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<mods:title id="1AD81B718819DFE145E43628E1E3F6EB">Revision of the Pachycondyla sikorae species-group (Hymenoptera: Formicidae) in Madagascar</mods:title>
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<subSubSection id="498244C6FFB7424AFF54469CFC12FC53" box="[151,951,943,969]" pageId="4" pageNumber="451" type="nomenclature">
<paragraph id="0127174DFFB7424AFF54469CFC12FC53" blockId="4.[151,951,943,969]" box="[151,951,943,969]" pageId="4" pageNumber="451">
<heading id="5A6FA021FFB7424AFF54469CFC12FC53" bold="true" box="[151,951,943,969]" fontSize="11" level="1" pageId="4" pageNumber="451" reason="1">
<emphasis id="33ECCB5FFFB7424AFF54469CFC12FC53" bold="true" box="[151,951,943,969]" pageId="4" pageNumber="451">
Synopsis of the taxonomic history of the
<taxonomicName id="C6986CCEFFB7424AFD53469CFCA1FC53" ID-CoL="4BQXY" ID-ENA="411771" authority="Fore, 1891" box="[656,772,943,969]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB7424AFD53469CFCA1FC53" bold="true" box="[656,772,943,969]" italics="true" pageId="4" pageNumber="451">P. sikorae</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="498244C6FFB7424AFF5446C8FC5FFB47" pageId="4" pageNumber="451" type="discussion">
<paragraph id="0127174DFFB7424AFF5446C8FAD6FB0D" blockId="4.[151,1436,1018,1176]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF5446C8FE36FB89" box="[151,403,1019,1043]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB7424AFF5446C8FE36FB89" box="[151,403,1019,1043]" italics="true" pageId="4" pageNumber="451">Pachycondyla sikorae</emphasis>
</taxonomicName>
was first described as a subgenus (
<taxonomicName id="C6986CCEFFB7424AFCFF46CFFC0FFB89" box="[828,938,1020,1043]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFCFF46CFFC0FFB89" box="[828,938,1020,1043]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
) of
<taxonomicName id="C6986CCEFFB7424AFC2046CFFB93FB89" box="[995,1078,1020,1043]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFC2046CFFB93FB89" box="[995,1078,1020,1043]" italics="true" pageId="4" pageNumber="451">Ponera</emphasis>
</taxonomicName>
by
<bibRefCitation id="65096ABCFFB7424AFBAB46C9FA82FB88" author="Forel, A." bookContentInfo="237 pp." box="[1128,1319,1018,1042]" editor="Grandidier, A." firstAuthor="Forel" journalOrPublisher="Hachette et Cie, Paris" page="127" pageId="4" pageNumber="451" refId="ref19650" refString="Forel, A. (1891) Les Formicides. [part]. In: Grandidier, A. 1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule). Hachette et Cie, Paris, v + 237 pp." title="Les Formicides. [part]" type="book" volumeTitle="1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule)" year="1891">Forel 1891: 127</bibRefCitation>
. The new subgenus
<taxonomicName id="C6986CCEFFB7424AFECA412EFEDDFBAE" box="[265,376,1053,1076]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFECA412EFEDDFBAE" box="[265,376,1053,1076]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
was later raised to genus (
<bibRefCitation id="65096ABCFFB7424AFD72412FFCCCFBAE" author="Emery, C." box="[689,873,1052,1076]" firstAuthor="Emery" journalOrPublisher="Annales de la Societe Entomologique de Belgique" page="46" pageId="4" pageNumber="451" pagination="32 - 54" part="45" refId="ref19541" refString="Emery, C. (1901) Notes sur les sous-familles des Dorylines et Ponerines (Famille des Formicides). Annales de la Societe Entomologique de Belgique, 45, 32 - 54." title="Notes sur les sous-familles des Dorylines et Ponerines (Famille des Formicides)" type="journal article" year="1901">Emery, 1901:46</bibRefCitation>
) and by 1911, included a heterogenous group of species placed in four subgenera:
<taxonomicName id="C6986CCEFFB7424AFDD8410DFC70FBCF" authority="Mesoponera, Brachyponera" authorityName="Mesoponera, Brachyponera" box="[539,981,1085,1109]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFDD8410DFC70FBCF" box="[539,981,1085,1109]" italics="true" pageId="4" pageNumber="451">Euponera, Mesoponera, Brachyponera</emphasis>
</taxonomicName>
, and
<taxonomicName id="C6986CCEFFB7424AFBD5410EFB61FBCF" box="[1046,1220,1085,1109]" class="Insecta" family="Formicidae" genus="Trachymesopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFBD5410EFB61FBCF" box="[1046,1220,1085,1109]" italics="true" pageId="4" pageNumber="451">Trachymesopus</emphasis>
</taxonomicName>
(
<bibRefCitation id="65096ABCFFB7424AFB16410EFF12FBEC" author="Emery, C." firstAuthor="Emery" journalOrPublisher="Genera Insectorum" page="79 - 86" pageId="4" pageNumber="451" pagination="1 - 125" part="118" refId="ref19624" refString="Emery, C. (1911) Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118, 1 - 125." title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae" type="journal article" year="1911">Emery 1911: 79 86</bibRefCitation>
). Brown in
<bibRefCitation id="65096ABCFFB7424AFE87416DFE43FBEC" author="Bolton, B." bookContentInfo="222 pp." box="[324,486,1118,1142]" firstAuthor="Bolton" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="4" pageNumber="451" refId="ref19129" refString="Bolton, B. (1994) Identification Guide to the Ant Genera of the World. Harvard University Press, Cambridge, Massachusetts, 222 pp." title="Identification Guide to the Ant Genera of the World" type="book" year="1994">Bolton (1994)</bibRefCitation>
synonymized the genus as a junior synonym of
<taxonomicName id="C6986CCEFFB7424AFBE2416DFB65FBEC" box="[1057,1216,1118,1142]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFBE2416DFB65FBEC" box="[1057,1216,1118,1142]" italics="true" pageId="4" pageNumber="451">Pachycondyla</emphasis>
</taxonomicName>
, which is where it remains today though molecular results suggest that
<taxonomicName id="C6986CCEFFB7424AFD2741B3FC26FB02" box="[740,899,1152,1176]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFD2741B3FC26FB02" box="[740,899,1152,1176]" italics="true" pageId="4" pageNumber="451">Pachycondyla</emphasis>
</taxonomicName>
is paraphyletic (
<bibRefCitation id="65096ABCFFB7424AFBF8414CFB1FFB0D" author="Ward, P. S." box="[1083,1210,1151,1175]" firstAuthor="Ward" journalOrPublisher="Myrmecological News" pageId="4" pageNumber="451" pagination="21 - 29" part="15" refId="ref19973" refString="Ward, P. S. (2011) Integrating molecular phylogenetic results into ant taxonomy (Hymenoptera: Formicidae). Myrmecological News, 15, 21 - 29." title="Integrating molecular phylogenetic results into ant taxonomy (Hymenoptera: Formicidae)" type="journal article" year="2011">Ward 2011</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFB05414CFAC2FB0D" author="Schmidt, C." box="[1222,1383,1151,1175]" firstAuthor="Schmidt" journalOrPublisher="Zootaxa" pageId="4" pageNumber="451" pagination="201" part="3647" refId="ref19931" refString="Schmidt, C. (2013) Molecular phylogenetics of ponerine ants (Hymenoptera: Formicidae: Ponerinae). Zootaxa, 3647, 201 - 250. http: // dx. doi. org / 10.11646 / zootaxa. 3647.2.1" title="Molecular phylogenetics of ponerine ants (Hymenoptera: Formicidae: Ponerinae)" type="journal article" year="2013">Schmidt 2013</bibRefCitation>
).
</paragraph>
<paragraph id="0127174DFFB7424AFF5441F6FC5FFB47" blockId="4.[151,1018,1221,1246]" box="[151,1018,1221,1246]" pageId="4" pageNumber="451">
<emphasis id="33ECCB5FFFB7424AFF5441F6FC54FB44" bold="true" box="[151,1009,1221,1246]" pageId="4" pageNumber="451">
Detailed taxonomic history of
<taxonomicName id="C6986CCEFFB7424AFE3D41F5FDD4FB47" box="[510,625,1222,1245]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFE3D41F5FDD4FB47" bold="true" box="[510,625,1222,1245]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
modified from AntCat.org 2013
</emphasis>
:
</paragraph>
</subSubSection>
<subSubSection id="498244C6FFB7424AFF54403DFB74F9B5" pageId="4" pageNumber="451" type="reference_group">
<paragraph id="0127174DFFB7424AFF54403DFB7FFABC" blockId="4.[151,1436,1293,1584]" box="[151,1242,1293,1318]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF54403DFE63FABF" authority="Forel, 1891: 126" authorityName="Forel" authorityPageNumber="126" authorityYear="1891" box="[151,454,1293,1317]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFF54403DFEA3FABF" box="[151,262,1294,1317]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
<bibRefCitation id="65096ABCFFB7424AFECF403EFE63FABF" author="Forel, A." bookContentInfo="237 pp." box="[268,454,1293,1317]" editor="Grandidier, A." firstAuthor="Forel" journalOrPublisher="Hachette et Cie, Paris" page="126" pageId="4" pageNumber="451" refId="ref19650" refString="Forel, A. (1891) Les Formicides. [part]. In: Grandidier, A. 1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule). Hachette et Cie, Paris, v + 237 pp." title="Les Formicides. [part]" type="book" volumeTitle="1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule)" year="1891">Forel, 1891: 126</bibRefCitation>
</taxonomicName>
[as subgenus of
<taxonomicName id="C6986CCEFFB7424AFD41403DFD71FABF" ID-CoL="63NZ7" authority=" Latreille, 1804" box="[642,724,1294,1317]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFD41403DFD71FABF" box="[642,724,1294,1317]" italics="true" pageId="4" pageNumber="451">Ponera</emphasis>
</taxonomicName>
] Type-species:
<taxonomicName id="C6986CCEFFB7424AFC46403DFB94FABF" box="[901,1073,1293,1317]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB7424AFC46403DFB94FABF" box="[901,1073,1293,1317]" italics="true" pageId="4" pageNumber="451">Ponera sikorae</emphasis>
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="0127174DFFB7424AFF544003FBD0FAFD" blockId="4.[151,1436,1293,1584]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF544003FEA3FADD" authority="Forel, 1891" box="[151,262,1328,1351]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFF544003FEA3FADD" box="[151,262,1328,1351]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
in
<taxonomicName id="C6986CCEFFB7424AFEEF401DFE3AFADC" ID-CoL="K6Z" box="[300,415,1326,1350]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="subFamily" subFamily="Ponerinae">Ponerinae</taxonomicName>
,
<taxonomicName id="C6986CCEFFB7424AFE6F401DFDAAFADC" ID-CoL="L68" box="[428,527,1326,1350]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="tribe" tribe="Ponerini">Ponerini</taxonomicName>
:
<bibRefCitation id="65096ABCFFB7424AFDD8401CFD4CFADC" author="Emery, C." box="[539,745,1326,1351]" firstAuthor="Emery" journalOrPublisher="Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere" page="767" pageId="4" pageNumber="451" pagination="685" part="8" refId="ref19506" refString="Emery, C. (1895) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere 8, 685 - 778." title="Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden" type="journal article" year="1895">Emery, 1895: 767</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFD34401CFC17FADC" author="Emery, C." box="[759,946,1326,1351]" firstAuthor="Emery" journalOrPublisher="Genera Insectorum" page="79" pageId="4" pageNumber="451" pagination="1 - 125" part="118" refId="ref19624" refString="Emery, C. (1911) Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118, 1 - 125." title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae" type="journal article" year="1911">Emery, 1911: 79</bibRefCitation>
[subtribe
<taxonomicName id="C6986CCEFFB7424AFBE4401DFB79FADC" box="[1063,1244,1326,1350]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="tribe" tribe="Pachycondylini">Pachycondylini</taxonomicName>
];
<bibRefCitation id="65096ABCFFB7424AFB2C401DFF43FAFD" author="Wheeler, W. M." firstAuthor="Wheeler" journalOrPublisher="Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg" page="37" pageId="4" pageNumber="451" pagination="1 - 142" part="55" refId="ref20068" refString="Wheeler, W. M. (1915) The ants of the Baltic Amber. Schriften der Physikalisch-Okonomischen Gesellschaft zu Konigsberg, 55, 1 - 142." title="The ants of the Baltic Amber" type="journal article" year="1915">Wheeler, 1915: 37</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFF31407CFE10FAFD" author="Arnold, G." box="[242,437,1359,1383]" firstAuthor="Arnold" journalOrPublisher="Annals of the South African Museum" page="63" pageId="4" pageNumber="451" pagination="1 - 159" part="14" refId="ref19061" refString="Arnold, G. (1915) A monograph of the Formicidae of South Africa Part I. Ponerinae, Dorylinae. Annals of the South African Museum, 14, 1 - 159." title="A monograph of the Formicidae of South Africa Part I. Ponerinae, Dorylinae" type="journal article" year="1915">Arnold, 1915: 63</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFE03407CFDD8FAFD" author="Forel, A." box="[448,637,1359,1383]" firstAuthor="Forel" journalOrPublisher="Bulletin de la Societe Vaudoise des Sciences Naturelles" page="237" pageId="4" pageNumber="451" pagination="229" part="51" refId="ref19709" refString="Forel, A. (1917) Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Societe Vaudoise des Sciences Naturelles 51, 229 - 253." title="Cadre synoptique actuel de la faune universelle des fourmis" type="journal article" year="1917">Forel, 1917: 237</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFD4A407CFCCDFAFD" author="Wheeler, W. M." box="[649,872,1359,1383]" firstAuthor="Wheeler" journalOrPublisher="Bulletin of the American Museum of Natural History" page="648" pageId="4" pageNumber="451" pagination="1005 - 1055" part="45" refId="ref20097" refString="Wheeler, W. M. (1922) Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region. Bulletin of the American Museum of Natural History, 45, 1005 - 1055." title="Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region" type="journal article" year="1922">Wheeler, 1922: 648</bibRefCitation>
; all subsequent authors.
</paragraph>
<paragraph id="0127174DFFB7424AFF544041FC14FA13" blockId="4.[151,1436,1293,1584]" box="[151,945,1393,1418]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF544041FEA3FA13" box="[151,262,1394,1417]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFF544041FEA3FA13" box="[151,262,1394,1417]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
as subgenus of
<taxonomicName id="C6986CCEFFB7424AFE7B4041FD73FA13" ID-CoL="63NZ7" authority="Forel, 1891: 126" authorityName="Forel" authorityPageNumber="126" authorityYear="1891" box="[440,726,1393,1417]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFE7B4041FDAEFA13" box="[440,523,1394,1417]" italics="true" pageId="4" pageNumber="451">Ponera</emphasis>
:
<bibRefCitation id="65096ABCFFB7424AFDDA4042FD73FA13" author="Forel, A." bookContentInfo="237 pp." box="[537,726,1393,1417]" editor="Grandidier, A." firstAuthor="Forel" journalOrPublisher="Hachette et Cie, Paris" page="126" pageId="4" pageNumber="451" refId="ref19650" refString="Forel, A. (1891) Les Formicides. [part]. In: Grandidier, A. 1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule). Hachette et Cie, Paris, v + 237 pp." title="Les Formicides. [part]" type="book" volumeTitle="1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule)" year="1891">Forel, 1891: 126</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="65096ABCFFB7424AFD214041FC08FA13" author="Emery, C." box="[738,941,1393,1418]" firstAuthor="Emery" journalOrPublisher="Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere" page="767" pageId="4" pageNumber="451" pagination="685" part="8" refId="ref19506" refString="Emery, C. (1895) Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbucher Abteilung fur Systematik Okologie und Geographie der Tiere 8, 685 - 778." title="Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden" type="journal article" year="1895">Emery, 1895: 767</bibRefCitation>
.
</paragraph>
<paragraph id="0127174DFFB7424AFF5440A7FC85F994" blockId="4.[151,1436,1293,1584]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF5440A7FEA3FA31" box="[151,262,1428,1451]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFF5440A7FEA3FA31" box="[151,262,1428,1451]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
as genus:
<bibRefCitation id="65096ABCFFB7424AFEBF40A0FD9FFA30" author="Emery, C." box="[380,570,1426,1451]" firstAuthor="Emery" journalOrPublisher="Annales de la Societe Entomologique de Belgique" page="46" pageId="4" pageNumber="451" pagination="32 - 54" part="45" refId="ref19541" refString="Emery, C. (1901) Notes sur les sous-familles des Dorylines et Ponerines (Famille des Formicides). Annales de la Societe Entomologique de Belgique, 45, 32 - 54." title="Notes sur les sous-familles des Dorylines et Ponerines (Famille des Formicides)" type="journal article" year="1901">Emery, 1901: 46</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFD8540A0FCB4FA30" author="Emery, C." box="[582,785,1426,1451]" firstAuthor="Emery" journalOrPublisher="Deutsche Entomologische Zeitschrift" page="364" pageId="4" pageNumber="451" pagination="355" part="1909" refId="ref19576" refString="Emery, C. (1909) Beitrage zur Monographie der Formiciden des palaarktischen Faunengebietes. (Hym.). Teil VIII. Deutsche Entomologische Zeitschrift, 1909, 355 - 376." title="Beitrage zur Monographie der Formiciden des palaarktischen Faunengebietes. (Hym.). Teil VIII" type="journal article" year="1909">Emery, 1909: 364</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFCDD40A1FC5BFA30" author="Wheeler, W. M." bookContentInfo="663 pp." box="[798,1022,1426,1450]" firstAuthor="Wheeler" journalOrPublisher="Columbia University Press, New York, xxv" page="135" pageId="4" pageNumber="451" refId="ref20038" refString="Wheeler, W. M. (1910) Ants: Their Structure, Development and Behavior. Columbia University Press, New York, xxv + 663 pp." title="Ants: Their Structure, Development and Behavior" type="book" year="1910">Wheeler, 1910: 135</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFBC940A0FB62FA30" author="Emery, C." box="[1034,1223,1426,1451]" firstAuthor="Emery" journalOrPublisher="Genera Insectorum" page="79" pageId="4" pageNumber="451" pagination="1 - 125" part="118" refId="ref19624" refString="Emery, C. (1911) Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum, 118, 1 - 125." title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae" type="journal article" year="1911">Emery, 1911: 79</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFB1040A1FA32FA30" author="Arnold, G." box="[1235,1431,1426,1450]" firstAuthor="Arnold" journalOrPublisher="Annals of the South African Museum" page="63" pageId="4" pageNumber="451" pagination="1 - 159" part="14" refId="ref19061" refString="Arnold, G. (1915) A monograph of the Formicidae of South Africa Part I. Ponerinae, Dorylinae. Annals of the South African Museum, 14, 1 - 159." title="A monograph of the Formicidae of South Africa Part I. Ponerinae, Dorylinae" type="journal article" year="1915">Arnold, 1915: 63</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFF044087FE22FA56" author="Forel, A." box="[199,391,1460,1484]" firstAuthor="Forel" journalOrPublisher="Bulletin de la Societe Vaudoise des Sciences Naturelles" page="237" pageId="4" pageNumber="451" pagination="229" part="51" refId="ref19709" refString="Forel, A. (1917) Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Societe Vaudoise des Sciences Naturelles 51, 229 - 253." title="Cadre synoptique actuel de la faune universelle des fourmis" type="journal article" year="1917">Forel, 1917: 237</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFE574087FDC9FA56" author="Gallardo, A." box="[404,620,1460,1484]" firstAuthor="Gallardo" journalOrPublisher="Anales del Museo Nacional de Buenos Aires" page="64" pageId="4" pageNumber="451" pagination="1 - 12" part="30" refId="ref19740" refString="Gallardo, A. (1918) Las hormigas de la Republica Argentina. Subfamilia ponerinas. Anales del Museo Nacional de Buenos Aires, 30, 1 - 12." title="Las hormigas de la Republica Argentina. Subfamilia ponerinas" type="journal article" year="1918">Gallardo, 1918: 64</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFDB94087FCF9FA56" author="Wheeler, W. M." box="[634,860,1460,1484]" firstAuthor="Wheeler" journalOrPublisher="Bulletin of the American Museum of Natural History" page="648" pageId="4" pageNumber="451" pagination="1005 - 1055" part="45" refId="ref20097" refString="Wheeler, W. M. (1922) Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region. Bulletin of the American Museum of Natural History, 45, 1005 - 1055." title="Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region" type="journal article" year="1922">Wheeler, 1922: 648</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFCAA4087FBF3FA56" author="Borgmeier, T." box="[873,1110,1460,1484]" firstAuthor="Borgmeier" journalOrPublisher="Archivo do Museu Nacional (Rio de Janeiro)" page="70" pageId="4" pageNumber="451" pagination="33 - 103" part="24" refId="ref19212" refString="Borgmeier, T. (1923) Catalogo systematico e synonymico das formigas do Brasil. 1 parte. Subfam. Dorylinae, Cerapachyinae, Ponerinae, Dolichoderinae. Archivo do Museu Nacional (Rio de Janeiro), 24, 33 - 103." title="Catalogo systematico e synonymico das formigas do Brasil. 1 parte. Subfam. Dorylinae, Cerapachyinae, Ponerinae, Dolichoderinae" type="journal article" year="1923">Borgmeier, 1923: 70</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFBA04087FABDFA56" author="Clark, J." box="[1123,1304,1460,1484]" firstAuthor="Clark" journalOrPublisher="Memoirs of the National Museum of Victoria" page="30" pageId="4" pageNumber="451" pagination="21 - 47" part="8" refId="ref19303" refString="Clark, J. (1934) New Australian ants. Memoirs of the National Museum of Victoria, 8, 21 - 47." title="New Australian ants" type="journal article" year="1934">Clark, 1934: 30</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFAE64087FE94FA77" author="Creighton, W. S." firstAuthor="Creighton" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" page="44" pageId="4" pageNumber="451" pagination="1 - 585" part="104" refId="ref19328" refString="Creighton, W. S. (1950) The ants of North America. Bulletin of the Museum of Comparative Zoology, 104, 1 - 585." title="The ants of North America" type="journal article" year="1950">Creighton, 1950: 44</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFE8340E6FD3FFA77" author="Chapman, J. W. &amp; Capco, S. R." box="[320,666,1493,1517]" firstAuthor="Chapman" journalOrPublisher="Monographs of the Institute of Science and Technology. Manila" page="63" pageId="4" pageNumber="451" pagination="1 - 327" part="1" refId="ref19257" refString="Chapman, J. W. &amp; Capco, S. R. (1951) Check list of the ants (Hymenoptera: Formicidae) of Asia. Monographs of the Institute of Science and Technology. Manila, 1, 1 - 327." title="Check list of the ants (Hymenoptera: Formicidae) of Asia" type="journal article" year="1951">Chapman &amp; Capco, 1951: 63</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFD6940E6FC2BFA77" author="Bernard, F." box="[682,910,1493,1517]" firstAuthor="Bernard" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" page="189" pageId="4" pageNumber="451" pagination="165" part="19" refId="ref19096" refString="Bernard, F. (1953) La reserve naturelle integrale du Mt Nimba. XI. Hymenopteres Formicidae. Memoires de l'Institut Francais d'Afrique Noire, 19, 165 - 270." title="La reserve naturelle integrale du Mt Nimba. XI. Hymenopteres Formicidae" type="journal article" year="1953">Bernard, 1953: 189</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFC5D40E6FAB5FA77" author="Wheeler, G. C. &amp; Wheeler, J." box="[926,1296,1493,1517]" firstAuthor="Wheeler" journalOrPublisher="Transactions of the American Entomological Society" page="256" pageId="4" pageNumber="451" pagination="255" part="111" refId="ref20004" refString="Wheeler, G. C. &amp; Wheeler, J. (1985) A simplified conspectus of the Formicidae. Transactions of the American Entomological Society, 111, 255 - 264." title="A simplified conspectus of the Formicidae" type="journal article" year="1985">Wheeler &amp; Wheeler, 1985: 256</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFADC40E6FE15F994" author="Dlussky, G. M. &amp; Fedoseeva, E. B." editor="Ponomarenko, A. G." firstAuthor="Dlussky" journalOrPublisher="Moskva, Nauka" page="78" pageId="4" pageNumber="451" pagination="70 - 144" refId="ref19393" refString="Dlussky, G. M. &amp; Fedoseeva, E. B. (1988) Origin and early stages of evolution in ants. [In Russian.]. In: Ponomarenko, A. G. (ed.). Cretaceous biocenotic crisis and insect evolution. [In Russian.]. Moskva, Nauka, pp. 70 - 144." title="Origin and early stages of evolution in ants. [In Russian.]" type="book chapter" volumeTitle="Cretaceous biocenotic crisis and insect evolution. [In Russian.]" year="1988">Dlussky &amp; Fedoseeva, 1988: 78</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFE7F40C5FCB9F994" author="Holldobler, B. &amp; Wilson E. O." bookContentInfo="732 pp." box="[444,796,1526,1550]" firstAuthor="Holldobler" journalOrPublisher="Harvard University Press. Cambridge, Massachusetts, USA" page="11" pageId="4" pageNumber="451" refId="ref19824" refString="Holldobler, B. &amp; Wilson E. O. (1990) The Ants. Harvard University Press. Cambridge, Massachusetts, USA. 732 pp." title="The Ants" type="book" year="1990">Hölldobler &amp; Wilson, 1990: 11</bibRefCitation>
.
</paragraph>
<paragraph id="0127174DFFB7424AFF54432AFB74F9B5" blockId="4.[151,1436,1293,1584]" box="[151,1233,1559,1584]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF54432AFEA3F9AA" box="[151,262,1561,1584]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFF54432AFEA3F9AA" box="[151,262,1561,1584]" italics="true" pageId="4" pageNumber="451">Euponera</emphasis>
</taxonomicName>
as junior synonym of
<taxonomicName id="C6986CCEFFB7424AFDC3432BFD58F9B5" ID-CoL="6BRJ" authority="Brown" authorityName="Brown" box="[512,765,1560,1584]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="33ECCB5FFFB7424AFDC3432BFD3AF9AA" box="[512,671,1560,1584]" italics="true" pageId="4" pageNumber="451">Pachycondyla</emphasis>
: Brown
</taxonomicName>
, in
<bibRefCitation id="65096ABCFFB7424AFCE64324FC56F9B5" author="Bolton, B." bookContentInfo="222 pp." box="[805,1011,1559,1583]" firstAuthor="Bolton" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" page="164" pageId="4" pageNumber="451" refId="ref19129" refString="Bolton, B. (1994) Identification Guide to the Ant Genera of the World. Harvard University Press, Cambridge, Massachusetts, 222 pp." title="Identification Guide to the Ant Genera of the World" type="book" year="1994">Bolton, 1994: 164</bibRefCitation>
;
<bibRefCitation id="65096ABCFFB7424AFC3C4324FB68F9B5" author="Bolton, B." box="[1023,1229,1559,1583]" firstAuthor="Bolton" journalOrPublisher="Memoirs of the American Entomological Institute" page="166" pageId="4" pageNumber="451" pagination="1 - 370" part="71" refId="ref19186" refString="Bolton, B. (2003) Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute, 71, 1 - 370." title="Synopsis and classification of Formicidae" type="journal article" year="2003">Bolton, 2003: 166</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="498244C6FFB7424BFF5443B0FDF3FE36" lastPageId="5" lastPageNumber="452" pageId="4" pageNumber="451" type="multiple">
<paragraph id="0127174DFFB7424AFF5443B0FCADF907" blockId="4.[151,776,1667,1694]" box="[151,776,1667,1694]" pageId="4" pageNumber="451">
<heading id="5A6FA021FFB7424AFF5443B0FCADF907" bold="true" box="[151,776,1667,1694]" fontSize="11" level="1" pageId="4" pageNumber="451" reason="1">
<emphasis id="33ECCB5FFFB7424AFF5443B0FCADF907" bold="true" box="[151,776,1667,1694]" pageId="4" pageNumber="451">
Checklist of the
<collectingCountry id="798F57DDFFB7424AFEA243B0FE7FF907" box="[353,474,1667,1693]" name="Madagascar" pageId="4" pageNumber="451">Malagasy</collectingCountry>
<emphasis id="33ECCB5FFFB7424AFE2243B7FDF7F904" bold="true" box="[481,594,1668,1694]" italics="true" pageId="4" pageNumber="451">P. s i k o r a e</emphasis>
species-group
</emphasis>
</heading>
</paragraph>
<paragraph id="0127174DFFB7424AFF5443F8FEFCF978" blockId="4.[151,672,1738,2015]" box="[151,345,1738,1763]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF5443F8FF4CF979" box="[151,233,1739,1763]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB7424AFF5443F8FF4CF979" box="[151,233,1739,1763]" italics="true" pageId="4" pageNumber="451">sikorae</emphasis>
</taxonomicName>
complex:
</paragraph>
<paragraph id="0127174DFFB7424AFF0443DCFD29F89C" blockId="4.[151,672,1738,2015]" box="[199,652,1774,1798]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF0443DCFDE0F89C" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[199,581,1774,1798]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="gorogota" status="sp. nov.">
<emphasis id="33ECCB5FFFB7424AFF0443DCFE88F89C" bold="true" box="[199,301,1775,1798]" italics="true" pageId="4" pageNumber="451">gorogota</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB7424AFD9343DCFD29F89C" box="[592,652,1775,1798]" pageId="4" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB7424AFF044221FD05F8B0" blockId="4.[151,672,1738,2015]" box="[199,672,1810,1835]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF044221FDFCF8B0" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[199,601,1810,1835]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="haratsingy" status="sp. nov.">
<emphasis id="33ECCB5FFFB7424AFF044221FEE7F8B1" bold="true" box="[199,322,1810,1835]" italics="true" pageId="4" pageNumber="451">haratsingy</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB7424AFDA74220FD05F8B0" box="[612,672,1811,1834]" pageId="4" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB7424AFF044205FDF8F8D4" blockId="4.[151,672,1738,2015]" box="[199,605,1846,1871]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF044205FDB3F8D4" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[199,534,1846,1871]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="ivolo" status="sp. nov.">
<emphasis id="33ECCB5FFFB7424AFF044205FF5AF8D5" bold="true" box="[199,255,1846,1871]" italics="true" pageId="4" pageNumber="451">ivolo</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB7424AFDE24204FDF8F8D4" box="[545,605,1847,1870]" pageId="4" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB7424AFF044268FDCAF8E8" blockId="4.[151,672,1738,2015]" box="[199,623,1882,1906]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF044268FD8CF8E8" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[199,553,1882,1906]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="maeva" status="sp. nov.">
<emphasis id="33ECCB5FFFB7424AFF044268FEB7F8E8" bold="true" box="[199,274,1883,1906]" italics="true" pageId="4" pageNumber="451">maeva</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB7424AFDF74268FDCAF8E8" box="[564,623,1883,1906]" pageId="4" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB7424AFF04424DFDC0F80C" blockId="4.[151,672,1738,2015]" box="[199,613,1918,1943]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF04424DFDBBF80C" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[199,542,1918,1943]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="mialy" status="sp. nov.">
<emphasis id="33ECCB5FFFB7424AFF04424DFEA2F80D" bold="true" box="[199,263,1918,1943]" italics="true" pageId="4" pageNumber="451">mialy</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB7424AFDEA424CFDC0F80C" box="[553,613,1919,1942]" pageId="4" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB7424AFF044290FDFFF820" blockId="4.[151,672,1738,2015]" box="[199,602,1954,1978]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF044290FDB6F820" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[199,531,1954,1978]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="nosy" status="sp. nov.">
<emphasis id="33ECCB5FFFB7424AFF044290FF59F820" bold="true" box="[199,252,1955,1978]" italics="true" pageId="4" pageNumber="451">nosy</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB7424AFDDD4290FDFFF820" box="[542,602,1955,1978]" pageId="4" pageNumber="451" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB7424AFF0442F5FE39F844" blockId="4.[151,672,1738,2015]" box="[199,412,1990,2015]" pageId="4" pageNumber="451">
<taxonomicName id="C6986CCEFFB7424AFF0442F5FE39F844" authority="Forel 1891" authorityName="Forel" authorityYear="1891" box="[199,412,1990,2015]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="451" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB7424AFF0442F5FEBEF845" bold="true" box="[199,283,1990,2015]" italics="true" pageId="4" pageNumber="451">sikorae</emphasis>
<bibRefCitation id="65096ABCFFB7424AFEE142F5FE39F844" author="Forel, A." bookContentInfo="237 pp." box="[290,412,1990,2014]" editor="Grandidier, A." firstAuthor="Forel" journalOrPublisher="Hachette et Cie, Paris" pageId="4" pageNumber="451" refId="ref19650" refString="Forel, A. (1891) Les Formicides. [part]. In: Grandidier, A. 1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule). Hachette et Cie, Paris, v + 237 pp." title="Les Formicides. [part]" type="book" volumeTitle="1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule)" year="1891">Forel 1891</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="0127174DFFB6424BFF5445ABFED7FF35" blockId="5.[151,657,151,428]" box="[151,370,151,176]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF5445ABFEA7FF2A" box="[151,258,152,176]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="vohitravo">
<emphasis id="33ECCB5FFFB6424BFF5445ABFEA7FF2A" box="[151,258,152,176]" italics="true" pageId="5" pageNumber="452">vohitravo</emphasis>
</taxonomicName>
complex:
</paragraph>
<paragraph id="0127174DFFB6424BFF064588FDD7FF4E" blockId="5.[151,657,151,428]" box="[197,626,187,212]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF064588FD8EFF4E" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,555,187,212]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="agnivo" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF064588FEB6FF4E" bold="true" box="[197,275,187,212]" italics="true" pageId="5" pageNumber="452">agnivo</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFDF5458EFDD7FF4E" box="[566,626,189,212]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB6424BFF0645ECFD35FF6D" blockId="5.[151,657,151,428]" box="[197,656,223,248]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF0645ECFDECFF6D" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,585,223,248]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="antsiraka" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF0645ECFE94FF62" bold="true" box="[197,305,223,248]" italics="true" pageId="5" pageNumber="452">antsiraka</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFD9745D3FD35FF6D" box="[596,656,224,247]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB6424BFF064430FDDBFE86" blockId="5.[151,657,151,428]" box="[197,638,259,284]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF064430FD92FE86" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,567,259,284]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="daraina" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF064430FEBAFE86" bold="true" box="[197,287,259,284]" italics="true" pageId="5" pageNumber="452">daraina</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFD814436FDDBFE86" box="[578,638,261,284]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB6424BFF06441BFDD0FEA5" blockId="5.[151,657,151,428]" box="[197,629,295,319]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF06441BFD8BFEA5" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,558,295,319]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="rovana" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF06441BFEB3FEA5" bold="true" box="[197,278,296,319]" italics="true" pageId="5" pageNumber="452">rovana</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFDFA441BFDD0FEA5" box="[569,629,296,319]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB6424BFF064478FDCBFEFE" blockId="5.[151,657,151,428]" box="[197,622,331,356]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF064478FD82FEFE" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,551,331,356]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="tahary" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF064478FEAAFEFE" bold="true" box="[197,271,331,356]" italics="true" pageId="5" pageNumber="452">tahary</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFDF1447EFDCBFEFE" box="[562,622,333,356]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB6424BFF06445CFD34FE1D" blockId="5.[151,657,151,428]" box="[197,657,367,392]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF06445CFDEFFE1D" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,586,367,392]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="vohitravo" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF06445CFE97FE12" bold="true" box="[197,306,367,392]" italics="true" pageId="5" pageNumber="452">vohitravo</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFD964443FD34FE1D" box="[597,657,368,391]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="0127174DFFB6424BFF0644A7FDF3FE36" blockId="5.[151,657,151,428]" box="[197,598,404,428]" pageId="5" pageNumber="452">
<taxonomicName id="C6986CCEFFB6424BFF0644A7FDAAFE36" authority="Rakotonirina and Fisher" authorityName="Rakotonirina and Fisher" box="[197,527,404,428]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="zoro" status="sp. nov.">
<emphasis id="33ECCB5FFFB6424BFF0644A7FF52FE31" bold="true" box="[197,247,404,427]" italics="true" pageId="5" pageNumber="452">zoro</emphasis>
Rakotonirina and Fisher
</taxonomicName>
,
<taxonomicNameLabel id="28DF7624FFB6424BFDD944A6FDF3FE36" box="[538,598,405,428]" pageId="5" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="498244C6FFB64244FF5444CCFA3EFDDD" lastPageId="10" lastPageNumber="457" pageId="5" pageNumber="452" type="key">
<paragraph id="0127174DFFB6424BFF5444CCFCB1FD83" blockId="5.[151,788,511,537]" box="[151,788,511,537]" pageId="5" pageNumber="452">
<heading id="5A6FA021FFB6424BFF5444CCFCB1FD83" bold="true" box="[151,788,511,537]" fontSize="11" level="1" pageId="5" pageNumber="452" reason="1">
<emphasis id="33ECCB5FFFB6424BFF5444CCFCB1FD83" bold="true" box="[151,788,511,537]" pageId="5" pageNumber="452">
Key to species of
<collectingCountry id="798F57DDFFB6424BFEAD44CCFE43FD83" box="[366,486,511,537]" name="Madagascar" pageId="5" pageNumber="452">Malagasy</collectingCountry>
<emphasis id="33ECCB5FFFB6424BFE2E44CCFDFBFD83" bold="true" box="[493,606,511,537]" italics="true" pageId="5" pageNumber="452">P. sikorae</emphasis>
species-group
</emphasis>
</heading>
</paragraph>
<paragraph id="0127174DFFB6424BFF544775FCA3FD18" blockId="5.[151,1436,582,642]" pageId="5" pageNumber="452">The function of the key is primarily to identify workers, but can also identify ergatoid and winged queens to species as long as the shape of the mesosoma is not considered.</paragraph>
<key id="7509407CFFB64244FF544781FA3EFDDD" lastPageId="10" lastPageNumber="457" pageId="5" pageNumber="452">
<keyStep id="BA6C0FE8FFB6424BFF544781FA39FC85" pageId="5" pageNumber="452">
<paragraph id="0127174DFFB6424BFF544781FA39FD7E" blockId="5.[151,1436,690,799]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF544781FA39FD7E" pageId="5" pageNumber="452">
1. With head in full-face view, anterior margin of clypeus widely transverse and more or less straight, with weak median notch (
<figureCitation id="99A30BC8FFB6424BFF2B47FCFE87FD7E" box="[232,290,719,740]" captionStart="FIGURES 2 3" captionStartId="5.[151,264,1370,1393]" captionTargetBox="[151,1435,852,1336]" captionTargetId="figure@5.[151,1435,852,1347]" captionTargetPageId="5" captionText="FIGURES 2 3. Shape of the clypeus with head in full-face view; Fig. 2: anterior margin widely transverse and straight with feeble median notch (CASENT 0497667); Fig. 3: anterior margin medially convex or bluntly angulate (CASENT 0050524)." httpUri="https://zenodo.org/record/247518/files/figure.png" pageId="5" pageNumber="452">Fig. 2</figureCitation>
) (
<taxonomicName id="C6986CCEFFB6424BFEF447E3FED8FD7F" box="[311,381,720,741]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB6424BFEF447E3FED8FD7F" box="[311,381,720,741]" italics="true" pageId="5" pageNumber="452">sikorae</emphasis>
</taxonomicName>
complex)............................................................................... 2
</keyLead>
</paragraph>
<paragraph id="0127174DFFB6424BFF5447DEFA39FC85" blockId="5.[151,1436,690,799]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF5447DEFA39FC85" pageId="5" pageNumber="452">
- With head in full-face view, anterior margin of clypeus generally convex or medially bluntly angulate; slight median notch may be present (
<figureCitation id="99A30BC8FFB6424BFEBC4639FE1DFC85" box="[383,440,778,799]" captionStart="FIGURES 2 3" captionStartId="5.[151,264,1370,1393]" captionTargetBox="[151,1435,852,1336]" captionTargetId="figure@5.[151,1435,852,1347]" captionTargetPageId="5" captionText="FIGURES 2 3. Shape of the clypeus with head in full-face view; Fig. 2: anterior margin widely transverse and straight with feeble median notch (CASENT 0497667); Fig. 3: anterior margin medially convex or bluntly angulate (CASENT 0050524)." httpUri="https://zenodo.org/record/247518/files/figure.png" pageId="5" pageNumber="452">Fig. 3</figureCitation>
) (
<taxonomicName id="C6986CCEFFB6424BFE0D4639FD8FFC85" box="[462,554,778,799]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="vohitravo">
<emphasis id="33ECCB5FFFB6424BFE0D4639FD8FFC85" box="[462,554,778,799]" italics="true" pageId="5" pageNumber="452">vohitravo</emphasis>
</taxonomicName>
complex)................................................................ 8
</keyLead>
</paragraph>
</keyStep>
<caption id="55E747C5FFB6424BFF544069FAD3FA0A" httpUri="https://zenodo.org/record/247518/files/figure.png" pageId="5" pageNumber="452" targetBox="[151,1435,852,1336]" targetPageId="5">
<paragraph id="0127174DFFB6424BFF544069FAD3FA0A" blockId="5.[151,1436,1370,1424]" pageId="5" pageNumber="452">
<emphasis id="33ECCB5FFFB6424BFF544069FE9EFAEA" bold="true" box="[151,315,1370,1393]" pageId="5" pageNumber="452">FIGURES 23.</emphasis>
Shape of the clypeus with head in full-face view; Fig. 2: anterior margin widely transverse and straight with feeble median notch (CASENT0497667); Fig. 3: anterior margin medially convex or bluntly angulate (CASENT0050524).
</paragraph>
</caption>
<keyStep id="BA6C0FE8FFB6424BFF54408FFA39F9B3" pageId="5" pageNumber="452">
<paragraph id="0127174DFFB6424BFF54408FFA39FA74" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF54408FFA39FA74" pageId="5" pageNumber="452">
2. Posterior surface of petiole node covered with numerous slender hairs and abundant pubescence; anterior surface of third abdominal segment concave, forming a shallow cavity to lodge the posterior surface of the petiole (
<figureCitation id="99A30BC8FFB6424BFB5940EAFB71FA74" box="[1178,1236,1497,1518]" captionStart="FIGURES 4 5" captionStartId="6.[151,264,684,707]" captionTargetBox="[151,1434,193,645]" captionTargetId="figure@6.[151,1436,193,660]" captionTargetPageId="6" captionText="FIGURES 4 5. Petiole and gaster in dorsal view; Fig. 4: posterior surface of petiole with abundant pubescence and anterior face of first gastral segment forming shallow cavity (CASENT 0195184); Fig. 5: abundant pubescence absent on posterior surface of petiole and anterior face of first gastral segment straight without shallow cavity (CASENT 0231239)." httpUri="https://zenodo.org/record/247519/files/figure.png" pageId="5" pageNumber="452">Fig. 4</figureCitation>
)............... 3
</keyLead>
</paragraph>
<paragraph id="0127174DFFB6424BFF5440C4FA39F9B3" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF5440C4FA39F9B3" pageId="5" pageNumber="452">
- Posterior surface of petiole node convex and not covered with numerous slender hairs and abundant pubescence; anterior surface of the third abdominal segment straight, without shallow cavity to lodge the posterior surface of the petiole (
<figureCitation id="99A30BC8FFB6424BFADD4327FAF2F9B3" box="[1310,1367,1556,1577]" captionStart="FIGURES 4 5" captionStartId="6.[151,264,684,707]" captionTargetBox="[151,1434,193,645]" captionTargetId="figure@6.[151,1436,193,660]" captionTargetPageId="6" captionText="FIGURES 4 5. Petiole and gaster in dorsal view; Fig. 4: posterior surface of petiole with abundant pubescence and anterior face of first gastral segment forming shallow cavity (CASENT 0195184); Fig. 5: abundant pubescence absent on posterior surface of petiole and anterior face of first gastral segment straight without shallow cavity (CASENT 0231239)." httpUri="https://zenodo.org/record/247519/files/figure.png" pageId="5" pageNumber="452">Fig. 5</figureCitation>
).... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFB6424BFF544302FA39F904" pageId="5" pageNumber="452">
<paragraph id="0127174DFFB6424BFF544302FA39F9FE" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF544302FA39F9FE" pageId="5" pageNumber="452">
3. Lateral surface of head and petiole node smooth and shiny, with scattered piligerous punctures (
<figureCitation id="99A30BC8FFB6424BFB524302FB6EF9DC" box="[1169,1227,1585,1606]" captionStart="FIGURES 6 7" captionStartId="6.[151,264,1306,1329]" captionTargetBox="[153,1433,844,1259]" captionTargetId="figure@6.[153,1433,844,1272]" captionTargetPageId="6" captionText="FIGURES 6 7. Lateral view showing the sculpture of lateral portion of head and petiole; Fig. 6: head and petiole smooth and shining (CASENT 0050330: holotype); Fig. 7: head densely and finely reticulate-rugulose, petiolar node densely and finely punctate (CASENT 0195184)." httpUri="https://zenodo.org/record/247520/files/figure.png" pageId="5" pageNumber="452">Fig. 6</figureCitation>
) [known from queen specimen]........................................................................................
<taxonomicName id="C6986CCEFFB6424BFAAF437CFA39F9FE" box="[1388,1436,1615,1636]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="ivolo">
<emphasis id="33ECCB5FFFB6424BFAAF437CFA39F9FE" bold="true" box="[1388,1436,1615,1636]" italics="true" pageId="5" pageNumber="452">ivolo</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0127174DFFB6424BFF54435FFA39F904" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF54435FFA39F904" pageId="5" pageNumber="452">
- Lateral portion of head densely and finely reticulate-rugulose, sides of petiolar node densely and finely punctate (
<figureCitation id="99A30BC8FFB6424BFAE3435FFAFEF91B" box="[1312,1371,1644,1665]" captionStart="FIGURES 6 7" captionStartId="6.[151,264,1306,1329]" captionTargetBox="[153,1433,844,1259]" captionTargetId="figure@6.[153,1433,844,1272]" captionTargetPageId="6" captionText="FIGURES 6 7. Lateral view showing the sculpture of lateral portion of head and petiole; Fig. 6: head and petiole smooth and shining (CASENT 0050330: holotype); Fig. 7: head densely and finely reticulate-rugulose, petiolar node densely and finely punctate (CASENT 0195184)." httpUri="https://zenodo.org/record/247520/files/figure.png" pageId="5" pageNumber="452">Fig. 7</figureCitation>
).................................................................................................
<taxonomicName id="C6986CCEFFB6424BFAF043BAFA39F904" box="[1331,1436,1673,1694]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="haratsingy">
<emphasis id="33ECCB5FFFB6424BFAF043BAFA39F904" bold="true" box="[1331,1436,1673,1694]" italics="true" pageId="5" pageNumber="452">haratsingy</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFB6424BFF544394FA39F88E" pageId="5" pageNumber="452">
<paragraph id="0127174DFFB6424BFF544394FA39F943" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF544394FA39F943" pageId="5" pageNumber="452">
4. With mesosoma in dorsal view, metanotal groove deeply impressed (
<figureCitation id="99A30BC8FFB6424BFC474394FC65F926" box="[900,960,1703,1724]" captionStart="FIGURES 8 9" captionStartId="6.[151,264,1979,2002]" captionTargetBox="[153,1436,1464,1943]" captionTargetId="figure@6.[151,1436,1464,1956]" captionTargetPageId="6" captionText="FIGURES 8 9. Mesosoma in dorsal view; Fig. 8: metanotal groove deeply impressed (CASENT 0034340: holotype); Fig. 9: metanotal groove lacking (CASENT 0497667)." httpUri="https://zenodo.org/record/247521/files/figure.png" pageId="5" pageNumber="452">Fig. 8</figureCitation>
); in lateral view the dorsal outline of mesosoma complex, at least with distinct convex mesonotum (
<figureCitation id="99A30BC8FFB6424BFD0D43F7FCBDF943" box="[718,792,1732,1753]" captionStart="FIGURES 37 39" captionStartId="16.[151,264,1645,1668]" captionTargetBox="[151,1436,183,1622]" captionTargetId="figure@16.[151,1436,183,1639]" captionTargetPageId="16" captionText="FIGURES 37 39. Worker caste of Pachycondyla gorogota: lateral and dorsal view of body and full-face view of head, holotype CASENT 0166009." httpUri="https://zenodo.org/record/247534/files/figure.png" pageId="5" pageNumber="452">Figs 37</figureCitation>
,
<figureCitation id="99A30BC8FFB6424BFCE143F7FC9EF943" box="[802,827,1732,1753]" captionStart="FIGURES 46 48" captionStartId="21.[151,264,1981,2004]" captionTargetBox="[151,1434,399,1980]" captionTargetId="figure@21.[151,1436,399,1982]" captionTargetPageId="21" captionText="FIGURES 46 48. Worker caste of Pachycondyla maeva: lateral and dorsal view of body and full-face view of head, holotype CASENT 0034340." httpUri="https://zenodo.org/record/247537/files/figure.png" pageId="5" pageNumber="452">46</figureCitation>
)................................................. 5
</keyLead>
</paragraph>
<paragraph id="0127174DFFB6424BFF5443D2FA39F88E" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF5443D2FA39F88E" pageId="5" pageNumber="452">
- With mesosoma in dorsal view, metanotal groove not impressed or absent (
<figureCitation id="99A30BC8FFB6424BFC7543D2FC54F96C" box="[950,1009,1761,1782]" captionStart="FIGURES 8 9" captionStartId="6.[151,264,1979,2002]" captionTargetBox="[153,1436,1464,1943]" captionTargetId="figure@6.[151,1436,1464,1956]" captionTargetPageId="6" captionText="FIGURES 8 9. Mesosoma in dorsal view; Fig. 8: metanotal groove deeply impressed (CASENT 0034340: holotype); Fig. 9: metanotal groove lacking (CASENT 0497667)." httpUri="https://zenodo.org/record/247521/files/figure.png" pageId="5" pageNumber="452">Fig. 9</figureCitation>
); in lateral view, the dorsal outline of mesosoma almost straight or slightly evenly convex, without breaks (
<figureCitation id="99A30BC8FFB6424BFC8443CCFC11F88E" box="[839,948,1791,1812]" captionStart="FIGURES 49 51" captionStartId="23.[151,264,1901,1924]" captionTargetBox="[151,1436,411,1861]" captionTargetId="figure@23.[151,1436,411,1878]" captionTargetPageId="23" captionText="FIGURES 49 51. Worker caste of Pachycondyla mialy: lateral and dorsal view of body and full-face view of head, holotype CASENT 0497667." httpUri="https://zenodo.org/record/247538/files/figure.png" pageId="5" pageNumber="452">Figs 49, 51</figureCitation>
)....................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFB6424BFF54422FFA39F85E" pageId="5" pageNumber="452">
<paragraph id="0127174DFFB6424BFF54422FFA39F8F6" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF54422FFA39F8F6" pageId="5" pageNumber="452">
5. Petiole node anteroposteriorly flattened in profile (
<figureCitation id="99A30BC8FFB6424BFD16422FFCBBF8AB" box="[725,798,1820,1841]" captionStart="FIGURES 10 11" captionStartId="7.[151,264,736,759]" captionTargetBox="[153,1436,195,700]" captionTargetId="figure@7.[151,1436,195,712]" captionTargetPageId="7" captionText="FIGURES 10 11. Form of petiole in profile; Fig. 10: anteroposteriorly flattened (CASENT 0034340); Fig. 11: nodiform about as long as broad (CASENT 0497202)." httpUri="https://zenodo.org/record/247522/files/figure.png" pageId="5" pageNumber="452">Fig. 10</figureCitation>
), in dorsal view its posterior margin medially notched; posterior margin of head slightly concave; antennal scape not surpassing the concave posterior cephalic margin (
<figureCitation id="99A30BC8FFB6424BFB02420AFAAEF8D4" box="[1217,1291,1849,1870]" captionStart="FIGURES 37 39" captionStartId="16.[151,264,1645,1668]" captionTargetBox="[151,1436,183,1622]" captionTargetId="figure@16.[151,1436,183,1639]" captionTargetPageId="16" captionText="FIGURES 37 39. Worker caste of Pachycondyla gorogota: lateral and dorsal view of body and full-face view of head, holotype CASENT 0166009." httpUri="https://zenodo.org/record/247534/files/figure.png" pageId="5" pageNumber="452">Figs 38</figureCitation>
,
<figureCitation id="99A30BC8FFB6424BFAD5420AFA94F8D4" box="[1302,1329,1849,1870]" captionStart="FIGURES 46 48" captionStartId="21.[151,264,1981,2004]" captionTargetBox="[151,1434,399,1980]" captionTargetId="figure@21.[151,1436,399,1982]" captionTargetPageId="21" captionText="FIGURES 46 48. Worker caste of Pachycondyla maeva: lateral and dorsal view of body and full-face view of head, holotype CASENT 0034340." httpUri="https://zenodo.org/record/247537/files/figure.png" pageId="5" pageNumber="452">47</figureCitation>
); mandible smooth between small punctures; tegument densely and finely reticulate-punctate (
<figureCitation id="99A30BC8FFB6424BFC384264FBE3F8F6" box="[1019,1094,1879,1900]" captionStart="FIGURES 37 39" captionStartId="16.[151,264,1645,1668]" captionTargetBox="[151,1436,183,1622]" captionTargetId="figure@16.[151,1436,183,1639]" captionTargetPageId="16" captionText="FIGURES 37 39. Worker caste of Pachycondyla gorogota: lateral and dorsal view of body and full-face view of head, holotype CASENT 0166009." httpUri="https://zenodo.org/record/247534/files/figure.png" pageId="5" pageNumber="452">Figs 37</figureCitation>
,
<figureCitation id="99A30BC8FFB6424BFB934264FBCCF8F6" box="[1104,1129,1879,1900]" captionStart="FIGURES 46 48" captionStartId="21.[151,264,1981,2004]" captionTargetBox="[151,1434,399,1980]" captionTargetId="figure@21.[151,1436,399,1982]" captionTargetPageId="21" captionText="FIGURES 46 48. Worker caste of Pachycondyla maeva: lateral and dorsal view of body and full-face view of head, holotype CASENT 0034340." httpUri="https://zenodo.org/record/247537/files/figure.png" pageId="5" pageNumber="452">46</figureCitation>
)........................ 6
</keyLead>
</paragraph>
<paragraph id="0127174DFFB6424BFF544247FA39F85E" blockId="5.[151,1436,1468,1988]" pageId="5" pageNumber="452">
<keyLead id="BA69B478FFB6424BFF544247FA39F85E" pageId="5" pageNumber="452">
- Petiole node thick in profile (
<figureCitation id="99A30BC8FFB6424BFDCB4247FDF4F813" box="[520,593,1908,1929]" captionStart="FIGURES 10 11" captionStartId="7.[151,264,736,759]" captionTargetBox="[153,1436,195,700]" captionTargetId="figure@7.[151,1436,195,712]" captionTargetPageId="7" captionText="FIGURES 10 11. Form of petiole in profile; Fig. 10: anteroposteriorly flattened (CASENT 0034340); Fig. 11: nodiform about as long as broad (CASENT 0497202)." httpUri="https://zenodo.org/record/247522/files/figure.png" pageId="5" pageNumber="452">Fig. 11</figureCitation>
); in dorsal view, the posterior margin broadly convex; antennal scape surpassing the broadly convex posterior cephalic margin (
<figureCitation id="99A30BC8FFB6424BFD4442A2FD74F83C" box="[647,721,1937,1958]" captionStart="FIGURES 58 60" captionStartId="28.[151,264,1531,1554]" captionTargetBox="[151,1436,195,1491]" captionTargetId="figure@28.[151,1436,195,1507]" captionTargetPageId="28" captionText="FIGURES 58 60. Worker caste of Pachycondyla sikorae: lateral and dorsal view of body and full-face view of head, CASENT 0497202." httpUri="https://zenodo.org/record/247541/files/figure.png" pageId="5" pageNumber="452">Fig. 59</figureCitation>
); mandible striate-punctate; black species with mostly smooth and shiny tegument (
<figureCitation id="99A30BC8FFB6424BFE84429CFE28F85E" box="[327,397,1967,1988]" captionStart="FIGURES 58 60" captionStartId="28.[151,264,1531,1554]" captionTargetBox="[151,1436,195,1491]" captionTargetId="figure@28.[151,1436,195,1507]" captionTargetPageId="28" captionText="FIGURES 58 60. Worker caste of Pachycondyla sikorae: lateral and dorsal view of body and full-face view of head, CASENT 0497202." httpUri="https://zenodo.org/record/247541/files/figure.png" pageId="5" pageNumber="452">Fig. 58</figureCitation>
)................................................................................
<taxonomicName id="C6986CCEFFB6424BFA97429CFA39F85E" box="[1364,1436,1967,1988]" class="Insecta" family="Formicidae" genus="Ponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="33ECCB5FFFB6424BFA97429CFA39F85E" bold="true" box="[1364,1436,1967,1988]" italics="true" pageId="5" pageNumber="452">sikorae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="55E747C5FFB54248FF54479FFB5AFC9A" httpUri="https://zenodo.org/record/247519/files/figure.png" pageId="6" pageNumber="453" targetBox="[151,1434,193,645]" targetPageId="6">
<paragraph id="0127174DFFB54248FF54479FFB5AFC9A" blockId="6.[151,1436,684,768]" pageId="6" pageNumber="453">
<emphasis id="33ECCB5FFFB54248FF54479FFE99FD5B" bold="true" box="[151,316,684,707]" pageId="6" pageNumber="453">FIGURES 45.</emphasis>
Petiole and gaster in dorsal view; Fig. 4: posterior surface of petiole with abundant pubescence and anterior face of first gastral segment forming shallow cavity (CASENT0195184); Fig. 5: abundant pubescence absent on posterior surface of petiole and anterior face of first gastral segment straight without shallow cavity (CASENT0231239).
</paragraph>
</caption>
<caption id="55E747C5FFB54248FF544029FE63FAF4" httpUri="https://zenodo.org/record/247520/files/figure.png" pageId="6" pageNumber="453" targetBox="[153,1433,844,1259]" targetPageId="6">
<paragraph id="0127174DFFB54248FF544029FE63FAF4" blockId="6.[151,1436,1306,1390]" pageId="6" pageNumber="453">
<emphasis id="33ECCB5FFFB54248FF544029FE9EFAAA" bold="true" box="[151,315,1306,1329]" pageId="6" pageNumber="453">FIGURES 67.</emphasis>
Lateral view showing the sculpture of lateral portion of head and petiole; Fig. 6: head and petiole smooth and shining (CASENT0050330: holotype); Fig. 7: head densely and finely reticulate-rugulose, petiolar node densely and finely punctate (CASENT0195184).
</paragraph>
</caption>
<caption id="55E747C5FFB54248FF544288FDD4F86A" httpUri="https://zenodo.org/record/247521/files/figure.png" pageId="6" pageNumber="453" targetBox="[153,1436,1464,1943]" targetPageId="6">
<paragraph id="0127174DFFB54248FF544288FDD4F86A" blockId="6.[151,1436,1979,2033]" pageId="6" pageNumber="453">
<emphasis id="33ECCB5FFFB54248FF544288FE9EF84B" bold="true" box="[151,315,1979,2002]" pageId="6" pageNumber="453">FIGURES 89.</emphasis>
Mesosoma in dorsal view; Fig. 8: metanotal groove deeply impressed (CASENT0034340: holotype); Fig. 9: metanotal groove lacking (CASENT0497667).
</paragraph>
</caption>
<caption id="55E747C5FFB44249FF5447D3FDB1FC8F" httpUri="https://zenodo.org/record/247522/files/figure.png" pageId="7" pageNumber="454" targetBox="[153,1436,195,700]" targetPageId="7">
<paragraph id="0127174DFFB44249FF5447D3FDB1FC8F" blockId="7.[151,1436,736,789]" pageId="7" pageNumber="454">
<emphasis id="33ECCB5FFFB44249FF5447D3FEF6FD6F" bold="true" box="[151,339,736,759]" pageId="7" pageNumber="454">FIGURES 1011.</emphasis>
Form of petiole in profile; Fig. 10: anteroposteriorly flattened (CASENT0034340); Fig. 11: nodiform about as long as broad (CASENT0497202).
</paragraph>
</caption>
<keyStep id="BA6C0FE8FFB44249FF54460CFA39FB99" pageId="7" pageNumber="454">
<paragraph id="0127174DFFB44249FF54460CFA3EFC31" blockId="7.[151,1436,831,1028]" pageId="7" pageNumber="454">
<keyLead id="BA69B478FFB44249FF54460CFA3EFC31" pageId="7" pageNumber="454">
6. Posterolateral margin of propodeum bordered with broad lamellae which project as tubercles at about the level of propodeal spiracle; antennal scape and outer surface of each tibia covered with erect, slender hairs (
<figureCitation id="99A30BC8FFB44249FBEC466FFBD3FCEB" box="[1071,1142,860,881]" captionStart="FIGURES 12 13" captionStartId="7.[151,264,1595,1618]" captionTargetBox="[151,1436,1095,1556]" captionTargetId="figure@7.[151,1436,1095,1571]" captionTargetPageId="7" captionText="FIGURES 12 13. Hairs on the outer surface of tibiae and the width of lamellae on posterior margin of propodeum; Fig. 12: outer surface of each tibia covered with erect hairs and posterior margin of propodeum with broad lamellae that project as a tubercle at about the level of the propodeal spiracle (CASENT 0166009: holotype); Fig. 13: outer surface of tibiae without erect hairs, posterior margin of propodeum with narrow lamellae that form an obtuse angle at about the level of propodeal spiracle (CASENT 0034340)." httpUri="https://zenodo.org/record/247523/files/figure.png" pageId="7" pageNumber="454">Fig. 12</figureCitation>
); dorsum of head near the posterior margin and rest of body covered with numerous slender long hairs; posterior cephalic margin more or less straight...................................................................................................
<taxonomicName id="C6986CCEFFB44249FA8746A5FA3EFC31" box="[1348,1435,918,939]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="454" phylum="Arthropoda" rank="species" species="gorogota">
<emphasis id="33ECCB5FFFB44249FA8746A5FA3EFC31" bold="true" box="[1348,1435,918,939]" italics="true" pageId="7" pageNumber="454">gorogota</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0127174DFFB44249FF544687FA39FB99" blockId="7.[151,1436,831,1028]" pageId="7" pageNumber="454">
<keyLead id="BA69B478FFB44249FF544687FA39FB99" pageId="7" pageNumber="454">
- Posterolateral margin of propodeum bordered with narrow lamellae which form an obtuse angle at about the level of propodeal spiracle; antennal scape and outer surface of each tibia lacking erect, slender hairs (
<figureCitation id="99A30BC8FFB44249FBC446E2FBEAFC7C" box="[1031,1103,977,998]" captionStart="FIGURES 12 13" captionStartId="7.[151,264,1595,1618]" captionTargetBox="[151,1436,1095,1556]" captionTargetId="figure@7.[151,1436,1095,1571]" captionTargetPageId="7" captionText="FIGURES 12 13. Hairs on the outer surface of tibiae and the width of lamellae on posterior margin of propodeum; Fig. 12: outer surface of each tibia covered with erect hairs and posterior margin of propodeum with broad lamellae that project as a tubercle at about the level of the propodeal spiracle (CASENT 0166009: holotype); Fig. 13: outer surface of tibiae without erect hairs, posterior margin of propodeum with narrow lamellae that form an obtuse angle at about the level of propodeal spiracle (CASENT 0034340)." httpUri="https://zenodo.org/record/247523/files/figure.png" pageId="7" pageNumber="454">Fig. 13</figureCitation>
); dorsum of head and rest of body covered with few erect, slender hairs; posterior cephalic margin concave......................................
<taxonomicName id="C6986CCEFFB44249FA9F46DDFA39FB99" box="[1372,1436,1006,1027]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="454" phylum="Arthropoda" rank="species" species="maeva">
<emphasis id="33ECCB5FFFB44249FA9F46DDFA39FB99" bold="true" box="[1372,1436,1006,1027]" italics="true" pageId="7" pageNumber="454">maeva</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="55E747C5FFB44249FF544308FECFF957" httpUri="https://zenodo.org/record/247523/files/figure.png" pageId="7" pageNumber="454" targetBox="[151,1436,1095,1556]" targetPageId="7">
<paragraph id="0127174DFFB44249FF544308FECFF957" blockId="7.[151,1436,1595,1741]" pageId="7" pageNumber="454">
<emphasis id="33ECCB5FFFB44249FF544308FEF0F9CA" bold="true" box="[151,341,1595,1618]" pageId="7" pageNumber="454">FIGURES 1213.</emphasis>
Hairs on the outer surface of tibiae and the width of lamellae on posterior margin of propodeum; Fig. 12: outer surface of each tibia covered with erect hairs and posterior margin of propodeum with broad lamellae that project as a tubercle at about the level of the propodeal spiracle (CASENT0166009: holotype); Fig. 13: outer surface of tibiae without erect hairs, posterior margin of propodeum with narrow lamellae that form an obtuse angle at about the level of propodeal spiracle (CASENT0034340).
</paragraph>
</caption>
<keyStep id="BA6C0FE8FFB44249FF5443CBFA3EF8FF" pageId="7" pageNumber="454">
<paragraph id="0127174DFFB44249FF5443CBFA3EF8B1" blockId="7.[151,1437,1784,2011]" pageId="7" pageNumber="454">
<keyLead id="BA69B478FFB44249FF5443CBFA3EF8B1" pageId="7" pageNumber="454">
7. Smaller species (HW: 1.15, HL&lt;1.5, WL: 1.78); eyes smaller, with seven to eight ommatidia; metanotal groove absent (
<figureCitation id="99A30BC8FFB44249FAB643CBFF59F8B1" captionStart="FIGURES 14 15" captionStartId="8.[151,264,773,796]" captionTargetBox="[152,1434,203,733]" captionTargetId="figure@8.[152,1435,203,743]" captionTargetPageId="8" captionText="FIGURES 14 15. Mesosoma in dorsal view; Fig. 14: metanotal groove absent (CASENT 0497667); Fig. 15: metanotal groove as a dotted line (CASENT 0231239: holotype)." httpUri="https://zenodo.org/record/247524/files/figure.png" pageId="7" pageNumber="454">Fig. 14</figureCitation>
); mesopleural suture lacking......................................................................
<taxonomicName id="C6986CCEFFB44249FAA64225FA3EF8B1" box="[1381,1435,1814,1835]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="454" phylum="Arthropoda" rank="species" species="mialy">
<emphasis id="33ECCB5FFFB44249FAA64225FA3EF8B1" bold="true" box="[1381,1435,1814,1835]" italics="true" pageId="7" pageNumber="454">mialy</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0127174DFFB44249FF544200FA3EF8FF" blockId="7.[151,1437,1784,2011]" pageId="7" pageNumber="454">
<keyLead id="BA69B478FFB44249FF544200FA3EF8FF" pageId="7" pageNumber="454">
- Larger species (HW: 1.44, HL: 1.62, WL: 2.34); eyes larger, with 13 ommatidia; metanotal groove present as a dotted line (
<figureCitation id="99A30BC8FFB44249FAB64200FF5EF8FF" captionStart="FIGURES 14 15" captionStartId="8.[151,264,773,796]" captionTargetBox="[152,1434,203,733]" captionTargetId="figure@8.[152,1435,203,743]" captionTargetPageId="8" captionText="FIGURES 14 15. Mesosoma in dorsal view; Fig. 14: metanotal groove absent (CASENT 0497667); Fig. 15: metanotal groove as a dotted line (CASENT 0231239: holotype)." httpUri="https://zenodo.org/record/247524/files/figure.png" pageId="7" pageNumber="454">Fig. 15</figureCitation>
); mesopleural suture present........................................................................
<taxonomicName id="C6986CCEFFB44249FAAD4263FA3EF8FF" box="[1390,1435,1872,1893]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="454" phylum="Arthropoda" rank="species" species="nosy">
<emphasis id="33ECCB5FFFB44249FAAD4263FA3EF8FF" bold="true" box="[1390,1435,1872,1893]" italics="true" pageId="7" pageNumber="454">nosy</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFB44249FF54425DFA39F841" pageId="7" pageNumber="454">
<paragraph id="0127174DFFB44249FF54425DFA39F83A" blockId="7.[151,1437,1784,2011]" pageId="7" pageNumber="454">
<keyLead id="BA69B478FFB44249FF54425DFA39F83A" pageId="7" pageNumber="454">
8. Anterior surface of the third abdominal segment concave (
<figureCitation id="99A30BC8FFB44249FCCE425DFCF4F819" box="[781,849,1902,1923]" captionStart="FIGURES 16 17" captionStartId="8.[151,264,1509,1532]" captionTargetBox="[151,1436,889,1470]" captionTargetId="figure@8.[151,1436,889,1480]" captionTargetPageId="8" captionText="FIGURES 16 17. First gastral segment in dorsal view; Fig. 16: anterior face forming shallow cavity (CASENT 0317589: holotype); Fig. 17: anterior face without shallow cavity (CASENT 0247289)." httpUri="https://zenodo.org/record/247525/files/figure.png" pageId="7" pageNumber="454">Fig. 16</figureCitation>
), forming a shallow cavity to lodge the median portion of the posterior surface of petiole node.....................................................................
<taxonomicName id="C6986CCEFFB44249FA9A42B8FA39F83A" box="[1369,1436,1931,1952]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="454" phylum="Arthropoda" rank="species" species="agnivo">
<emphasis id="33ECCB5FFFB44249FA9A42B8FA39F83A" bold="true" box="[1369,1436,1931,1952]" italics="true" pageId="7" pageNumber="454">agnivo</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0127174DFFB44249FF54429BFA39F841" blockId="7.[151,1437,1784,2011]" pageId="7" pageNumber="454">
<keyLead id="BA69B478FFB44249FF54429BFA39F841" pageId="7" pageNumber="454">
- Anterior surface of the third abdominal segment straight, not forming a shallow cavity to lodge the posterior surface of petiole node (
<figureCitation id="99A30BC8FFB44249FEDF42F5FEC7F841" box="[284,354,1990,2011]" captionStart="FIGURES 16 17" captionStartId="8.[151,264,1509,1532]" captionTargetBox="[151,1436,889,1470]" captionTargetId="figure@8.[151,1436,889,1480]" captionTargetPageId="8" captionText="FIGURES 16 17. First gastral segment in dorsal view; Fig. 16: anterior face forming shallow cavity (CASENT 0317589: holotype); Fig. 17: anterior face without shallow cavity (CASENT 0247289)." httpUri="https://zenodo.org/record/247525/files/figure.png" pageId="7" pageNumber="454">Fig. 17</figureCitation>
)........................................................................................ 9
</keyLead>
</paragraph>
</keyStep>
<caption id="55E747C5FFBB4246FF544636FDCAFCA0" httpUri="https://zenodo.org/record/247524/files/figure.png" pageId="8" pageNumber="455" targetBox="[152,1434,203,733]" targetPageId="8">
<paragraph id="0127174DFFBB4246FF544636FDCAFCA0" blockId="8.[151,1436,773,826]" pageId="8" pageNumber="455">
<emphasis id="33ECCB5FFFBB4246FF544636FEF6FC80" bold="true" box="[151,339,773,796]" pageId="8" pageNumber="455">FIGURES 1415.</emphasis>
Mesosoma in dorsal view; Fig. 14: metanotal groove absent (CASENT0497667); Fig. 15: metanotal groove as a dotted line (CASENT0231239: holotype).
</paragraph>
</caption>
<caption id="55E747C5FFBB4246FF5440D6FC05F981" httpUri="https://zenodo.org/record/247525/files/figure.png" pageId="8" pageNumber="455" targetBox="[151,1436,889,1470]" targetPageId="8">
<paragraph id="0127174DFFBB4246FF5440D6FC05F981" blockId="8.[151,1437,1509,1563]" pageId="8" pageNumber="455">
<emphasis id="33ECCB5FFFBB4246FF5440D6FEF2FA61" bold="true" box="[151,343,1509,1532]" pageId="8" pageNumber="455">FIGURES 1617.</emphasis>
First gastral segment in dorsal view; Fig. 16: anterior face forming shallow cavity (CASENT0317589: holotype); Fig. 17: anterior face without shallow cavity (CASENT0247289).
</paragraph>
</caption>
<keyStep id="BA6C0FE8FFBB4246FF544374FA39F9E3" pageId="8" pageNumber="455">
<paragraph id="0127174DFFBB4246FF544374FA3EF9C6" blockId="8.[151,1436,1607,1921]" box="[151,1435,1607,1628]" pageId="8" pageNumber="455">
<keyLead id="BA69B478FFBB4246FF544374FA3EF9C6" box="[151,1435,1607,1628]" pageId="8" pageNumber="455">
9. Posterolateral margins of petiole node without a series of sharp teeth or tubercles (
<figureCitation id="99A30BC8FFBB4246FC224374FB82F9C6" box="[993,1063,1607,1628]" captionStart="FIGURES 18 19" captionStartId="9.[151,264,713,736]" captionTargetBox="[151,1436,195,678]" captionTargetId="figure@9.[151,1436,195,692]" captionTargetPageId="9" captionText="FIGURES 18 19. Petiolar node in profile; Fig. 18: posterolateral margin without a series of sharp teeth or tubercles (CASENT 0043301: holotype); Fig. 19: posterolateral margin armed with a series of sharp teeth or tubercles (CASENT 0317590: holotype)." httpUri="https://zenodo.org/record/247526/files/figure.png" pageId="8" pageNumber="455">Fig. 18</figureCitation>
); dorsum of node not tubercuate.... 10
</keyLead>
</paragraph>
<paragraph id="0127174DFFBB4246FF544357FA39F9E3" blockId="8.[151,1436,1607,1921]" box="[151,1436,1636,1657]" pageId="8" pageNumber="455">
<keyLead id="BA69B478FFBB4246FF544357FA39F9E3" box="[151,1436,1636,1657]" pageId="8" pageNumber="455">
- Posterolateral margins of petiole node armed with a series of sharp teeth or tubercles (
<figureCitation id="99A30BC8FFBB4246FBC74357FBECF9E3" box="[1028,1097,1636,1657]" captionStart="FIGURES 18 19" captionStartId="9.[151,264,713,736]" captionTargetBox="[151,1436,195,678]" captionTargetId="figure@9.[151,1436,195,692]" captionTargetPageId="9" captionText="FIGURES 18 19. Petiolar node in profile; Fig. 18: posterolateral margin without a series of sharp teeth or tubercles (CASENT 0043301: holotype); Fig. 19: posterolateral margin armed with a series of sharp teeth or tubercles (CASENT 0317590: holotype)." httpUri="https://zenodo.org/record/247526/files/figure.png" pageId="8" pageNumber="455">Fig. 19</figureCitation>
); dorsum of node tuberculate... 12
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</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFBB4246FF5443B2FA3EF974" pageId="8" pageNumber="455">
<paragraph id="0127174DFFBB4246FF5443B2FA3EF92E" blockId="8.[151,1436,1607,1921]" pageId="8" pageNumber="455">
<keyLead id="BA69B478FFBB4246FF5443B2FA3EF92E" pageId="8" pageNumber="455">
10. In dorsal view, petiole node almost twice as broad as long (
<figureCitation id="99A30BC8FFBB4246FCE543B2FCCAF90C" box="[806,879,1665,1686]" captionStart="FIGURES 20 21" captionStartId="9.[151,264,1346,1369]" captionTargetBox="[151,1436,854,1306]" captionTargetId="figure@9.[151,1436,854,1319]" captionTargetPageId="9" captionText="FIGURES 20 21. Petiole in dorsal view; Fig. 20: node approximately twice broader than long (CASENT 0043301); Fig. 21: node nearly as broad as long (CASENT 0045619: holotype)." httpUri="https://zenodo.org/record/247527/files/figure.png" pageId="8" pageNumber="455">Fig. 20</figureCitation>
); in lateral view, subpetiolar process simple with only an anterior triangular lobe (
<figureCitation id="99A30BC8FFBB4246FE0843ACFDB5F92E" box="[459,528,1695,1716]" captionStart="FIGURES 18 19" captionStartId="9.[151,264,713,736]" captionTargetBox="[151,1436,195,678]" captionTargetId="figure@9.[151,1436,195,692]" captionTargetPageId="9" captionText="FIGURES 18 19. Petiolar node in profile; Fig. 18: posterolateral margin without a series of sharp teeth or tubercles (CASENT 0043301: holotype); Fig. 19: posterolateral margin armed with a series of sharp teeth or tubercles (CASENT 0317590: holotype)." httpUri="https://zenodo.org/record/247526/files/figure.png" pageId="8" pageNumber="455">Fig. 18</figureCitation>
)....................................................................
<taxonomicName id="C6986CCEFFBB4246FA8D43ACFA3EF92E" box="[1358,1435,1695,1716]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="455" phylum="Arthropoda" rank="species" species="daraina">
<emphasis id="33ECCB5FFFBB4246FA8D43ACFA3EF92E" bold="true" box="[1358,1435,1695,1716]" italics="true" pageId="8" pageNumber="455">daraina</emphasis>
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</paragraph>
<paragraph id="0127174DFFBB4246FF54438FFA3EF974" blockId="8.[151,1436,1607,1921]" pageId="8" pageNumber="455">
<keyLead id="BA69B478FFBB4246FF54438FFA3EF974" pageId="8" pageNumber="455">
- In dorsal view, petiole nodenearly as broad as long (
<figureCitation id="99A30BC8FFBB4246FD13438FFCB2F94B" box="[720,791,1724,1745]" captionStart="FIGURES 20 21" captionStartId="9.[151,264,1346,1369]" captionTargetBox="[151,1436,854,1306]" captionTargetId="figure@9.[151,1436,854,1319]" captionTargetPageId="9" captionText="FIGURES 20 21. Petiole in dorsal view; Fig. 20: node approximately twice broader than long (CASENT 0043301); Fig. 21: node nearly as broad as long (CASENT 0045619: holotype)." httpUri="https://zenodo.org/record/247527/files/figure.png" pageId="8" pageNumber="455">Fig. 21</figureCitation>
); in lateral view subpetiolar process with an anterior hook-like process and a posterior subtriangular process separated by a concavity (
<figureCitation id="99A30BC8FFBB4246FCAC43EAFC10F974" box="[879,949,1753,1774]" captionStart="FIGURES 55 57" captionStartId="26.[151,264,1788,1811]" captionTargetBox="[151,1436,503,1748]" captionTargetId="figure@26.[151,1436,503,1764]" captionTargetPageId="26" captionText="FIGURES 55 57. Worker caste of Pachycondyla rovana: lateral and dorsal view of body and full-face view of head, CASENT 0247289." httpUri="https://zenodo.org/record/247540/files/figure.png" pageId="8" pageNumber="455">Fig. 55</figureCitation>
)...................................... 11
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</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFBB4246FF5443C4FA3EF81B" pageId="8" pageNumber="455">
<paragraph id="0127174DFFBB4246FF5443C4FA39F8DC" blockId="8.[151,1436,1607,1921]" pageId="8" pageNumber="455">
<keyLead id="BA69B478FFBB4246FF5443C4FA39F8DC" pageId="8" pageNumber="455">
11. Larger ant (HW: 1.221.32, WL: 1.852.05); mesopleuron and lower half of the side of propodeum mostly smooth and shiny with scattered small punctures in the center and fine rugulae at the junction of anepisternum and katepisternum and the lower portion of the metapleuron near the opening of the metapleural gland (
<figureCitation id="99A30BC8FFBB4246FC474202FC6CF8DC" box="[900,969,1841,1862]" captionStart="FIGURES 22 23" captionStartId="9.[151,264,1940,1963]" captionTargetBox="[151,1436,1466,1903]" captionTargetId="figure@9.[151,1436,1466,1918]" captionTargetPageId="9" captionText="FIGURES 22 23. Sculpture of the lateral surface of mesosoma; Fig. 22: smooth and shiny mesopleuron and metapleuron with scattered small punctures (CASENT 0045619); Fig. 23: densely rugulose mesopleuron and metapleuron superimposed with punctures (CASENT 0247289)." httpUri="https://zenodo.org/record/247528/files/figure.png" pageId="8" pageNumber="455">Fig. 22</figureCitation>
)..................................
<taxonomicName id="C6986CCEFFBB4246FAB24202FA39F8DC" box="[1393,1436,1841,1862]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="455" phylum="Arthropoda" rank="species" species="zoro">
<emphasis id="33ECCB5FFFBB4246FAB24202FA39F8DC" bold="true" box="[1393,1436,1841,1862]" italics="true" pageId="8" pageNumber="455">zoro</emphasis>
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</paragraph>
<paragraph id="0127174DFFBB4246FF54427CFA3EF81B" blockId="8.[151,1436,1607,1921]" pageId="8" pageNumber="455">
<keyLead id="BA69B478FFBB4246FF54427CFA3EF81B" pageId="8" pageNumber="455">
- Smaller species (HW: 1.011.14, WL: 1.581.77); mesopleuron and lower half of the side of propodeum densely rugulose and superimposed with punctures or foveolae (
<figureCitation id="99A30BC8FFBB4246FDBA425FFD1AF81B" box="[633,703,1900,1921]" captionStart="FIGURES 22 23" captionStartId="9.[151,264,1940,1963]" captionTargetBox="[151,1436,1466,1903]" captionTargetId="figure@9.[151,1436,1466,1918]" captionTargetPageId="9" captionText="FIGURES 22 23. Sculpture of the lateral surface of mesosoma; Fig. 22: smooth and shiny mesopleuron and metapleuron with scattered small punctures (CASENT 0045619); Fig. 23: densely rugulose mesopleuron and metapleuron superimposed with punctures (CASENT 0247289)." httpUri="https://zenodo.org/record/247528/files/figure.png" pageId="8" pageNumber="455">Fig. 23</figureCitation>
)......................................................
<taxonomicName id="C6986CCEFFBB4246FA95425FFA3EF81B" box="[1366,1435,1900,1921]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="455" phylum="Arthropoda" rank="species" species="rovana">
<emphasis id="33ECCB5FFFBB4246FA95425FFA3EF81B" bold="true" box="[1366,1435,1900,1921]" italics="true" pageId="8" pageNumber="455">rovana</emphasis>
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</paragraph>
</keyStep>
<caption id="55E747C5FFBA4247FF5447FAFE75FC87" httpUri="https://zenodo.org/record/247526/files/figure.png" pageId="9" pageNumber="456" targetBox="[151,1436,195,678]" targetPageId="9">
<paragraph id="0127174DFFBA4247FF5447FAFE75FC87" blockId="9.[151,1436,713,797]" pageId="9" pageNumber="456">
<emphasis id="33ECCB5FFFBA4247FF5447FAFEFFFD45" bold="true" box="[151,346,713,736]" pageId="9" pageNumber="456">FIGURES 1819.</emphasis>
Petiolar node in profile; Fig. 18: posterolateral margin without a series of sharp teeth or tubercles (CASENT0043301: holotype); Fig. 19: posterolateral margin armed with a series of sharp teeth or tubercles (CASENT0317590: holotype).
</paragraph>
</caption>
<caption id="55E747C5FFBA4247FF544071FD52FAED" httpUri="https://zenodo.org/record/247527/files/figure.png" pageId="9" pageNumber="456" targetBox="[151,1436,854,1306]" targetPageId="9">
<paragraph id="0127174DFFBA4247FF544071FD52FAED" blockId="9.[151,1436,1346,1399]" pageId="9" pageNumber="456">
<emphasis id="33ECCB5FFFBA4247FF544071FEF0FAC2" bold="true" box="[151,341,1346,1369]" pageId="9" pageNumber="456">FIGURES 2021.</emphasis>
Petiole in dorsal view; Fig. 20: node approximately twice broader than long (CASENT0043301); Fig. 21: node nearly as broad as long (CASENT0045619: holotype).
</paragraph>
</caption>
<caption id="55E747C5FFBA4247FF5442A7FE76F872" httpUri="https://zenodo.org/record/247528/files/figure.png" pageId="9" pageNumber="456" targetBox="[151,1436,1466,1903]" targetPageId="9">
<paragraph id="0127174DFFBA4247FF5442A7FE76F872" blockId="9.[151,1436,1940,2024]" pageId="9" pageNumber="456">
<emphasis id="33ECCB5FFFBA4247FF5442A7FEF1F833" bold="true" box="[151,340,1940,1963]" pageId="9" pageNumber="456">FIGURES 2223.</emphasis>
Sculpture of the lateral surface of mesosoma; Fig. 22: smooth and shiny mesopleuron and metapleuron with scattered small punctures (CASENT0045619); Fig. 23: densely rugulose mesopleuron and metapleuron superimposed with punctures (CASENT0247289).
</paragraph>
</caption>
<keyStep id="BA6C0FE8FFB94244FF5445A4FA3EFEA5" pageId="10" pageNumber="457">
<paragraph id="0127174DFFB94244FF5445A4FA3EFF7D" blockId="10.[151,1436,151,583]" pageId="10" pageNumber="457">
<keyLead id="BA69B478FFB94244FF5445A4FA3EFF7D" pageId="10" pageNumber="457">
12. Second gastral (fourth abdominal) tergite punctate to sparsely punctuate; pubescence absent or reduced in abundance on first and second gastral segments, distance between two adjacent pubescences generally equal to or greater than the length of the pubescence itself (
<figureCitation id="99A30BC8FFB94244FE5145E1FE7DFF7D" box="[402,472,210,231]" captionStart="FIGURES 24 25" captionStartId="10.[151,264,1166,1189]" captionTargetBox="[151,1436,652,1129]" captionTargetId="figure@10.[151,1436,652,1142]" captionTargetPageId="10" captionText="FIGURES 24 25. Sculpture and pubescence of the gaster; Fig. 24: first and second gastral tergites with few pubescence and second gastral tergite punctate to sparsely punctate (CASENT 0317590); Fig. 25: first and second gastral tergites with abundant pubescence and second gastral tergite densely punctate (CASENT 0050524)." httpUri="https://zenodo.org/record/247529/files/figure.png" pageId="10" pageNumber="457">Fig. 24</figureCitation>
)............................................................................. 13
</keyLead>
</paragraph>
<paragraph id="0127174DFFB94244FF5445DCFA3EFEA5" blockId="10.[151,1436,151,583]" pageId="10" pageNumber="457">
<keyLead id="BA69B478FFB94244FF5445DCFA3EFEA5" pageId="10" pageNumber="457">
- Second gastral (fourth abdominal) tergite densely punctate, with small punctures between larger ones; first and second gastral segments with abundant pubescence, distance between two adjacent pubescences always less than the length of the pubescence itself (
<figureCitation id="99A30BC8FFB94244FEDD4419FEC1FEA5" box="[286,356,298,319]" captionStart="FIGURES 24 25" captionStartId="10.[151,264,1166,1189]" captionTargetBox="[151,1436,652,1129]" captionTargetId="figure@10.[151,1436,652,1142]" captionTargetPageId="10" captionText="FIGURES 24 25. Sculpture and pubescence of the gaster; Fig. 24: first and second gastral tergites with few pubescence and second gastral tergite punctate to sparsely punctate (CASENT 0317590); Fig. 25: first and second gastral tergites with abundant pubescence and second gastral tergite densely punctate (CASENT 0050524)." httpUri="https://zenodo.org/record/247529/files/figure.png" pageId="10" pageNumber="457">Fig. 25</figureCitation>
).................................................................................
<taxonomicName id="C6986CCEFFB94244FAFD4419FA3EFEA5" box="[1342,1435,298,319]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="457" phylum="Arthropoda" rank="species" species="vohitravo">
<emphasis id="33ECCB5FFFB94244FAFD4419FA3EFEA5" bold="true" box="[1342,1435,298,319]" italics="true" pageId="10" pageNumber="457">vohitravo</emphasis>
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</paragraph>
</keyStep>
<keyStep id="BA6C0FE8FFB94244FF544474FA3EFDDD" pageId="10" pageNumber="457">
<paragraph id="0127174DFFB94244FF544474FA39FE2E" blockId="10.[151,1436,151,583]" pageId="10" pageNumber="457">
<keyLead id="BA69B478FFB94244FF544474FA39FE2E" pageId="10" pageNumber="457">
13. Mesopleural suture distinctly continuous and narrowly impressed (
<figureCitation id="99A30BC8FFB94244FCB54474FC64FEC6" box="[886,961,327,348]" captionStart="FIGURES 26 27" captionStartId="10.[151,264,1846,1869]" captionTargetBox="[151,1436,1321,1812]" captionTargetId="figure@10.[151,1436,1321,1825]" captionTargetPageId="10" captionText="FIGURES 26 27. Mesosoma in lateral view; Fig. 26: mesopleural suture distinctly continuous and narrowly impressed (CASENT 0317590); Fig. 27: mesopleural suture indistinct (CASENT 0162668: holotype)." httpUri="https://zenodo.org/record/247530/files/figure.png" pageId="10" pageNumber="457">Fig. 26</figureCitation>
); head densely and finely reticulate-rugulose or reticulate-punctate behind the level of eye; with head in full-face view, the lateral borders at the level of anterior margin of eye not strongly curving towards the base of mandibles (
<figureCitation id="99A30BC8FFB94244FD1944B1FC87FE0D" box="[730,802,386,407]" captionStart="FIGURES 31 33" captionStartId="13.[151,264,1839,1862]" captionTargetBox="[151,1436,375,1807]" captionTargetId="figure@13.[151,1436,375,1823]" captionTargetPageId="13" captionText="FIGURES 31 33. Worker caste of Pachycondyla antsiraka: lateral and dorsal view of body and full-face view of head, holotype CASENT 0317590." httpUri="https://zenodo.org/record/247532/files/figure.png" pageId="10" pageNumber="457">Fig. 32</figureCitation>
); in profile, distance between eyes and base of mandibles roughly three times or more than the maximum diameter of eye.................................................
<taxonomicName id="C6986CCEFFB94244FAFC44ACFA39FE2E" box="[1343,1436,415,436]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="457" phylum="Arthropoda" rank="species" species="antsiraka">
<emphasis id="33ECCB5FFFB94244FAFC44ACFA39FE2E" bold="true" box="[1343,1436,415,436]" italics="true" pageId="10" pageNumber="457">antsiraka</emphasis>
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</keyLead>
</paragraph>
<paragraph id="0127174DFFB94244FF54448EFA3EFDDD" blockId="10.[151,1436,151,583]" pageId="10" pageNumber="457">
<keyLead id="BA69B478FFB94244FF54448EFA3EFDDD" pageId="10" pageNumber="457">
- Mesopleural suture indistinct, with broad and discontinuous concavities or impressions in this region, but no obvious suture (
<figureCitation id="99A30BC8FFB94244FF2B44E9FE95FE75" box="[232,304,474,495]" captionStart="FIGURES 26 27" captionStartId="10.[151,264,1846,1869]" captionTargetBox="[151,1436,1321,1812]" captionTargetId="figure@10.[151,1436,1321,1825]" captionTargetPageId="10" captionText="FIGURES 26 27. Mesosoma in lateral view; Fig. 26: mesopleural suture distinctly continuous and narrowly impressed (CASENT 0317590); Fig. 27: mesopleural suture indistinct (CASENT 0162668: holotype)." httpUri="https://zenodo.org/record/247530/files/figure.png" pageId="10" pageNumber="457">Fig. 27</figureCitation>
); head mostly smooth and shiny with faintly effaced reticulate-punctate sculpture behind the level of eyes; with head in full-face view, the lateral margins from the level of anterior margin of eye abruptly curving toward the base of mandibles (
<figureCitation id="99A30BC8FFB94244FF2B4726FE95FDB0" box="[232,304,533,554]" captionStart="FIGURES 61 63" captionStartId="30.[151,264,1732,1755]" captionTargetBox="[151,1436,260,1704]" captionTargetId="figure@30.[151,1436,260,1721]" captionTargetPageId="30" captionText="FIGURES 61 63. Worker caste of Pachycondyla tahary: lateral and dorsal view of body and full-face view of head, holotype CASENT 0162668." httpUri="https://zenodo.org/record/247542/files/figure.png" pageId="10" pageNumber="457">Fig. 62</figureCitation>
); in lateral view, distance between eye and base of mandible slightly less than twice the maximum diameter of eye...................................................................................................
<taxonomicName id="C6986CCEFFB94244FA9F4701FA3EFDDD" box="[1372,1435,562,583]" class="Insecta" family="Formicidae" genus="Pachycondyla" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="457" phylum="Arthropoda" rank="species" species="tahary">
<emphasis id="33ECCB5FFFB94244FA9F4701FA3EFDDD" bold="true" box="[1372,1435,562,583]" italics="true" pageId="10" pageNumber="457">tahary</emphasis>
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