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<document id="3431876297F73EAA7BCA80C0F2BC709F" ID-DOI="10.3897/zookeys.1210.123986" ID-ZooBank="6972216E-2356-4DF7-BD24-2B6B68E639B4" ID-publisher-id="123986" URI-arpha="B6184626-6862-59C4-9047-EF9B7AC24E35" URI-zoobank="http://zoobank.org/6972216E-2356-4DF7-BD24-2B6B68E639B4" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1723830749198" checkinUser="pensoft" docAuthor="Xu, Xin, Zhan, Yi, Aung, Khin Pyae Pyae, Liu, Fengxiang, Yu, Long &amp; Li, Daiqin" docDate="2024" docId="BE9AF2F05BBA5A87B548EEDB4256870E" docLanguage="en" docName="ZooKeys 1210: 99-115" docOrigin="ZooKeys 1210" docSource="https://zookeys.pensoft.net/article/123986/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Liphistius rostratus Zhan &amp; Xu 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="115" masterDocId="B6184626686259C49047EF9B7AC24E35" masterDocTitle="Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar" masterLastPageNumber="115" masterPageNumber="99" pageNumber="99" updateTime="1723832703504" updateUser="admin">
<mods:mods id="A3E49628CE4CD01C46CDF602BB334872" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="73F757A674A873B66985BDD7F60C940F">Four new species of the spider genus Liphistius (Araneae, Mesothelae, Liphistiidae, Liphistiinae) from Myanmar</mods:title>
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<mods:name id="871D267F6FA37BFD88AB30F5769B2AA4" type="personal">
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<mods:namePart id="3DAA57A666899666D48BEF750B441902">Xu, Xin</mods:namePart>
<mods:nameIdentifier id="C3D72520C61B759C6544C9255B912090" type="ORCID">0000-0001-5632-6622</mods:nameIdentifier>
<mods:affiliation id="63DB8CDD5F8BA1686D8F1516FD2507E5">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:namePart id="22174C8B61FBAE0572A7D4810828FC8A">Zhan, Yi</mods:namePart>
<mods:affiliation id="EC1F13E4D5B2B39639D91D5F595B3C83">College of Life Sciences, Hunan Normal University, 36 Lushan Road, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="070FDAF8B5B1898AB6816E4C5A0F3702">Aung, Khin Pyae Pyae</mods:namePart>
<mods:affiliation id="D830C15AFE24D53BA9B5DD1FD9ED0A9D">Department of Zoology, University of Yangon, Kamayut Township, Pyay Road, Yangon, 11041, Myanmar &amp; Department of Biology, Taungoo Education Degree College, Taungoo, 08101, Myanmar</mods:affiliation>
</mods:name>
<mods:name id="F6A39FF57219E6F71373938BEC8E9987" type="personal">
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</mods:role>
<mods:namePart id="4A083BC1216DBB06A5D0AEBC828793F7">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="C8397D0918CC7C5430685EED5C2CF618">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
</mods:name>
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<mods:namePart id="E2A13D1F9D634B76EC453EEF6C47781C">Yu, Long</mods:namePart>
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<mods:affiliation id="6260CEAA65EC4D630AA9BD445DC94536">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China</mods:affiliation>
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<mods:namePart id="7C6F04D0EC417F5AC313A9A18C75EEA9">Li, Daiqin</mods:namePart>
<mods:nameIdentifier id="FD009D50B365D69E1EAEC125D5C8D547" type="ORCID">0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation id="1B6AB3CC2F3801945CE327FA15AC51B1">State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan 430062, Hubei Province, China &amp; Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543</mods:affiliation>
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<mods:typeOfResource id="D6514313C0E743D362F2208EFDA5ABB4">text</mods:typeOfResource>
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<mods:date id="7638250CEA064B8606F78551AE5EB608">2024</mods:date>
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<mods:number id="F590F0753CF01986144B21FFE882F091">2024-08-16</mods:number>
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<treatment id="BE9AF2F05BBA5A87B548EEDB4256870E" ID-DOI="http://doi.org/10.5281/zenodo.13333484" ID-Zenodo-Dep="13333484" ID-arpha="BE9AF2F0-5BBA-5A87-B548-EEDB4256870E" ID-zoobank="https://zoobank.org/35E36D61-84CB-4C01-830F-BCFE53FBAA6E" LSID="urn:lsid:plazi:treatment:BE9AF2F05BBA5A87B548EEDB4256870E" LSID-ZBK="urn:lsid:zoobank.org:act:35E36D61-84CB-4C01-830F-BCFE53FBAA6E" httpUri="http://treatment.plazi.org/id/BE9AF2F05BBA5A87B548EEDB4256870E">
<subSubSection id="0145F8F88E6DA1E1558D672278BAC01D" type="nomenclature">
<paragraph id="262AFC0357A6F4368FBCEE1F125FDBB7">
<taxonomicName id="E8597CBC1292B2409900E2E3EE7C47D5" authority="Zhan &amp; Xu" authorityName="Zhan &amp; Xu" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="09EC6147A3D2562E225F29B9BD346948" italics="true">Liphistius rostratus</emphasis>
Zhan &amp; Xu
</taxonomicName>
<taxonomicNameLabel id="CB335354974F3FA422691FBCB2719DF2" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="311A172FCD5516F7D55B47E8AD657749" type="description">
<paragraph id="BD270E97EA6B21E53BDAF7D12AD3C2E3">
<figureCitation id="77611BDFE971920F33CD9C4238A7BA83" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">Figs 9</figureCitation>
,
<figureCitation id="EA2FE05A41A0AF18E590CCF09DE2F640" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vulva of Liphistius rostratus sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure10" httpUri="https://binary.pensoft.net/fig/1115136">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1KBG" type="type material">
<paragraph id="5559C4DD5D7B5F47CF46371F79E07F58">
<heading id="5DBECA201F1CEBFA7709FD2BC42EC9DE" reason="title">Type material.</heading>
</paragraph>
<paragraph id="815DA4A2B33723FD4D0CDA7AB43B8172">
<materialsCitation id="42AAFE8EA0C379F3CE00162A028F1D6B" collectingDate="2019-07-25" collectionCode="XUX" collectorName="D. Li" country="Myanmar" elevation="553" latitude="22.88" location="War Phyu Taung Vill." longLatPrecision="754" longitude="96.12" specimenCode="XUX-2019-038" specimenCount="1" specimenCount-male="1" stateProvince="Mandalay Region" typeStatus="holotype">
<emphasis id="CBF8BB9F608FC32E55E57B086EC5EF5D" bold="true" italics="true">
<typeStatus id="A7FBEB8069606D6FFA4390DB953A83B9" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="18643CDD17069BD6AEA450A8CACF5459" name="Myanmar">Myanmar</collectingCountry>
<specimenCount id="E4D36A634ABA1F69646DCBCE707584E6" count="1" type="male"></specimenCount>
,
<collectingRegion id="F0B7438BB556196C9ADE58665E480187" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
,
<location id="3E14B34CC73F492074D3C5F0F6D52622" LSID="urn:lsid:plazi:treatment:BE9AF2F05BBA5A87B548EEDB4256870E:3E14B34CC73F492074D3C5F0F6D52622" country="Myanmar" latitude="22.88" longLatPrecision="754" longitude="96.12" name="War Phyu Taung Vill." stateProvince="Mandalay Region">War Phyu Taung Vill.</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EILBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[96.120000,22.880000]}">
<geoCoordinate id="CB0FE2FF1669ABDFCB357CAE878797A4" degrees="22.88" direction="north" orientation="latitude" precision="555" value="22.88">22.88 ° N</geoCoordinate>
,
<geoCoordinate id="D37014C72A308503B5608047F3A79C48" degrees="96.12" direction="east" orientation="longitude" precision="555" value="96.12">96.12 ° E</geoCoordinate>
</named-content>
, alt.
<quantity id="E28B7169FE62F39A92FC18A21F689EFC" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">
<elevation id="002D392D7BB1ED6E1EEE1F20705BDDEE" metricMagnitude="2" metricUnit="m" metricValue="5.529999999999999" unit="m" value="553.0">553 m</elevation>
</quantity>
;
<date id="F82CAB061B07ADAC4009B99B6F14EA09" value="2019-07-25">
<collectingDate id="BD2D02078B30BA8E4E3BF1B07B33480A" value="2019-07-25">25. 07. 2019</collectingDate>
</date>
;
<collectorName id="9134E72FA719992C3F02B0B1F9B1EEB6">D. Li</collectorName>
et al. leg.;
<specimenCode id="EB28C391C46F707E5D607231D689C2F0">XUX-2019-038</specimenCode>
</materialsCitation>
.
<materialsCitation id="6044F475C40C9C2E9181C9CD819C233F" collectingDate="2019-07-25" collectionCode="XUX" collectorName="D. Li" country="Myanmar" elevation="553" latitude="22.88" location="War Phyu Taung Vill." longLatPrecision="754" longitude="96.12" specimenCode="XUX-2019-034" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Mandalay Region" typeStatus="paratype">
<emphasis id="3FA207C1E0A8C04308C09D253E0A8691" bold="true" italics="true">
<typeStatus id="3C4CA49C29155610F512511819E9A11B" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="ECF665EA5B1D05C6C7A96123315342CD" count="2" type="male">2 ♂♂</specimenCount>
<specimenCount id="8232C0E4DDB0F1CD5EF13D05C65B2B48" count="3" type="female">3 ♀♀</specimenCount>
, same data as for holotype;
<specimenCode id="CD643F3B9BF2A00F184507BCA27CAA4F">XUX-2019-034</specimenCode>
037 / 041
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQLBG" type="diagnosis">
<paragraph id="91CDBF6F21FAD11643F8D6145A2A45A4">
<heading id="20B55ABE211E805F1644DEA0F5B7E9C6" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FA021153CBB28AC9FF888FE0396F38C8">
The male of
<taxonomicName id="5EE438C63650F296A788D8EBE97DB57A" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="326FDEED752920213DC44168082F3F31" italics="true">L. rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EF3FD97276FA1CA5B2C609688B4EFAD3" rank="species">sp. nov.</taxonomicNameLabel>
resembles those of
<taxonomicName id="806EFDE3EF784101464218C6B884297E" authorityName="Schwendinger &amp; Huber" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="8B745EC089BF21D40A0788EC3620A681" italics="true">L. cupreus</emphasis>
</taxonomicName>
,
<taxonomicName id="B17090B61F98652838B2D3D50D6BE3DB" authorityName="Yu, Zhang &amp; Zhang" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="nabang">
<emphasis id="2298C0E783E79AE07CE56EDF535BD6FC" italics="true">L. nabang</emphasis>
</taxonomicName>
, and
<taxonomicName id="D783036F8103731304717CE8FF57FCF4" authorityName="Schwendinger &amp; Huber" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="platnicki">
<emphasis id="C2A8F1308FA34F6242B05AAB1F04640C" italics="true">L. platnicki</emphasis>
</taxonomicName>
in having distinctly elevated cumulus (
<abbrev id="ABBRID0ECNBG" xlink_title="cumulus">Cu</abbrev>
) (Fig.
<figureCitation id="4E1A1B43485D9D404C97FAF8B75ACD57" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 C</figureCitation>
), but it can be distinguished from them by the relatively larger paraembolic plate (PeP) (Fig.
<figureCitation id="B76081F6297AD279FCACCC0CE1589A36" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 A, E</figureCitation>
vs figs 21 A, 23 K in
<bibRefCitation id="E2C75F6D6D0679E2CB4BE737950037DF" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
; fig. 3 A in
<bibRefCitation id="CEB0280A19B688A0583B0B98F79BE71A" DOI="10.3969/j.issn.1005-9628.2021.01.006" author="Yu" etAl="et al." firstAuthor="Yu" refId="B10" refString="Yu K, Zhang F, Zhang JX (2021) First new species of the genus Liphistius Schiödte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30: 3640. https://doi.org/10.3969/j.issn.1005-9628.2021.01.006" year="2021">Yu et al. 2021</bibRefCitation>
), and an indistinct contrategular process (
<abbrev id="ABBRID0EWNBG" xlink_title="contrategular process">cp</abbrev>
) (Fig.
<figureCitation id="CCBD1A1031F5CBF1A1B8A09A88D036A0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B</figureCitation>
vs relatively large contrategular process, figs 21 E, 23 E in
<bibRefCitation id="D0F3F1DFD57EDEA751B6F1F6E5A81FA6" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
; fig. 3 B in
<bibRefCitation id="0DE6AD9633D08F12299E02222821F12F" DOI="10.3969/j.issn.1005-9628.2021.01.006" author="Yu" etAl="et al." firstAuthor="Yu" refId="B10" refString="Yu K, Zhang F, Zhang JX (2021) First new species of the genus Liphistius Schiödte, 1849 from China (Araneae: Liphistiidae). Acta Arachnologica Sinica 30: 3640. https://doi.org/10.3969/j.issn.1005-9628.2021.01.006" year="2021">Yu et al. 2021</bibRefCitation>
). The female of
<taxonomicName id="F0318219C914F1BCB24E470947000AED" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="7FA8BC96D2354B8BFFF62E13B3BCACB4" italics="true">L. rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0758C94BA8C2283E146ADBA70F364A58" rank="species">sp. nov.</taxonomicNameLabel>
resembles to that of
<taxonomicName id="C41C835EBC262ACDB92DE9F47F619683" authorityName="Schwendinger &amp; Huber" authorityYear="2022" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="cupreus">
<emphasis id="391714BD7BEC57302065ED389C3A5C35" italics="true">L. cupreus</emphasis>
</taxonomicName>
in having vesicle clusters along the anterior margin of the poreplate (
<abbrev id="ABBRID0E3OBG" xlink_title="sclerotized poreplate">PPl</abbrev>
), but it can be distinguished by the relatively longer posterior stalk (
<collectionCode id="450895A77A2275F4574CB105711F65BC">PS</collectionCode>
), with a length / width ratio 0.40.6 (Fig.
<figureCitation id="2C11D1F39D5BC7932BDF22C55DAB05F1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vulva of Liphistius rostratus sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure10" httpUri="https://binary.pensoft.net/fig/1115136">10</figureCitation>
vs length / width ratio about 0.20.4, fig.
<quantity id="DCBEA25F36C8526B3EBA95F6C94D7995" metricMagnitude="-1" metricUnit="m" metricValue="5.588" unit="in" value="22.0">22 in</quantity>
<bibRefCitation id="17F5A1ADA3061878F531AEAB8F61D438" DOI="10.35929/RSZ.0083" author="Schwendinger" etAl="et al." firstAuthor="Schwendinger" issue="2" journalOrPublisher="Revue Suisse de Zoologie" pagination="375-424" refId="B5" refString="Schwendinger PJ, Huber S, Lehmann-Graber C, Ono H, Aung MM, Hongpadharakiree K (2022) A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species. Revue Suisse de Zoologie 129 (2): 375424. https://doi.org/10.35929/RSZ.0083" title="A taxonomic revision of the Liphistius birmanicus - group (Araneae: Liphistiidae) with the description of five new species." volume="129" year="2022">Schwendinger et al. 2022</bibRefCitation>
).
</paragraph>
<caption id="C997D08E1B159160B1111B4367F6CB80" ID-DOI="10.3897/zookeys.1210.123986.figure9" ID-arpha="EE41BB75-A320-5356-97B2-C4FE897FAE94" httpUri="https://binary.pensoft.net/fig/1115135" startId="F9">
<paragraph id="0B8918B9F376B296EA199C5BA95B021C">
<label id="A74677FAF2EE11C49ED98FC00B948041">Figure 9.</label>
</paragraph>
<paragraph id="E4E7A17E2483CB868FB73476506F0EDB">
Male palp of
<taxonomicName id="C2BD0368F33DAB0CC5DCB577EA022CC2" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="5A93A17998703565FA74E33CAB344E46" italics="true">Liphistius rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EEF583394BB7FDEF2CE6D8F072D19351" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3A9CA78A8EA9C297E4DB53F04F4B1926" bold="true">A, E</emphasis>
prolateral view
<emphasis id="9B6B17F6047FF04203177E70EC9D8295" bold="true">B</emphasis>
ventral view
<emphasis id="568E02C705575F2866D9051E81F0BB98" bold="true">C, F</emphasis>
retrolateral view
<emphasis id="1EEB3159C4B6601AD0628B05921C0E49" bold="true">D</emphasis>
dorsal view
<emphasis id="E9B8CECB46EEBB67947497C325673E2F" bold="true">G</emphasis>
distal view, red arrow shows beak-like contrategular projection. Abbreviations:
<abbrev id="ABBRID0EFQBG" xlink_title="contrategulum">CT</abbrev>
= contrategulum;
<abbrev id="ABBRID0EJQBG" xlink_title="contrategular process">cp</abbrev>
= contrategular process;
<abbrev id="ABBRID0ENQBG" xlink_title="cumulus">Cu</abbrev>
= cumulus; E = embolus;
<abbrev id="ABBRID0ERQBG" xlink_title="paracymbium">PC</abbrev>
= paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum;
<abbrev id="ABBRID0EVQBG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
= tibial apophysis. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="5874242F0D3072C948265B27F6789200" ID-DOI="10.3897/zookeys.1210.123986.figure10" ID-arpha="CC0BDFD5-5D2E-5F2C-8AD4-7292A822B05C" httpUri="https://binary.pensoft.net/fig/1115136" startId="F10">
<paragraph id="C208B9D425864655700D5C5F10C19B72">
<label id="4811C0E7F56A72BED1E4D5CA07417F31">Figure 10.</label>
</paragraph>
<paragraph id="8331AD0BBE8B9D2E9AFE25EDABE9A38C">
Vulva of
<taxonomicName id="8F21898EC6B18F97129EFC8C00B880C6" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus" status="sp. nov.">
<emphasis id="78ACCF574389A7EA04C50AC7E663E281" italics="true">Liphistius rostratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0898230BBC5D53A55CD93726DA6A1F75" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="6187C14FC0B43D9414700CAF988C3C98" bold="true">A, C</emphasis>
dorsal view
<emphasis id="9D80BAE9E565F11389839EB7C457D5A8" bold="true">B, D</emphasis>
ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk;
<abbrev id="ABBRID0EVRBG" xlink_title="sclerotized poreplate">PPl</abbrev>
= poreplate;
<abbrev id="ABBRID0EZRBG" xlink_title="receptacular cluster">RC</abbrev>
= receptacular cluster. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ECSBG" type="description">
<paragraph id="F9882118EB337685EFEA08690C49D4F2">
<heading id="883A1ACE69E9534BEE4740E1852B23D1" reason="title">Description.</heading>
</paragraph>
<paragraph id="CE23517DE7E8169960F48501E2F16111">
<emphasis id="3A3AEF8E20E8D44AAC607CE574F7DFD0" bold="true">
Male
<typeStatus id="D7BF5644204985CFDF002223F9135C5F" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace yellowish brown, with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 26
<superScript id="D36AD013E6027708BEE4BA99B83B2CFD">th</superScript>
larger than others, 4
<superScript id="248A3F61E9222084A6B2E905A609018B">th</superScript>
largest; chelicerae robust, promargin of cheliceral groove with 12 denticles of variable size; labium and sternum yellowish brown, sternum with few short setae on anterior tip and many long setae on elongated posterior tip; legs yellowish brown, with strong setae and spines. Measurements:
<abbrev id="ABBRID0EOSBG" xlink_title="body length">BL</abbrev>
9.34,
<abbrev id="ABBRID0ESSBG" xlink_title="carapace length">CL</abbrev>
4.57,
<abbrev id="ABBRID0EWSBG" xlink_title="carapace width">CW</abbrev>
4.26,
<abbrev id="ABBRID0E1SBG" xlink_title="opisthosoma length">OL</abbrev>
5.02,
<abbrev id="ABBRID0E5SBG" xlink_title="opisthosoma width">OW</abbrev>
3.59; eye sizes and interdistances:
<abbrev id="ABBRID0ECTBG" xlink_title="anterior median eyes">AME</abbrev>
reduced,
<abbrev id="ABBRID0EGTBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.40,
<abbrev id="ABBRID0EKTBG" xlink_title="posterior median eyes">PME</abbrev>
reduced,
<abbrev id="ABBRID0EOTBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.24,
<abbrev id="ABBRID0ESTBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EWTBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.07,
<abbrev id="ABBRID0E1TBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0E5TBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.05,
<abbrev id="ABBRID0ECUBG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0EGUBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.30; labium 0.98 long and 0.48 wide; sternum 2.09 long and 0.83 wide; legs: I 12.63 (3.80, 1.32, 2.60, 3.07, 1.84), II 13.19 (3.80, 1.32, 2.60, 3.07, 1.84), III 14.35 (3.79, 1.58, 3.33, 3.44, 2.21), IV 19.76 (4.76, 1.66, 4.13, 6.01, 3.20).
</paragraph>
<paragraph id="D191393659F1496FACD56540BD510CEC">
Palp: tibial apophysis (
<abbrev id="ABBRID0EMUBG" xlink_title="male palps bearing tibial apophysis">TiA</abbrev>
) with 4 setae of similar length (Fig.
<figureCitation id="FF1E6D481C3B87FC8A6303E15D731FAC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 A C</figureCitation>
); paracymbium (
<abbrev id="ABBRID0EUUBG" xlink_title="paracymbium">PC</abbrev>
) with several setae situated on tip (Fig.
<figureCitation id="7C097F7273A1C825966B474F60897A40" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, C</figureCitation>
); cumulus (
<abbrev id="ABBRID0E3UBG" xlink_title="cumulus">Cu</abbrev>
) distinct elevated with 5 tapering spines (Fig.
<figureCitation id="714D27869EFE0A63388481EF36FC6C71" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 C</figureCitation>
); subtegulum (ST) without apophysis (Fig.
<figureCitation id="1A5986685701DDF1168BD26D037F6AE0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 F</figureCitation>
); contrategulum (
<abbrev id="ABBRID0EIVBG" xlink_title="contrategulum">CT</abbrev>
) with beak-like projection distally in distal view and small process in ventral view (Fig.
<figureCitation id="08E6A40F6B8C646A7FD7F2373472D156" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, G</figureCitation>
); tegulum (
<collectionCode id="F59DBCE55A561ABEE3F32EEAE4A84996" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
) semicircular with flat surface and serrated edges, and proximal tegular margin with an elevated transverse ridge (Fig.
<figureCitation id="D1C454F4665E54D898464D1BA7F320B7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, C, F</figureCitation>
); paraembolic plate (PeP) sclerotized, narrow (Fig.
<figureCitation id="5F3882DE8B0D654A0D33C9B87653B029" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 A, E</figureCitation>
); embolus (E) with 3 distinct longitudinal ridges reaching tip retrolaterally, and several denticles along these longitudinal ridges (Fig.
<figureCitation id="A4ABB38C28D297CA1E131C2A3ED6AC01" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male palp of Liphistius rostratus sp. nov. A, E prolateral view B ventral view C, F retrolateral view D dorsal view G distal view, red arrow shows beak-like contrategular projection. Abbreviations: CT = contrategulum; cp = contrategular process; Cu = cumulus; E = embolus; PC = paracymbium; PeP = paraembolic plate; ST = subtegulum; T = tegulum; TiA = tibial apophysis. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure9" httpUri="https://binary.pensoft.net/fig/1115135">9 B, F</figureCitation>
).
</paragraph>
<paragraph id="293BE238132A85EFC7869AD75DB4DC08">
<emphasis id="A9033D247D702D04B5FE4497DD818650" bold="true">
Female
<typeStatus id="1F980AEAE0707D0397515DE3B5AE9A91" type="paratype">paratype</typeStatus>
(
<specimenCode id="F26C952CE3818E156A9513AC8318CA0F">XUX- 2019-034</specimenCode>
).
</emphasis>
Carapace brown with few short, scattered bristles; opisthosoma brown, with 12 brown tergites, close to each other, 26
<superScript id="A2E8FFE5CC3DE05F5CA69360B180BE24">th</superScript>
larger than others, 5
<superScript id="A6E39A9204A4DD25FDA26DF5C2CD5511">th</superScript>
largest; chelicerae robust, reddish brown; promargin of cheliceral groove with 12 denticles of variable size; labium yellowish brown, sternum yellowish brown with several setae; legs with strong setae and spines. Measurements:
<abbrev id="ABBRID0EEWBG" xlink_title="body length">BL</abbrev>
10.54,
<abbrev id="ABBRID0EIWBG" xlink_title="carapace length">CL</abbrev>
4.5,
<abbrev id="ABBRID0EMWBG" xlink_title="carapace width">CW</abbrev>
3.93,
<abbrev id="ABBRID0EQWBG" xlink_title="opisthosoma length">OL</abbrev>
6.47,
<abbrev id="ABBRID0EUWBG" xlink_title="opisthosoma width">OW</abbrev>
4.74; eye sizes and interdistances:
<abbrev id="ABBRID0EYWBG" xlink_title="anterior median eyes">AME</abbrev>
0.05,
<abbrev id="ABBRID0E3WBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.37,
<abbrev id="ABBRID0EAXBG" xlink_title="posterior median eyes">PME</abbrev>
0.19,
<abbrev id="ABBRID0EEXBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.29,
<abbrev id="ABBRID0EIXBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EMXBG" xlink_title="anterior median eyes">AME</abbrev>
0.07,
<abbrev id="ABBRID0EQXBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EUXBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.12,
<abbrev id="ABBRID0EYXBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0E3XBG" xlink_title="posterior median eyes">PME</abbrev>
0.06,
<abbrev id="ABBRID0EAYBG" xlink_title="posterior median eyes">PME</abbrev>
<abbrev id="ABBRID0EEYBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.06,
<abbrev id="ABBRID0EIYBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EMYBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.09,
<abbrev id="ABBRID0EQYBG" xlink_title="anterior lateral eyes">ALE</abbrev>
<abbrev id="ABBRID0EUYBG" xlink_title="anterior lateral eyes">ALE</abbrev>
0.10,
<abbrev id="ABBRID0EYYBG" xlink_title="posterior lateral eyes">PLE</abbrev>
<abbrev id="ABBRID0E3YBG" xlink_title="posterior lateral eyes">PLE</abbrev>
0.32,
<abbrev id="ABBRID0EAZBG" xlink_title="anterior median eyes">AME</abbrev>
<abbrev id="ABBRID0EEZBG" xlink_title="posterior median eyes">PME</abbrev>
0.08; labium 0.95 long and 0.38 wide; sternum 2.41 long and 0.99 wide; palp 7.10 (2.44, 0.98, 1.86, 1.82), leg I 8.89 (2.94, 1.34, 2.00, 1.62, 0.99), II 9.69 (3.40, 1.09, 2.02, 1.98, 1.20), III 9.48 (2.29, 1.05, 2.59, 2.38, 1.17), IV 13.71 (3.21, 1.44, 3.11, 3.80, 2.21).
</paragraph>
<paragraph id="33580FF4BFA1C1C1D14C8E77933699A4">
Vulva: about rectangular poreplate (
<abbrev id="ABBRID0EKZBG" xlink_title="sclerotized poreplate">PPl</abbrev>
) with pair of anterior lobes; central dorsal opening (
<collectionCode id="905D1A264670DAA4400755019B26DA0F">CDO</collectionCode>
) located below center of poreplate (
<abbrev id="ABBRID0EOZBG" xlink_title="sclerotized poreplate">PPl</abbrev>
); racemose receptacular cluster (
<abbrev id="ABBRID0ESZBG" xlink_title="receptacular cluster">
<collectionCode id="235FC8924BD81EFE2297B3C9243BB828">RC</collectionCode>
</abbrev>
) small; posterior stalk (
<collectionCode id="03B36BD368534646B7262D7B23756475">PS</collectionCode>
) as wide as poreplate (
<abbrev id="ABBRID0EWZBG" xlink_title="sclerotized poreplate">PPl</abbrev>
), lateral margins of genital atrium (GA) slightly sclerotized (Fig.
<figureCitation id="A94D011792C7BE6AB12129E711F2D17D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Vulva of Liphistius rostratus sp. nov. A, C dorsal view B, D ventral view. Abbreviations: CDO = central dorsal opening; GA = genital atrium; PS = posterior stalk; PPl = poreplate; RC = receptacular cluster. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1210.123986.figure10" httpUri="https://binary.pensoft.net/fig/1115136">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5ZBG" type="variation">
<paragraph id="B6590AE953A64F095D8115141E43C726">
<heading id="912DC78384B05B2CDE02C031DA46A89C" reason="title">Variation.</heading>
</paragraph>
<paragraph id="7FF2073944B35C35DC7D414C8FBCE720">
Males (
<emphasis id="817B360FBD7EF3EF703834D6433B6453" italics="true">N</emphasis>
= 3):
<abbrev id="ABBRID0EG1BG" xlink_title="body length">BL</abbrev>
9.3410.31,
<abbrev id="ABBRID0EK1BG" xlink_title="carapace length">CL</abbrev>
4.575.24,
<abbrev id="ABBRID0EO1BG" xlink_title="carapace width">CW</abbrev>
4.265.83,
<abbrev id="ABBRID0ES1BG" xlink_title="opisthosoma length">OL</abbrev>
4.735.10,
<abbrev id="ABBRID0EW1BG" xlink_title="opisthosoma width">OW</abbrev>
3.403.40; females (
<emphasis id="52702B85236FB06A351F605BE28BF7E4" italics="true">N</emphasis>
= 4):
<abbrev id="ABBRID0E31BG" xlink_title="body length">BL</abbrev>
10.5412.35,
<abbrev id="ABBRID0EA2BG" xlink_title="carapace length">CL</abbrev>
4.506.31,
<abbrev id="ABBRID0EE2BG" xlink_title="carapace width">CW</abbrev>
3.935.47,
<abbrev id="ABBRID0EI2BG" xlink_title="opisthosoma length">OL</abbrev>
6.476.87,
<abbrev id="ABBRID0EM2BG" xlink_title="opisthosoma width">OW</abbrev>
4.644.79. The number of denticles on the cheliceral promargin varies from 12 to 13 (
<emphasis id="089289E164CCBD9ED51C231FD76CB1E3" italics="true">N</emphasis>
= 6). In addition, the male lacking an
<abbrev id="ABBRID0ES2BG" xlink_title="anterior median eyes">AME</abbrev>
and
<abbrev id="ABBRID0EW2BG" xlink_title="posterior median eyes">PME</abbrev>
, which may be degenerate during molting, is chosen as the
<typeStatus id="005E9B890AE75A15F04DAAF85D5DE9CC" type="holotype">holotype</typeStatus>
, because all
<typeStatus id="1684F557AB335E52DF1FE705B4814689" type="paratype">paratype</typeStatus>
male palps are deformed.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E12BG" type="etymology">
<paragraph id="295AED6F79390A21DAA2A02E5F4D8876">
<heading id="3778F46B2963AC25B42089628846C180" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="3968C4DF81B826F56791C1ECA0647ECD">
The species name is derived from the Latin word
<taxonomicName id="79903805B446A136631194A4653ADB56" authorityName="Xu &amp; Zhan &amp; Aung &amp; Liu &amp; Yu &amp; Li" authorityYear="2024" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="4C8DD574F58FCD9534F44EE8B3CA1A62" italics="true">rostratus</emphasis>
</taxonomicName>
”, referring to the beak-like contrategulum of the male palp in distal view.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK3BG" type="distribution">
<paragraph id="C4859CBF17DBAD7600019C4D24EF553D">
<heading id="3C6513F111C92CF0310A9F28FA73C9C9" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="DEAD99E9BC2F569366116E4CA1C1B3F8">
<collectingRegion id="B90F85A46ABE6D42BD1BC3BD739E8E33" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
(War Hpyu Taung),
<collectingCountry id="5F0FBAF34B9B2ADBA5B93D55D286CDFD" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>