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<document id="A7175F374C3B7F9C9F518347925577BB" ID-CLB-Dataset="32249" ID-DOI="10.11646/zootaxa.4313.1.1" ID-GBIF-Dataset="d8cc2cd8-8410-49aa-a75d-7a41d9f52b26" ID-ISSN="1175-5326" ID-Zenodo-Dep="883161" ID-ZooBank="A5Fdfba5-F992-44A8-84C2-1756C943C19B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504182989959" checkinUser="plazi" docAuthor="Bush, Sarah E." docDate="2017" docId="832187E9FF0FFF4BFF7464C8FC09FB84" docLanguage="en" docName="zootaxa.4313.1.1.pdf" docOrigin="Zootaxa 4313 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Titanomessor sexloba Bush 2017, new species" docType="treatment" docVersion="16" lastPageNumber="137" masterDocId="7F18FF91FF8AFFC3FFE36200FFC9FFA3" masterDocTitle="Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key" masterLastPageNumber="443" masterPageNumber="1" pageNumber="134" updateTime="1698467717597" updateUser="ExternalLinkService">
<mods:mods id="659FE553ADA05A1A3F408B13BFB5BF51" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1500BDB2E3082884EACE9452B8D5DCF6">
<mods:title id="64D711ED6B553A4D61510601EFC57A37">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
</mods:titleInfo>
<mods:name id="CEC28A7374ED6B2BFDB631F850176FEA" type="personal">
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<mods:roleTerm id="5BE915EE75CA7E822B4F3262A06F9BE7">Author</mods:roleTerm>
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<mods:namePart id="9301363EB80F81A6806D858DC9141238">Bush, Sarah E.</mods:namePart>
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<mods:title id="0AA21342E6EA1B1AB4AA4FEC7288D7DE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5A177B5321051133EB6AB1E2673D99AB">
<mods:date id="318F34801B52BE91AD3745B8D0388808">2017</mods:date>
<mods:detail id="D6F9C20C4BC84A81E44DBB65DD9EC365" type="pubDate">
<mods:number id="CB277598751D9FAC03D141C1466FF295">2017-08-31</mods:number>
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<mods:number id="23A20A5039799722F9979916DE400E74">4313</mods:number>
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<mods:classification id="DEB778300D0F32B19E4E7CD656D97D93">journal article</mods:classification>
<mods:identifier id="0D5F761709A9631CF23C14183806154C" type="CLB-Dataset">32249</mods:identifier>
<mods:identifier id="CBD0707BB47FADC330E0B685EF4312A2" type="DOI">10.11646/zootaxa.4313.1.1</mods:identifier>
<mods:identifier id="80EA12AD887DC99C3B7D72F97455BBBB" type="GBIF-Dataset">d8cc2cd8-8410-49aa-a75d-7a41d9f52b26</mods:identifier>
<mods:identifier id="7DDA9180A9E2AE3377FC7A163269E6C8" type="ISSN">1175-5326</mods:identifier>
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<treatment id="832187E9FF0FFF4BFF7464C8FC09FB84" ID-DOI="http://doi.org/10.5281/zenodo.5296945" ID-GBIF-Taxon="133100181" ID-Zenodo-Dep="5296945" LSID="urn:lsid:plazi:treatment:832187E9FF0FFF4BFF7464C8FC09FB84" httpUri="http://treatment.plazi.org/id/832187E9FF0FFF4BFF7464C8FC09FB84" lastPageId="136" lastPageNumber="137" pageId="133" pageNumber="134">
<subSubSection id="43926574FF0FFF46FF7464C8FE8AF8A0" pageId="133" pageNumber="134" type="nomenclature">
<paragraph id="0B3736FFFF0FFF46FF7464C8FCFAF941" blockId="133.[151,819,1736,1795]" box="[151,819,1736,1762]" pageId="133" pageNumber="134">
<heading id="507F8193FF0FFF46FF7464C8FCFAF941" bold="true" box="[151,819,1736,1762]" fontSize="11" level="1" pageId="133" pageNumber="134" reason="1">
<emphasis id="39FCEAEDFF0FFF46FF7464C8FCFAF941" bold="true" box="[151,819,1736,1762]" pageId="133" pageNumber="134">
<taxonomicName id="CC884D7CFF0FFF46FF7464C8FD5FF941" authority="Gustafsson &amp; Bush" authorityName="Bush" authorityYear="2017" box="[151,662,1736,1762]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sexloba" status="sp. nov.">
<emphasis id="39FCEAEDFF0FFF46FF7464C8FE55F941" bold="true" box="[151,412,1736,1762]" italics="true" pageId="133" pageNumber="134">Titanomessor sexloba</emphasis>
Gustafsson &amp; Bush
</taxonomicName>
,
<taxonomicNameLabel id="22CF5796FF0FFF46FD4064C8FCFAF941" box="[675,819,1736,1762]" pageId="133" pageNumber="134" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B3736FFFF0FFF46FF7464EAFE8AF8A0" blockId="133.[151,819,1736,1795]" box="[151,323,1770,1795]" pageId="133" pageNumber="134">
(
<figureCitation id="93B32A7AFF0FFF46FF7C64EAFEF2F8A0" box="[159,315,1770,1795]" captionStart-0="FIGURES 210 – 211" captionStart-1="FIGURES 212 – 216" captionStartId-0="134.[151,264,1837,1859]" captionStartId-1="135.[151,264,1883,1905]" captionTargetBox-0="[193,1373,229,1811]" captionTargetBox-1="[219,1376,221,1841]" captionTargetId-0="figure@134.[151,1436,193,1815]" captionTargetId-1="figure@135.[191,1396,193,1862]" captionTargetPageId-0="134" captionTargetPageId-1="135" captionText-0="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." captionText-1="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri-0="https://zenodo.org/record/883272/files/figure.png" httpUri-1="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Figs 210–216</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43926574FF0FFF46FF746532FE4BF8C4" pageId="133" pageNumber="134" type="materials_examined">
<paragraph id="0B3736FFFF0FFF46FF746532FE4BF8C4" blockId="133.[151,978,1842,1895]" pageId="133" pageNumber="134">
<materialsCitation id="BBE03CA2FF0FFF46FF746532FE4BF8C4" ID-GBIF-Occurrence="1587279392" country="Uganda" location="Uganda" pageId="133" pageNumber="134" specimenCount="2" typeStatus="holotype">
<emphasis id="39FCEAEDFF0FFF46FF746532FECCF8EB" bold="true" box="[151,261,1842,1864]" pageId="133" pageNumber="134">
<typeStatus id="D433885DFF0FFF46FF746532FF05F8EB" box="[151,204,1842,1864]" pageId="133" pageNumber="134">Type</typeStatus>
host.
</emphasis>
<taxonomicName id="CC884D7CFF0FFF46FEEF6533FD06F8EB" authority="(Cretzschmar, 1829)" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[268,719,1842,1864]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="133" pageNumber="134" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF0FFF46FEEF6533FE37F8EB" box="[268,510,1842,1864]" italics="true" pageId="133" pageNumber="134">Laniarius erythrogaster</emphasis>
(Cretzschmar, 1829)
</taxonomicName>
—black-headed gonolek.
<emphasis id="39FCEAEDFF0FFF46FF746551FEECF8C4" bold="true" box="[151,293,1873,1895]" pageId="133" pageNumber="134">
<typeStatus id="D433885DFF0FFF46FF746551FF05F8C4" box="[151,204,1873,1895]" pageId="133" pageNumber="134">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="739F766FFF0FFF46FECF6551FEB7F8C4" box="[300,382,1873,1895]" name="Uganda" pageId="133" pageNumber="134">Uganda</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43926574FF0FFF46FF746594FB67F856" pageId="133" pageNumber="134" type="diagnosis">
<paragraph id="0B3736FFFF0FFF46FF746594FB67F856" blockId="133.[151,1436,1940,2037]" pageId="133" pageNumber="134">
<emphasis id="39FCEAEDFF0FFF46FF746594FED8F80E" bold="true" box="[151,273,1940,1965]" pageId="133" pageNumber="134">Diagnosis.</emphasis>
<taxonomicName id="CC884D7CFF0FFF46FEFA6595FDC7F80E" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[281,526,1940,1965]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="sexloba" status="sp. nov.">
<emphasis id="39FCEAEDFF0FFF46FEFA6595FDC7F80E" box="[281,526,1940,1965]" italics="true" pageId="133" pageNumber="134">Titanomessor sexloba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF0FFF46FDF66594FD9CF80E" box="[533,597,1940,1965]" pageId="133" pageNumber="134" rank="species">
<emphasis id="39FCEAEDFF0FFF46FDF66594FD9CF80E" bold="true" box="[533,597,1940,1965]" pageId="133" pageNumber="134">n. sp.</emphasis>
</taxonomicNameLabel>
is separated from all other known species of the
<taxonomicName id="CC884D7CFF0FFF46FB676594FB2DF80E" authorityName="Keler" authorityYear="1936" box="[1156,1252,1940,1965]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFF0FFF46FB676594FB2DF80E" box="[1156,1252,1940,1965]" italics="true" pageId="133" pageNumber="134">Brueelia</emphasis>
</taxonomicName>
-complex by the shape and structure of the male parameres (
<figureCitation id="93B32A7AFF0FFF46FD6865B8FD27F872" box="[651,750,1976,2001]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 215</figureCitation>
), the transversally continuous dorsal preantennal suture that does not reach the margin of the head (
<figureCitation id="93B32A7AFF0FFF46FDA965DCFD64F857" box="[586,685,2012,2037]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="133" pageNumber="134">Fig. 212</figureCitation>
), and the sexually dimorphic
<taxonomicName id="CC884D7CFF0FFF46FC1465DDFBD7F857" box="[1015,1054,2013,2036]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="133" pageNumber="134" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF0FFF46FC1465DDFBD7F857" box="[1015,1054,2013,2036]" italics="true" pageId="133" pageNumber="134">mts</emphasis>
</taxonomicName>
(see below).
</paragraph>
</subSubSection>
<caption id="5FF76677FF0CFF45FF74652DFD66F8C1" httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="134" pageNumber="135" targetBox="[193,1373,229,1811]" targetPageId="134">
<paragraph id="0B3736FFFF0CFF45FF74652DFD66F8C1" blockId="134.[151,1435,1837,1890]" pageId="134" pageNumber="135">
<emphasis id="39FCEAEDFF0CFF45FF74652DFEA2F8E0" bold="true" box="[151,363,1837,1859]" pageId="134" pageNumber="135">FIGURES 210–211.</emphasis>
<taxonomicName id="CC884D7CFF0CFF45FE90652EFD86F8E0" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[371,591,1837,1859]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="134" pageNumber="135" phylum="Arthropoda" rank="species" species="sexloba" status="gen. nov.">
<emphasis id="39FCEAEDFF0CFF45FE90652EFD86F8E0" box="[371,591,1837,1859]" italics="true" pageId="134" pageNumber="135">Titanomessor sexloba</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFF0CFF45FDB4652DFD35F8E0" bold="true" box="[599,764,1837,1859]" pageId="134" pageNumber="135">
<taxonomicNameLabel id="22CF5796FF0CFF45FDB4652DFD56F8E0" box="[599,671,1837,1859]" pageId="134" pageNumber="135" rank="genus">n. gen.</taxonomicNameLabel>
&amp; n. sp.
</emphasis>
ex
<taxonomicName id="CC884D7CFF0CFF45FCC1652EFBDCF8E0" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[802,1045,1837,1859]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="134" pageNumber="135" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF0CFF45FCC1652EFBDCF8E0" box="[802,1045,1837,1859]" italics="true" pageId="134" pageNumber="135">Laniarius erythrogaster</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFF0CFF45FBC0652DFB86F8E0" bold="true" box="[1059,1103,1837,1859]" pageId="134" pageNumber="135">210,</emphasis>
male habitus, dorsal and ventral views.
<emphasis id="39FCEAEDFF0CFF45FF3D654CFEC3F8C1" bold="true" box="[222,266,1868,1890]" pageId="134" pageNumber="135">211,</emphasis>
female habitus, dorsal and ventral views.
</paragraph>
</caption>
<caption id="5FF76677FF0DFF44FF74655BFC0AF80D" httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="135" pageNumber="136" targetBox="[219,1376,221,1841]" targetPageId="135">
<paragraph id="0B3736FFFF0DFF44FF74655BFC0AF80D" blockId="135.[151,1436,1883,1966]" pageId="135" pageNumber="136">
<emphasis id="39FCEAEDFF0DFF44FF74655BFEA7F8D2" bold="true" box="[151,366,1883,1905]" pageId="135" pageNumber="136">FIGURES 212–216.</emphasis>
<taxonomicName id="CC884D7CFF0DFF44FE94655CFD9CF8D2" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[375,597,1883,1905]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="sexloba" status="gen. nov.">
<emphasis id="39FCEAEDFF0DFF44FE94655CFD9CF8D2" box="[375,597,1883,1905]" italics="true" pageId="135" pageNumber="136">Titanomessor sexloba</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFF0DFF44FDBD655BFCC0F8D2" bold="true" box="[606,777,1883,1905]" pageId="135" pageNumber="136">
<taxonomicNameLabel id="22CF5796FF0DFF44FDBD655BFD61F8D2" box="[606,680,1883,1905]" pageId="135" pageNumber="136" rank="genus">n. gen.</taxonomicNameLabel>
&amp; n. sp.
</emphasis>
ex
<taxonomicName id="CC884D7CFF0DFF44FCD1655CFBEFF8D2" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[818,1062,1883,1905]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="135" pageNumber="136" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF0DFF44FCD1655CFBEFF8D2" box="[818,1062,1883,1905]" italics="true" pageId="135" pageNumber="136">Laniarius erythrogaster</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFF0DFF44FBD6655BFBA8F8D2" bold="true" box="[1077,1121,1883,1905]" pageId="135" pageNumber="136">212,</emphasis>
male head, dorsal and ventral views.
<emphasis id="39FCEAEDFF0DFF44FF3D657AFEC3F833" bold="true" box="[222,266,1914,1936]" pageId="135" pageNumber="136">213,</emphasis>
male genitalia, dorsal view.
<emphasis id="39FCEAEDFF0DFF44FDC9657AFD9FF833" bold="true" box="[554,598,1914,1936]" pageId="135" pageNumber="136">214,</emphasis>
mesosome, ventral view.
<emphasis id="39FCEAEDFF0DFF44FCB8657AFC4EF833" bold="true" box="[859,903,1914,1936]" pageId="135" pageNumber="136">215,</emphasis>
parameres, dorsal view.
<emphasis id="39FCEAEDFF0DFF44FB63657AFB65F833" bold="true" box="[1152,1196,1914,1936]" pageId="135" pageNumber="136">216,</emphasis>
female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia.
</paragraph>
</caption>
<subSubSection id="43926574FF02FF4BFF246297FC4EFC3A" pageId="136" pageNumber="137" type="description">
<paragraph id="0B3736FFFF02FF4BFF246297FC48FE53" blockId="136.[151,1437,151,1063]" pageId="136" pageNumber="137">
<emphasis id="39FCEAEDFF02FF4BFF246297FE14FF0C" bold="true" box="[199,477,151,176]" pageId="136" pageNumber="137">
Description.
<emphasis id="39FCEAEDFF02FF4BFE826297FE14FF0C" bold="true" box="[353,477,151,176]" italics="true" pageId="136" pageNumber="137">Both sexes</emphasis>
</emphasis>
. Head shape, structure, and chaetotaxy as in genus description and
<figureCitation id="93B32A7AFF02FF4BFB006297FA8CFF13" box="[1251,1349,151,176]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="136" pageNumber="137">Fig. 212</figureCitation>
. Dorsal preantennal suture irregular, often with diffuse margins, but does not seem to reach either
<emphasis id="39FCEAEDFF02FF4BFB4262BBFB01FF77" box="[1185,1224,187,212]" italics="true" pageId="136" pageNumber="137">ads</emphasis>
or
<emphasis id="39FCEAEDFF02FF4BFB1262BBFAE0FF77" box="[1265,1321,187,212]" italics="true" pageId="136" pageNumber="137">dsms</emphasis>
. We have examined both male and female specimens in which the
<taxonomicName id="CC884D7CFF02FF4BFCC262E0FC81FF54" box="[801,840,224,247]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF02FF4BFCC262E0FC81FF54" box="[801,840,224,247]" italics="true" pageId="136" pageNumber="137">mts</emphasis>
</taxonomicName>
chaetotaxy is different from that given in the genus description. Specifically, in some female specimens
<emphasis id="39FCEAEDFF02FF4BFD076303FCD0FEB9" box="[740,793,259,282]" italics="true" pageId="136" pageNumber="137">
<taxonomicName id="CC884D7CFF02FF4BFD076303FCC2FEB9" box="[740,779,259,282]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
5
</emphasis>
is as long as
<emphasis id="39FCEAEDFF02FF4BFC516303FBCAFEB8" box="[946,1027,259,283]" italics="true" pageId="136" pageNumber="137">
<taxonomicName id="CC884D7CFF02FF4BFC516303FC11FEB8" box="[946,984,259,283]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3–4
</emphasis>
, and in some male specimens either
<emphasis id="39FCEAEDFF02FF4BFF746326FF05FE9D" box="[151,204,294,318]" italics="true" pageId="136" pageNumber="137">
<taxonomicName id="CC884D7CFF02FF4BFF746326FF77FE9D" box="[151,190,294,318]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
4
</emphasis>
or
<emphasis id="39FCEAEDFF02FF4BFF1B6326FEE4FE9E" box="[248,301,294,317]" italics="true" pageId="136" pageNumber="137">
<taxonomicName id="CC884D7CFF02FF4BFF1B6326FED6FE9E" box="[248,287,294,317]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
5
</emphasis>
are spine-like, but these specimens are aberrant only on one side of the head. Head largely pale, except median margin of marginal carina, parts of mandibular framework, preocular nodi, occipital carinae, and gular plate with medium-brown pigmentation. Pedicel and flagellomeres I–II with light brown pigmentation. Thoracic and abdominal segments as in genus description and
<figureCitation id="93B32A7AFF02FF4BFC8E6390FBC2FE0B" box="[877,1035,400,425]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="136" pageNumber="137">Figs 210–211</figureCitation>
. Thorax with only proepimera and metepisterna with brown pigmentation, otherwise translucent. Tergopleurites translucent. Sternal plates, pleurites, and subgenital plates of both sexes with light brown pigmentation.
</paragraph>
<paragraph id="0B3736FFFF02FF4BFF2463FAFE90FD02" blockId="136.[151,1437,151,1063]" pageId="136" pageNumber="137">
<collectingRegion id="C94CF81DFF02FF4BFF2463FAFECBFDB0" box="[199,258,506,531]" country="Maldives" name="Maale" pageId="136" pageNumber="137">
<emphasis id="39FCEAEDFF02FF4BFF2463FAFECBFDB0" bold="true" box="[199,258,506,531]" italics="true" pageId="136" pageNumber="137">Male</emphasis>
</collectingRegion>
. Distal part of subgenital plate translucent. Abdominal chaetotaxy as in
<tableCitation id="460A0344FF02FF4BFBBA63FAFB79FDB0" box="[1113,1200,506,531]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="136" pageNumber="137">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFF02FF4BFB0C63FAFA9AFDB0" box="[1263,1363,506,531]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="136" pageNumber="137">Fig. 210</figureCitation>
.
<collectingRegion id="C94CF81DFF02FF4BFA8163FAFA55FDB0" box="[1378,1436,506,531]" country="Maldives" name="Maale" pageId="136" pageNumber="137">Male</collectingRegion>
genitalia as in genus description and
<figureCitation id="93B32A7AFF02FF4BFDA2601DFD29FD95" box="[577,736,541,566]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="136" pageNumber="137">Figs 213–215</figureCitation>
. Proximal mesosome somewhat variable between specimens, rounded to flat;
<emphasis id="39FCEAEDFF02FF4BFEA96042FE4DFDFA" box="[330,388,578,601]" italics="true" pageId="136" pageNumber="137">pmes</emphasis>
not visible in all specimens, including
<typeStatus id="D433885DFF02FF4BFCDB6041FC57FDF9" box="[824,926,577,602]" pageId="136" pageNumber="137" type="holotype">holotype</typeStatus>
. Measurements ex
<taxonomicName id="CC884D7CFF02FF4BFB906042FAB4FDF9" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[1139,1405,577,602]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="136" pageNumber="137" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF02FF4BFB906042FAB4FDF9" box="[1139,1405,577,602]" italics="true" pageId="136" pageNumber="137">Laniarius erythrogaster</emphasis>
</taxonomicName>
(n = 6 except n = 5 for TL): TL = 1.75–1.92; HL = 0.47–0.50; HW = 0.53–0.56; PRW = 0.30–0.32; PTW = 0.48–0.51; AW = 0.65–0.72.
</paragraph>
<paragraph id="0B3736FFFF02FF4BFF2460ACFC4EFC3A" blockId="136.[151,1437,151,1063]" pageId="136" pageNumber="137">
<emphasis id="39FCEAEDFF02FF4BFF2460ACFED7FD66" bold="true" box="[199,286,684,709]" italics="true" pageId="136" pageNumber="137">Female</emphasis>
. Posterior and median sections of tergopleurite IX+X with pale brown pigmentation. Abdominal chaetotaxy as in
<tableCitation id="460A0344FF02FF4BFEBF60CFFE7BFD4B" box="[348,434,719,744]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="136" pageNumber="137">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFF02FF4BFE0C60CFFD9BFD4B" box="[495,594,719,744]" captionStart="FIGURES 210 – 211" captionStartId="134.[151,264,1837,1859]" captionTargetBox="[193,1373,229,1811]" captionTargetId="figure@134.[151,1436,193,1815]" captionTargetPageId="134" captionText="FIGURES 210 – 211. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 210, male habitus, dorsal and ventral views. 211, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883272/files/figure.png" pageId="136" pageNumber="137">Fig. 211</figureCitation>
. Subgenital plate irregularly rectangular, not reaching vulval marin (
<figureCitation id="93B32A7AFF02FF4BFA8C60CFFF0CFCAF" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="136" pageNumber="137">Fig. 216</figureCitation>
). Vulval margin gently rounded (
<figureCitation id="93B32A7AFF02FF4BFDDD60F3FD68FCAF" box="[574,673,755,780]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="136" pageNumber="137">Fig. 216</figureCitation>
), with 5–8 short, slender
<emphasis id="39FCEAEDFF02FF4BFC2160F4FC27FCA8" box="[962,1006,756,779]" italics="true" pageId="136" pageNumber="137">vms</emphasis>
on each side, and 12–16 short, thornlike
<emphasis id="39FCEAEDFF02FF4BFF246117FF20FC8D" box="[199,233,791,814]" italics="true" pageId="136" pageNumber="137">vss</emphasis>
on each side. 6–10 long, slender
<emphasis id="39FCEAEDFF02FF4BFDBE6117FD4BFC8D" box="[605,642,791,814]" italics="true" pageId="136" pageNumber="137">vos</emphasis>
; distal 1–2
<emphasis id="39FCEAEDFF02FF4BFCE66117FCE3FC8D" box="[773,810,791,814]" italics="true" pageId="136" pageNumber="137">vos</emphasis>
median to
<emphasis id="39FCEAEDFF02FF4BFC456117FC01FC8D" box="[934,968,791,814]" italics="true" pageId="136" pageNumber="137">vss</emphasis>
. Some examined females (3 of 7) with 1– 2 thorn-like setae lateral to
<emphasis id="39FCEAEDFF02FF4BFE01613BFDCEFCF1" box="[482,519,827,850]" italics="true" pageId="136" pageNumber="137">vos</emphasis>
on one side as in
<figureCitation id="93B32A7AFF02FF4BFD0E6139FC9AFCF1" box="[749,851,825,850]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="136" pageNumber="137">Fig. 216</figureCitation>
; these thorn-like setae are missing entirely in the remaining
<specimenCount id="1D8EFD76FF02FF4BFEF3615EFE48FCD5" box="[272,385,861,886]" pageId="136" pageNumber="137" type="female">4 females</specimenCount>
. Measurements ex
<taxonomicName id="CC884D7CFF02FF4BFDB8615EFCAEFCD5" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[603,871,861,886]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="136" pageNumber="137" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF02FF4BFDB8615EFCAEFCD5" box="[603,871,861,886]" italics="true" pageId="136" pageNumber="137">Laniarius erythrogaster</emphasis>
</taxonomicName>
(n = 7): TL = 1.92–2.20; HL = 0.52–0.56; HW = 0.58–0.62; PRW = 0.34–0.38; PTW = 0.52–0.57; AW = 0.69–0.80.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF02FF4BFF2461A4FDB0FC43" pageId="136" pageNumber="137" type="etymology">
<paragraph id="0B3736FFFF02FF4BFF2461A4FDB0FC43" blockId="136.[151,1437,151,1063]" pageId="136" pageNumber="137">
<emphasis id="39FCEAEDFF02FF4BFF2461A4FE87FC1E" bold="true" box="[199,334,932,957]" pageId="136" pageNumber="137">Etymology.</emphasis>
The species epithet is formed by Latin “
<emphasis id="39FCEAEDFF02FF4BFCF561A5FCF3FC1F" box="[790,826,933,956]" italics="true" pageId="136" pageNumber="137">sex</emphasis>
” for “six” and “
<emphasis id="39FCEAEDFF02FF4BFC1161A4FBE6FC1E" box="[1010,1071,932,957]" italics="true" pageId="136" pageNumber="137">lobus</emphasis>
” for “lobes”, referring to the six lobes of the male genitalia (
<figureCitation id="93B32A7AFF02FF4BFE2E61C7FDA4FC43" box="[461,621,967,992]" captionStart="FIGURES 212 – 216" captionStartId="135.[151,264,1883,1905]" captionTargetBox="[219,1376,221,1841]" captionTargetId="figure@135.[191,1396,193,1862]" captionTargetPageId="135" captionText="FIGURES 212 – 216. Titanomessor sexloba n. gen. &amp; n. sp. ex Laniarius erythrogaster: 212, male head, dorsal and ventral views. 213, male genitalia, dorsal view. 214, mesosome, ventral view. 215, parameres, dorsal view. 216, female subgenital plate and vulval margin, ventral view. Scale for parameres same as for male genitalia." httpUri="https://zenodo.org/record/883274/files/figure.png" pageId="136" pageNumber="137">Figs 213–215</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="43926574FF02FF4BFF2461EBFC09FB84" pageId="136" pageNumber="137" type="materials_examined">
<paragraph id="0B3736FFFF02FF4BFF2461EBFC09FB84" blockId="136.[151,1437,151,1063]" pageId="136" pageNumber="137">
<materialsCitation id="BBE03CA2FF02FF4BFF2461EBFEC9FB84" ID-GBIF-Occurrence="1587279375" collectingDate="1936-04" collectionCode="NHML" collectorName="R. Meinertzhagen" country="Uganda" location="Uganda" pageId="136" pageNumber="137" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="39FCEAEDFF02FF4BFF2461EBFEB2FBA7" bold="true" box="[199,379,1003,1028]" pageId="136" pageNumber="137">
<typeStatus id="D433885DFF02FF4BFF2461EBFECBFBA7" box="[199,258,1003,1028]" pageId="136" pageNumber="137">Type</typeStatus>
material.
</emphasis>
Ex
<emphasis id="39FCEAEDFF02FF4BFE5661EBFC9BFBA7" bold="true" box="[437,850,1003,1028]" pageId="136" pageNumber="137">
<taxonomicName id="CC884D7CFF02FF4BFE5661EBFD07FBA7" authorityName="Bush" authorityYear="2017" baseAuthorityName="Cretzschmar" baseAuthorityYear="1829" box="[437,718,1003,1028]" class="Aves" family="Malaconotidae" genus="Laniarius" kingdom="Animalia" order="Passeriformes" pageId="136" pageNumber="137" phylum="Chordata" rank="species" species="erythrogaster">
<emphasis id="39FCEAEDFF02FF4BFE5661EBFD07FBA7" bold="true" box="[437,718,1003,1028]" italics="true" pageId="136" pageNumber="137">Laniarius erythrogaster</emphasis>
</taxonomicName>
:
<typeStatus id="D433885DFF02FF4BFD0661EBFC9BFBA7" box="[741,850,1003,1028]" pageId="136" pageNumber="137" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="1D8EFD76FF02FF4BFC8361ECFCBEFBA0" box="[864,887,1004,1027]" pageId="136" pageNumber="137" type="male">♂</specimenCount>
,
<collectingCountry id="739F766FFF02FF4BFC6A61EBFC2AFBA7" box="[905,995,1003,1028]" name="Uganda" pageId="136" pageNumber="137">Uganda</collectingCountry>
,
<collectingDate id="6F72E9D7FF02FF4BFC1661EBFBA6FBA7" box="[1013,1135,1003,1028]" pageId="136" pageNumber="137" value="1936-04">
<date id="7F36103FFF02FF4BFC1661EBFBA6FBA7" box="[1013,1135,1003,1028]" pageId="136" pageNumber="137" value="1936-04">Apr. 1936</date>
</collectingDate>
,
<collectorName id="A67D5329FF02FF4BFB6361ECFA9BFBA7" box="[1152,1362,1003,1028]" pageId="136" pageNumber="137">R. Meinertzhagen</collectorName>
, 7736 (
<collectionCode id="6D99AE3AFF02FF4BFF7C660FFF31FB84" box="[159,248,1039,1063]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="136" pageNumber="137">NHML</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="BBE03CA2FF02FF4BFEE9660EFC75FB84" ID-GBIF-Occurrence="1587279387" box="[266,956,1038,1063]" collectingDate="1936-04" collectionCode="NHML" collectorName="R. Meinertzhagen" country="Uganda" location="Uganda" pageId="136" pageNumber="137" specimenCount="11" specimenCount-female="6" specimenCount-male="5" typeStatus="paratype">
<emphasis id="39FCEAEDFF02FF4BFEE9660EFE42FB84" bold="true" box="[266,395,1038,1063]" pageId="136" pageNumber="137">
<typeStatus id="D433885DFF02FF4BFEE9660EFE4FFB84" box="[266,390,1038,1063]" pageId="136" pageNumber="137" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="1D8EFD76FF02FF4BFE71660EFE7FFB84" box="[402,438,1038,1063]" pageId="136" pageNumber="137" type="male">5♂</specimenCount>
,
<specimenCount id="1D8EFD76FF02FF4BFE20660EFE2EFB84" box="[451,487,1038,1063]" pageId="136" pageNumber="137" type="female">6♀</specimenCount>
, same data as holotype, 7735–6 (
<collectionCode id="6D99AE3AFF02FF4BFCB8660FFC7DFB84" box="[859,948,1039,1063]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="136" pageNumber="137">NHML</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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