1954" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="12" pageNumber="493" phylum="Brachiopoda" rank="species" species="congregata">
<emphasis id="7E8BEA8CBB636132FAC7F50915BEFD94" box="[1280,1435,582,605]" italics="true" pageId="12" pageNumber="493">T. congregata</emphasis>
<bibRefCitation id="286E4B6FBB636132FF50F5261116FD48" author="Cooper, G. A." box="[151,307,616,641]" pageId="12" pageNumber="493" pagination="315 - 318" refId="ref14248" refString="Cooper, G. A. (1954) Recent brachiopods in Bikini and nearby atolls, Marshall Islands. United States Geological Survey Professional Paper, 260 G, 315 - 318. https: // doi. org / 10.3133 / pp 260 g" type="journal article" year="1954">Cooper, 1954</bibRefCitation>
</taxonomicName>
. Canopying spicules are different from the
<typeStatus id="9344883CBB636132FCE5F5261377FD48" box="[802,850,617,641]" pageId="12" pageNumber="493">type</typeStatus>
of
<emphasis id="7E8BEA8CBB636132FCBCF52613C9FD48" box="[891,1004,616,641]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
, being single and very massive in the anterior part of the brachial cavities and being extremely small and pointed in the posterior part. In
<emphasis id="7E8BEA8CBB636132FAE5F5C215B0FD6C" box="[1314,1429,652,677]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
, spicules have a much more regular shape. The right and left sides of the septum have no typical “curled flanges with serrate edges” as in
<emphasis id="7E8BEA8CBB636132FE68F59A123AFD25" box="[431,543,723,748]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
(
<bibRefCitation id="286E4B6FBB636132FDEAF59C12E7FD25" author="Hedley, C." box="[557,706,723,748]" pageId="12" pageNumber="493" pagination="491 - 510" refId="ref14930" refString="Hedley, C. (1899) The Mollusca of Funafuti. Part II. Pelecypoda and Brachiopoda. Memoirs of the Australian Museum, 3, 491 - 510. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1899.504" type="journal article" year="1899">Hedley 1899</bibRefCitation>
, fig. 57;
<bibRefCitation id="286E4B6FBB636132FCE2F59B139CFD25" author="Cooper, G. A." box="[805,953,723,748]" pageId="12" pageNumber="493" pagination="315 - 318" refId="ref14248" refString="Cooper, G. A. (1954) Recent brachiopods in Bikini and nearby atolls, Marshall Islands. United States Geological Survey Professional Paper, 260 G, 315 - 318. https: // doi. org / 10.3133 / pp 260 g" type="journal article" year="1954">Cooper 1954</bibRefCitation>
, pl. 81, fig. 10).
</paragraph>
</subSubSection>
<subSubSection id="04E56515BB636132FF00F5B712C6FA85" pageId="12" pageNumber="493" type="description">
<paragraph id="4C40369EBB636132FF00F5B711A2FC2F" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
<emphasis id="7E8BEA8CBB636132FF00F5B7114EFCD9" box="[199,363,759,784]" italics="true" pageId="12" pageNumber="493">Non T. maxilla</emphasis>
in
<bibRefCitation id="286E4B6FBB636132FE49F5B8130EFCD9" author="Bitner, M. A." box="[398,811,759,784]" pageId="12" pageNumber="493" pagination="1 - 39" refId="ref14064" refString="Bitner, M. A. (2009) Recent Brachiopoda from the Norfolk Ridge, New Caledonia, with description of four new species. Zootaxa, 2235, 1 - 39." type="journal article" year="2009">Bitner (2009, pp. 1718; fig. 12AJ)</bibRefCitation>
. These specimens display a peripheral ridge with strong tubercles and a median septum without lateral flanges. In
<emphasis id="7E8BEA8CBB636132FCE6F45313B5FCFD" box="[801,912,795,820]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
the peripheral ridge is nearly smooth or finely granulated. The space between the brachial bridge and intrabrachial ridge is larger than in
<emphasis id="7E8BEA8CBB636132FB5CF40F152EFC9E" box="[1179,1291,830,855]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
. The ventral valve is not emarginate as in the
<typeStatus id="9344883CBB636132FDD8F42C126AFCB2" box="[543,591,867,891]" pageId="12" pageNumber="493">type</typeStatus>
specimen and has a rough valve floor whereas it is nearly smooth in
<emphasis id="7E8BEA8CBB636132FA40F42B10CEFC56" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
. The cardinal process and socket ridges are much stronger than in the
<typeStatus id="9344883CBB636132FBC2F4C81410FC56" box="[1029,1077,903,927]" pageId="12" pageNumber="493">type</typeStatus>
population of
<emphasis id="7E8BEA8CBB636132FB1CF4C8156EFC56" box="[1243,1355,902,927]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
. These specimens could be representatives of another species derived from the same group as
<emphasis id="7E8BEA8CBB636132FB53F4E4152DFC0A" box="[1172,1288,938,963]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
but it needs genetic confirmation.
</paragraph>
<paragraph id="4C40369EBB636132FF00F4BD1533FB74" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
<emphasis id="7E8BEA8CBB636132FF00F4BD1155FBC3" box="[199,368,1009,1034]" italics="true" pageId="12" pageNumber="493">Non T. maxilla</emphasis>
in
<bibRefCitation id="286E4B6FBB636132FE5FF4BE133EFBC3" author="Bitner, M. A." box="[408,795,1009,1034]" pageId="12" pageNumber="493" pagination="245 - 265" refId="ref14151" refString="Bitner, M. A. (2014) Living brachiopods from French Polynesia, Central Pacific, with descriptions of two new species. Pacific Science, 68 (2), 245 - 265. https: // doi. org / 10.2984 / 68.2.6" type="journal article" year="2014">Bitner (2014, p. 259, fig. 11DE)</bibRefCitation>
. These ventral valves have a floor with extremely thick tubercles whereas this surface in the
<emphasis id="7E8BEA8CBB636132FD83F3591292FBE7" box="[580,695,1045,1070]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
is smooth or finely granulate. The gonad pits are situated more posteriorly and they are much smaller than in
<emphasis id="7E8BEA8CBB636132FD6AF375133AFB9B" box="[685,799,1081,1106]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
or in
<taxonomicName id="8BFF4D1DBB636132FCAFF3751426FB9B" authority="Simon &amp; Lüter &amp; Logan &amp; Mottequin, 2018" authorityName="Simon &amp; Lüter &amp; Logan &amp; Mottequin" authorityYear="2018" box="[872,1027,1081,1106]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="12" pageNumber="493" phylum="Brachiopoda" rank="species" species="mawaliana" status="sp. nov.">
<emphasis id="7E8BEA8CBB636132FCAFF3751426FB9B" box="[872,1027,1081,1106]" italics="true" pageId="12" pageNumber="493">T. mawaliana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65B857F7BB636132FBCBF376144DFB9B" box="[1036,1128,1081,1106]" pageId="12" pageNumber="493" rank="species">
<emphasis id="7E8BEA8CBB636132FBCBF376144DFB9B" bold="true" box="[1036,1128,1081,1106]" pageId="12" pageNumber="493">sp. nov.</emphasis>
</taxonomicNameLabel>
Moreover, it has a narrow median groove devoid of tubercles (not a ridge), a character not seen in
<emphasis id="7E8BEA8CBB636132FC0FF311141CFBBC" box="[968,1081,1116,1141]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
. They are also not similar to
<taxonomicName id="8BFF4D1DBB636132FA40F3111131FB50" authority="Simon &amp; Lüter &amp; Logan &amp; Mottequin, 2018" authorityName="Simon &amp; Lüter &amp; Logan &amp; Mottequin" authorityYear="2018" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="12" pageNumber="493" phylum="Brachiopoda" rank="species" species="mawaliana" status="sp. nov.">
<emphasis id="7E8BEA8CBB636132FA40F3111131FB50" italics="true" pageId="12" pageNumber="493">T. mawaliana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65B857F7BB636132FEDBF3CF115DFB50" box="[284,376,1152,1177]" pageId="12" pageNumber="493" rank="species">
<emphasis id="7E8BEA8CBB636132FEDBF3CF115DFB50" bold="true" box="[284,376,1152,1177]" pageId="12" pageNumber="493">sp. nov.</emphasis>
</taxonomicNameLabel>
: in the new species, the valve floor displays much smaller tubercles, the hemispondylium is attached to the bottom of the valve floor for only a short part of its length and the gonad pits are larger.
</paragraph>
<paragraph id="4C40369EBB636132FF00F38612C6FA85" blockId="12.[151,1437,151,1356]" pageId="12" pageNumber="493">
<emphasis id="7E8BEA8CBB636132FF00F3861149FB29" box="[199,364,1223,1248]" italics="true" pageId="12" pageNumber="493">Non T. maxilla</emphasis>
in
<bibRefCitation id="286E4B6FBB636132FE48F3881282FB29" author="Bitner, M. A." box="[399,679,1223,1248]" pageId="12" pageNumber="493" pagination="33 - 50" refId="ref14198" refString="Bitner, M. A. (2015) Checklist of recent brachiopod species collected during the Terrasses and Exbodi cruises in the New Caledonian region, SW Pacific. ZooKeys, 537, 33 - 50. https: // doi. org / 10.3897 / zookeys. 537.6567" type="journal article" year="2015">Bitner (2015, fig. 5AE)</bibRefCitation>
. Canopying spicules are different from the
<typeStatus id="9344883CBB636132FB4CF386149EFB28" box="[1163,1211,1225,1249]" pageId="12" pageNumber="493">type</typeStatus>
of
<emphasis id="7E8BEA8CBB636132FB27F3861575FB29" box="[1248,1360,1223,1248]" italics="true" pageId="12" pageNumber="493">T. maxilla</emphasis>
, being single and very massive in the anterior part of the brachial cavities and being extremely small and pointed in the posterior part. The typical “curled flanges with serrate edges” of the median septum are missing. These specimens are representatives of
<taxonomicName id="8BFF4D1DBB636132FE4AF27B120DFA82" authorityName=": Cooper" authorityYear="1954" box="[397,552,1332,1355]" class="Rhynchonellata" family="Thecidellinidae" genus="Thecidellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thecideida" pageId="12" pageNumber="493" phylum="Brachiopoda" rank="species" species="congregata">
<emphasis id="7E8BEA8CBB636132FE4AF27B120DFA82" box="[397,552,1332,1355]" italics="true" pageId="12" pageNumber="493">T. congregata</emphasis>
</taxonomicName>
(see hereunder).
</paragraph>
</subSubSection>
</treatment>
</document>