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<document ID-DOI="10.11646/phytotaxa.585.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="7681608" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1677506995801" checkinUser="plazi" docAuthor="Tagane, Shuichiro, Souladeth, Phetlasy, Tanaka, Nobuyuki, Phengmala, Kachonesak &amp; Yahara, Tetsukazu" docDate="2023" docId="731884124926FF98FF1D9812FB8FFEAD" docLanguage="en" docName="phytotaxa.585.2.5.pdf" docOrigin="Phytotaxa 585 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.4:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="4" docTitle="Lasianthus bolavenensis Tagane &amp; Soulad. 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="148" masterDocId="8F21FC6A4927FF9BFF95992EFFA7FFE5" masterDocTitle="Two new species and a new record of Lasianthus (Rubiaceae) in Laos" masterLastPageNumber="153" masterPageNumber="145" pageNumber="146" updateTime="1677507388309" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Two new species and a new record of Lasianthus (Rubiaceae) in Laos</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tagane, Shuichiro</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1974-7329</mods:nameIdentifier>
<mods:affiliation>The Kagoshima University Museum, Kagoshima University, Kagoshima, Japan &amp; stagane 29 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1974 - 7329</mods:affiliation>
<mods:nameIdentifier type="email">stagane29@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Souladeth, Phetlasy</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8563-1773</mods:nameIdentifier>
<mods:affiliation>Faculty of Forest Science, National University of Laos, Vientiane, Laos &amp; p. souladeth @ nuol. edu. la; https: // orcid. org / 0000 - 0001 - 8563 - 1773</mods:affiliation>
<mods:nameIdentifier type="email">p.souladeth@nuol.edu.la</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tanaka, Nobuyuki</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0229-8061</mods:nameIdentifier>
<mods:affiliation>Department of Botany, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan &amp; nobuyuki _ tanaka @ kahaku. go. jp; https: // orcid. org / 0000 - 0002 - 0229 - 8061</mods:affiliation>
<mods:nameIdentifier type="email">nobuyuki_tanaka@kahaku.go.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Phengmala, Kachonesak</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1508-0704</mods:nameIdentifier>
<mods:affiliation>Faculty of Forest Science, National University of Laos, Vientiane, Laos &amp; Kachonesak @ gmail. com; https: // orcid. org / 0000 - 0003 - 1508 - 0704</mods:affiliation>
<mods:nameIdentifier type="email">achonesak@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yahara, Tetsukazu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5105-7152</mods:nameIdentifier>
<mods:affiliation>Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan &amp; tet. yahara @ gmail. com; https: // orcid. org / 0000 - 0001 - 5105 - 7152</mods:affiliation>
<mods:nameIdentifier type="email">tet.yahara@gmail.com</mods:nameIdentifier>
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<mods:title>Phytotaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-27</mods:number>
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<mods:number>585</mods:number>
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<mods:number>2</mods:number>
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<mods:start>145</mods:start>
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<mods:identifier type="ISSN">1179-3163</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:731884124926FF98FF1D9812FB8FFEAD" httpUri="http://treatment.plazi.org/id/731884124926FF98FF1D9812FB8FFEAD" lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146">
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<paragraph blockId="1.[136,893,316,343]" box="[136,893,316,343]" pageId="1" pageNumber="146">
<heading box="[136,893,316,343]" fontSize="11" level="2" pageId="1" pageNumber="146" reason="3">
<taxonomicName authority="Tagane &amp; Soulad." authorityName="Tagane &amp; Soulad." authorityYear="2023" box="[136,631,316,343]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bolavenensis" status="sp. nov.">
<emphasis bold="true" box="[136,420,316,343]" italics="true" pageId="1" pageNumber="146">Lasianthus bolavenensis</emphasis>
Tagane &amp; Soulad.
</taxonomicName>
, 
<emphasis bold="true" box="[641,731,316,342]" pageId="1" pageNumber="146">
<taxonomicNameLabel box="[641,731,316,342]" pageId="1" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation box="[738,843,316,342]" captionStart="FIGURE 1" captionStartId="2.[136,229,1744,1766]" captionTargetBox="[258,1316,191,1711]" captionTargetId="figure-25@2.[255,1333,184,1716]" captionTargetPageId="2" captionText="FIGURE 1. Isotype of Lasianthus bolavenensis Tagane &amp; Soulad. (Souladeth et al. L3337 (KAG155702))." figureDoi="http://doi.org/10.5281/zenodo.7681612" httpUri="https://zenodo.org/record/7681612/files/figure.png" pageId="1" pageNumber="146">Figures 1</figureCitation>
&amp; 
<figureCitation box="[879,893,316,342]" captionStart="FIGURE 2" captionStartId="3.[136,229,1507,1529]" captionTargetBox="[283,1303,387,1483]" captionTargetId="figure-102@3.[283,1303,387,1483]" captionTargetPageId="3" captionText="FIGURE 2. Lasianthus bolavenensis Tagane &amp; Soulad.A. Flowering twig; B. Abaxial leaf surface; C. Stipule; D. Flower bud and calyces; E. Calyces after anthesis; F. Flower (side view); G. Flower (front view); H. Fruits. Materials A &amp; D from Souladeth et al. L2501; B, C, E–H from Souladeth et al. L3337." figureDoi="http://doi.org/10.5281/zenodo.7681614" httpUri="https://zenodo.org/record/7681614/files/figure.png" pageId="1" pageNumber="146">2</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="146" type="materials_examined">
<paragraph blockId="1.[136,1452,356,450]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[136,232,356,377]" pageId="1" pageNumber="146">TYPE:—</emphasis>
<materialsCitation collectionCode="FOF, TNS" collectorName="P. Souladeth &amp; S. Tagane &amp; D. Kongxaysavath &amp; S. Rueangruea &amp; S. Suddee &amp; Y. Suyama &amp; E. Suzuki" country="Laos" county="Dong Hua Sao National Park" elevation="1226" latitude="15.0754385" location="Nong Luang Village" longLatPrecision="1" longitude="106.20853" municipality="Paksong District" pageId="1" pageNumber="146" specimenCode="KAG155702" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
<collectingCountry box="[232,298,356,377]" name="Laos" pageId="1" pageNumber="146">LAOS</collectingCountry>
. 
<collectingRegion box="[308,514,356,378]" country="Laos" name="Champasak" pageId="1" pageNumber="146">Champasak Province</collectingRegion>
, 
<collectingCounty box="[525,808,356,378]" pageId="1" pageNumber="146">Dong Hua Sao National Park</collectingCounty>
[Bolaven Plateau], 
<collectingMunicipality box="[1001,1165,356,378]" pageId="1" pageNumber="146">Paksong District</collectingMunicipality>
, near 
<location LSID="urn:lsid:plazi:treatment:731884124926FF98FF1D9812FB8FFEAD:FE6E63DF4926FF9AFB52984AFA28FE9F" box="[1223,1423,356,378]" country="Laos" county="Dong Hua Sao National Park" latitude="15.0754385" longLatPrecision="1" longitude="106.20853" municipality="Paksong District" name="Nong Luang Village" pageId="1" pageNumber="146" stateProvince="Champasak">Nong Luang Village</location>
, in lower montane forest, near a stream, 
<geoCoordinate box="[547,694,390,413]" degrees="15" direction="north" minutes="04" orientation="latitude" pageId="1" pageNumber="146" precision="1" seconds="31.58" value="15.0754385">15°04′31.58″N</geoCoordinate>
, 
<geoCoordinate box="[703,860,390,413]" degrees="106" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="146" precision="1" seconds="30.72" value="106.20853">106°12′30.72″E</geoCoordinate>
, 
<quantity box="[869,942,392,414]" metricMagnitude="3" metricUnit="m" metricValue="1.226" pageId="1" pageNumber="146" unit="m" value="1226.0">
<elevation box="[869,942,392,414]" metricMagnitude="3" metricUnit="m" metricValue="1.226" pageId="1" pageNumber="146" unit="m" value="1226.0">1226 m</elevation>
</quantity>
elev., 
<emphasis italics="true" pageId="1" pageNumber="146">
<collectorName box="[1005,1127,392,413]" pageId="1" pageNumber="146">P. Souladeth</collectorName>
, 
<collectorName box="[1137,1233,392,413]" pageId="1" pageNumber="146">S. Tagane</collectorName>
, 
<collectorName box="[1243,1423,392,413]" pageId="1" pageNumber="146">D. Kongxaysavath</collectorName>
, 
<collectorName pageId="1" pageNumber="146">S. Rueangruea</collectorName>
, 
<collectorName box="[319,415,428,449]" pageId="1" pageNumber="146">S. Suddee</collectorName>
, 
<collectorName box="[424,525,428,449]" pageId="1" pageNumber="146">Y. Suyama</collectorName>
, 
<collectorName box="[535,625,428,449]" pageId="1" pageNumber="146">E. Suzuki</collectorName>
L3337
</emphasis>
[fl. &amp; fr.] (
<typeStatus box="[798,884,428,450]" pageId="1" pageNumber="146">holotype</typeStatus>
<collectionCode box="[890,941,428,449]" pageId="1" pageNumber="146">FOF</collectionCode>
!, isotype 
<specimenCode box="[1029,1160,428,450]" collectionCode="KAG" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="1" pageNumber="146" type="Herbarium">KAG155702</specimenCode>
!, 
<collectionCode box="[1173,1222,428,449]" country="Japan" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="1" pageNumber="146" type="Herbarium">TNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[136,1452,496,774]" pageId="1" pageNumber="146">
<taxonomicName box="[136,413,496,522]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[136,413,496,522]" italics="true" pageId="1" pageNumber="146">Lasianthus bolavenensis</emphasis>
</taxonomicName>
is characterized by glabrous twigs, glabrous leaves except with sparse hairs on midrib and secondary veins abaxially when young, 4 or 5 pairs of secondary veins, and semi-orbicular bracts, by which combination it can be distinguished from the other species of 
<taxonomicName box="[686,809,568,594]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="genus">
<emphasis box="[686,809,568,594]" italics="true" pageId="1" pageNumber="146">Lasianthus</emphasis>
</taxonomicName>
in 
<collectingCountry box="[847,901,569,595]" name="Laos" pageId="1" pageNumber="146">Laos</collectingCountry>
and its surrounding countries. In Indochina, it is similar to 
<taxonomicName authority="Dang &amp; Naiki" authorityName="Dang &amp; Naiki" box="[249,576,604,631]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bidoupensis">
<emphasis box="[249,412,604,630]" italics="true" pageId="1" pageNumber="146">L. bidoupensis</emphasis>
Dang &amp; Naiki
</taxonomicName>
in Dang 
<emphasis box="[679,734,604,630]" italics="true" pageId="1" pageNumber="146">et al.</emphasis>
(2019: 191), an endemic species to southern 
<collectingCountry box="[1238,1336,604,630]" name="Vietnam" pageId="1" pageNumber="146">Vietnam</collectingCountry>
, in having almost glabrous twigs and leaves, elliptic-oblong lamina, 5-merous flowers, pinkish-white calyx, and bluish fruits, but different in having smaller leaf size (4–11.3 × 
<quantity box="[653,778,676,703]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.8" metricValueMin="1.6" pageId="1" pageNumber="146" unit="cm" value="2.7" valueMax="3.8" valueMin="1.6">1.6–3.8 cm</quantity>
in 
<taxonomicName box="[814,988,676,702]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[814,988,676,702]" italics="true" pageId="1" pageNumber="146">L. bolavenensis</emphasis>
</taxonomicName>
vs. 16–24 × 
<quantity box="[1133,1236,676,703]" metricMagnitude="-2" metricUnit="m" metricValue="7.25" metricValueMax="8.5" metricValueMin="6.0" pageId="1" pageNumber="146" unit="cm" value="7.25" valueMax="8.5" valueMin="6.0">6–8.5 cm</quantity>
in 
<taxonomicName box="[1272,1435,676,702]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bidoupensis">
<emphasis box="[1272,1435,676,702]" italics="true" pageId="1" pageNumber="146">L. bidoupensis</emphasis>
</taxonomicName>
), fewer secondary veins (4 or 5 vs. 6 or 7), larger anthers (
<quantity box="[795,885,712,739]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="146" unit="mm" value="2.8">2.8 mm</quantity>
long vs. 
<quantity box="[993,1109,712,739]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="146" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
long), and glabrous style (vs. pubescent) (Table 1).
</paragraph>
<paragraph blockId="1.[136,1452,822,882]" box="[136,1452,822,846]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[136,226,822,846]" pageId="1" pageNumber="146">Table 1.</emphasis>
Comparison of 
<taxonomicName box="[399,661,822,846]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[399,661,822,846]" italics="true" pageId="1" pageNumber="146">Lasianthus bolavenensis</emphasis>
</taxonomicName>
with two morphologically similar species. Data on 
<taxonomicName box="[1215,1370,822,846]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bidoupensis">
<emphasis box="[1215,1370,822,846]" italics="true" pageId="1" pageNumber="146">L. bidoupensis</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[1430,1452,823,846]" pageId="1" pageNumber="146">
<emphasis box="[1430,1452,823,846]" italics="true" pageId="1" pageNumber="146">L.</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="1.[136,1452,822,882]" box="[136,1005,858,882]" pageId="1" pageNumber="146">
<emphasis box="[136,231,858,882]" italics="true" pageId="1" pageNumber="146">inodorus</emphasis>
subsp. 
<emphasis box="[312,407,858,882]" italics="true" pageId="1" pageNumber="146">inodorus</emphasis>
were retrieved from Dang et al. (2019) and 
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[877,1002,858,882]" pageId="1" pageNumber="146" pagination="1 - 102" refId="ref5152" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">Zhu (2012)</bibRefCitation>
.
</paragraph>
</subSubSection>
<paragraph pageId="1" pageNumber="146">
<table box="[144,1445,894,1230]" gridcols="4" gridrows="11" pageId="1" pageNumber="146">
<tr box="[144,1445,894,917]" gridrow="0" pageId="1" pageNumber="146">
<th box="[144,347,894,917]" gridcol="0" gridrow="0" pageId="1" pageNumber="146">
<emphasis bold="true" box="[144,259,895,917]" pageId="1" pageNumber="146">Characters</emphasis>
</th>
<th box="[427,740,894,917]" gridcol="1" gridrow="0" pageId="1" pageNumber="146">
<taxonomicName box="[427,670,894,916]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis bold="true" box="[427,670,894,916]" italics="true" pageId="1" pageNumber="146">Lasianthus bolavenensis</emphasis>
</taxonomicName>
</th>
<th box="[771,1073,894,917]" gridcol="2" gridrow="0" pageId="1" pageNumber="146">
<taxonomicName box="[771,915,894,916]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bidoupensis">
<emphasis bold="true" box="[771,915,894,916]" italics="true" pageId="1" pageNumber="146">L. bidoupensis</emphasis>
</taxonomicName>
</th>
<th box="[1116,1445,894,917]" gridcol="3" gridrow="0" pageId="1" pageNumber="146">
<emphasis bold="true" box="[1116,1395,894,916]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[1116,1230,894,916]" italics="true" pageId="1" pageNumber="146">L. inodorus</emphasis>
subsp. 
<emphasis bold="true" box="[1307,1395,894,916]" italics="true" pageId="1" pageNumber="146">inodorus</emphasis>
</emphasis>
</th>
</tr>
<tr box="[144,1445,926,949]" gridrow="1" pageId="1" pageNumber="146">
<th box="[144,347,926,949]" gridcol="0" gridrow="1" pageId="1" pageNumber="146">Plant height</th>
<td box="[427,740,926,949]" gridcol="1" gridrow="1" pageId="1" pageNumber="146">1.5 m tall</td>
<td box="[771,1073,926,949]" gridcol="2" gridrow="1" pageId="1" pageNumber="146">Up to 3 m tall</td>
<td box="[1116,1445,926,949]" gridcol="3" gridrow="1" pageId="1" pageNumber="146">2–3 m tall</td>
</tr>
<tr box="[144,1445,958,980]" gridrow="2" pageId="1" pageNumber="146">
<th box="[144,347,958,980]" gridcol="0" gridrow="2" pageId="1" pageNumber="146">Lamina size</th>
<td box="[427,740,958,980]" gridcol="1" gridrow="2" pageId="1" pageNumber="146">(4–)5.5–11.3 × (1.6–)2.1–3.8 cm</td>
<td box="[771,1073,958,980]" gridcol="2" gridrow="2" pageId="1" pageNumber="146">16–24 × 6–8.5 cm</td>
<td box="[1116,1445,958,980]" gridcol="3" gridrow="2" pageId="1" pageNumber="146">10–20 × 2.5–6 cm</td>
</tr>
<tr box="[144,1445,990,1012]" gridrow="3" pageId="1" pageNumber="146">
<th box="[144,740,990,1012]" colspan="2" colspanRight="1" gridcol="0" gridrow="3" pageId="1" pageNumber="146">Number of secondary veins 4 or 5 pairs</th>
<td box="[771,1073,990,1012]" gridcol="2" gridrow="3" pageId="1" pageNumber="146">6 or 7 pairs</td>
<td box="[1116,1445,990,1012]" gridcol="3" gridrow="3" pageId="1" pageNumber="146">6 or 7 pairs</td>
</tr>
<tr box="[144,1445,1022,1044]" gridrow="4" pageId="1" pageNumber="146">
<th box="[144,347,1022,1044]" gridcol="0" gridrow="4" pageId="1" pageNumber="146">Stipules</th>
<td box="[427,740,1022,1044]" gridcol="1" gridrow="4" pageId="1" pageNumber="146">triangular, 2–2.5 mm long</td>
<td box="[771,1073,1022,1044]" gridcol="2" gridrow="4" pageId="1" pageNumber="146">triangular to broadly triangular,</td>
<td box="[1116,1445,1022,1044]" gridcol="3" gridrow="4" pageId="1" pageNumber="146">triangular to ovate-lanceolate, 3–5</td>
</tr>
<tr box="[144,1445,1051,1073]" gridrow="5" pageId="1" pageNumber="146" rowspan-0="1" rowspan-1="1">
<td box="[771,1073,1051,1073]" gridcol="2" gridrow="5" pageId="1" pageNumber="146">5.5–6.5 mm long</td>
<td box="[1116,1445,1051,1073]" gridcol="3" gridrow="5" pageId="1" pageNumber="146">mm long</td>
</tr>
<tr box="[144,1445,1083,1105]" gridrow="6" pageId="1" pageNumber="146">
<th box="[144,347,1083,1105]" gridcol="0" gridrow="6" pageId="1" pageNumber="146">Bract size</th>
<td box="[427,740,1083,1105]" gridcol="1" gridrow="6" pageId="1" pageNumber="146">1–2 mm long</td>
<td box="[771,1073,1083,1105]" gridcol="2" gridrow="6" pageId="1" pageNumber="146">5–8 mm long</td>
<td box="[1116,1445,1083,1105]" gridcol="3" gridrow="6" pageId="1" pageNumber="146">3–6 mm long</td>
</tr>
<tr box="[144,1445,1115,1137]" gridrow="7" pageId="1" pageNumber="146">
<th box="[144,347,1115,1137]" gridcol="0" gridrow="7" pageId="1" pageNumber="146">Anther length</th>
<td box="[427,740,1115,1137]" gridcol="1" gridrow="7" pageId="1" pageNumber="146">2.8 mm long</td>
<td box="[771,1073,1115,1137]" gridcol="2" gridrow="7" pageId="1" pageNumber="146">1–1.2 mm long</td>
<td box="[1116,1445,1115,1137]" gridcol="3" gridrow="7" pageId="1" pageNumber="146">-</td>
</tr>
<tr box="[144,1445,1146,1168]" gridrow="8" pageId="1" pageNumber="146">
<th box="[144,347,1146,1168]" gridcol="0" gridrow="8" pageId="1" pageNumber="146">Hairiness of style</th>
<td box="[427,740,1146,1168]" gridcol="1" gridrow="8" pageId="1" pageNumber="146">glabrous</td>
<td box="[771,1073,1146,1168]" gridcol="2" gridrow="8" pageId="1" pageNumber="146">pubescent</td>
<td box="[1116,1445,1146,1168]" gridcol="3" gridrow="8" pageId="1" pageNumber="146">-</td>
</tr>
<tr box="[144,1445,1178,1200]" gridrow="9" pageId="1" pageNumber="146">
<th box="[144,347,1178,1200]" gridcol="0" gridrow="9" pageId="1" pageNumber="146">Fruit shape and color</th>
<td box="[427,740,1178,1200]" gridcol="1" gridrow="9" pageId="1" pageNumber="146">oblong, bluish</td>
<td box="[771,1073,1178,1200]" gridcol="2" gridrow="9" pageId="1" pageNumber="146">ovoid or ellipsoid, deep red</td>
<td box="[1116,1445,1178,1200]" gridcol="3" gridrow="9" pageId="1" pageNumber="146">ovoid, dark orange-red turning</td>
</tr>
<tr box="[144,1445,1208,1230]" gridrow="10" pageId="1" pageNumber="146" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td box="[1116,1445,1208,1230]" gridcol="3" gridrow="10" pageId="1" pageNumber="146">black when mature</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="1" pageNumber="146" type="description">
<paragraph blockId="1.[136,1452,1273,2092]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[189,327,1273,1299]" pageId="1" pageNumber="146">Description</emphasis>
:—Shrub, 
<quantity box="[447,513,1273,1300]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="146" unit="m" value="1.5">1.5 m</quantity>
tall. Twigs glabrous, blackish when dry, terete; internode 1.7–4(–7.5) cm. Leaves opposite, petiolate; petiole 
<quantity box="[456,587,1310,1336]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="1" pageNumber="146" unit="cm" value="1.05" valueMax="1.3" valueMin="0.8">0.8–1.3 cm</quantity>
long, glabrous; blade ovate-elliptic, elliptic-oblong, elliptic, (4–)5.5–11.3 × (1.6–) 
<quantity box="[228,361,1345,1372]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.8" metricValueMin="2.1" pageId="1" pageNumber="146" unit="cm" value="2.95" valueMax="3.8" valueMin="2.1">2.1–3.8 cm</quantity>
, coriaceous, glabrous except on midrib and secondary veins abaxially which is sparsely hairy, adaxially dark grayish brown, abaxially dark brownish yellow to gray-green, apex acuminate to aristate, tip pointed ca. 
<quantity box="[136,222,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="146" unit="mm" value="1.2">1.2 mm</quantity>
long, base cuneate, sometimes oblique, margin entire, midrib slightly concave adaxially, prominent abaxially, secondary veins 4 or 5 pairs, prominent abaxially, tertiary veins parallel. Stipule triangular, 
<quantity box="[1146,1259,1453,1480]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="146" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
long, apex acute, densely pubescent near margin. Inflorescence axillary, cymose, 1–4 flowered, sessile; bract semi-orbicular, 
<quantity box="[1338,1452,1490,1516]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="146" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
long, apex truncate, margin densely pubescent. Flowers subsessile; calyx pinkish white, ca. 
<quantity box="[1195,1263,1525,1552]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="146" unit="mm" value="6.0">6 mm</quantity>
long, tube ca. 
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long, glabrous, lobes triangular, ca. 
<quantity box="[592,657,1562,1588]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="146" unit="mm" value="2.0">2 mm</quantity>
long, apex acute, pubescent near apex; corolla pinkish white to white, cylindrical-campanulate, ca. 
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long, tube ca. 
<quantity box="[697,786,1597,1624]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="146" unit="mm" value="8.5">8.5 mm</quantity>
long, outside villous in upper 3/4, inside villous in upper 1/2, lobes 5, triangular, ca. 
<quantity box="[445,532,1633,1660]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="146" unit="mm" value="1.5">1.5 mm</quantity>
long, apex acute, villous. Stamens 5, inserted in corolla tube, anthers ca. 
<quantity box="[1365,1452,1634,1660]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="146" unit="mm" value="2.8">2.8 mm</quantity>
long, filament basally connate to corolla, free part ca. 
<quantity box="[735,821,1670,1696]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="146" unit="mm" value="0.2">0.2 mm</quantity>
long; ovary 5-locular, style 
<quantity box="[1133,1198,1670,1696]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="146" unit="mm" value="4.0">4 mm</quantity>
long, glabrous, stigma 4-lobed, ca. 
<quantity box="[276,342,1706,1732]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="146" unit="mm" value="2.0">2 mm</quantity>
long. Fruits subglobose or oblong, 
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long, ca. 
<quantity box="[983,1049,1705,1732]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="146" unit="mm" value="5.0">5 mm</quantity>
in diam., glabrous, blue when ripe; pyrenes 5. Seeds 5, ca. 
<quantity box="[397,462,1741,1768]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="146" unit="mm" value="6.0">6 mm</quantity>
long, blackish brown when dry.
</paragraph>
<paragraph blockId="1.[136,1452,1273,2092]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[189,596,1777,1803]" pageId="1" pageNumber="146">Additional specimen examined:—</emphasis>
<materialsCitation collectingDate="2019-02-19" collectionCode="BKF, FOF" collectorName="P. Souladeth" country="Laos" county="Dong Hua Sao National Protected Area" elevation="1147" latitude="15.067389" location="Ban Nong Luang" longLatPrecision="1" longitude="106.208305" municipality="Seua Waterfall" pageId="1" pageNumber="146" specimenCode="KAG128750" specimenCount="1" stateProvince="Champasak">
<collectingCountry box="[596,673,1778,1804]" name="Laos" pageId="1" pageNumber="146">LAOS</collectingCountry>
. 
<collectingRegion box="[684,924,1777,1803]" country="Laos" name="Champasak" pageId="1" pageNumber="146">Champasak Province</collectingRegion>
, 
<collectingCounty box="[937,1385,1777,1804]" pageId="1" pageNumber="146">Dong Hua Sao National Protected Area</collectingCounty>
, 
<collectingMunicipality pageId="1" pageNumber="146">Seua Waterfall</collectingMunicipality>
near 
<location LSID="urn:lsid:plazi:treatment:731884124926FF98FF1D9812FB8FFEAD:FE6E63DF4926FF9AFEA79E38FE5EF8D5" box="[306,505,1814,1840]" country="Laos" county="Dong Hua Sao National Protected Area" latitude="15.067389" longLatPrecision="1" longitude="106.208305" municipality="Seua Waterfall" name="Ban Nong Luang" pageId="1" pageNumber="146" stateProvince="Champasak">Ban Nong Luang</location>
, 
<geoCoordinate box="[519,676,1815,1842]" degrees="15" direction="north" minutes="04" orientation="latitude" pageId="1" pageNumber="146" precision="1" seconds="02.6" value="15.067389">15°04′02.6″N</geoCoordinate>
, 
<geoCoordinate box="[689,856,1815,1842]" degrees="106" direction="east" minutes="12" orientation="longitude" pageId="1" pageNumber="146" precision="1" seconds="29.9" value="106.208305">106°12′29.9″E</geoCoordinate>
, 
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<elevation box="[870,956,1814,1840]" metricMagnitude="3" metricUnit="m" metricValue="1.147" pageId="1" pageNumber="146" unit="m" value="1147.0">1147 m</elevation>
</quantity>
elev., 
<date box="[1024,1230,1813,1840]" pageId="1" pageNumber="146" value="2019-02-19">
<collectingDate box="[1024,1230,1813,1840]" pageId="1" pageNumber="146" value="2019-02-19">19 February 2019</collectingDate>
</date>
, 
<emphasis italics="true" pageId="1" pageNumber="146">
<collectorName box="[1243,1384,1813,1839]" pageId="1" pageNumber="146">P. Souladeth</collectorName>
et al. L2501
</emphasis>
[fl.] (
<collectionCode box="[272,329,1850,1876]" country="Thailand" name="National Park, Wildlife and Plant Conservation Department" pageId="1" pageNumber="146" type="Herbarium">BKF</collectionCode>
, 
<collectionCode box="[340,400,1850,1876]" pageId="1" pageNumber="146">FOF</collectionCode>
!, 
<specimenCode box="[415,570,1849,1876]" collectionCode="KAG" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="1" pageNumber="146" type="Herbarium">KAG128750</specimenCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="146" type="distribution">
<paragraph blockId="1.[136,1452,1273,2092]" box="[189,1226,1885,1912]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[189,373,1885,1911]" pageId="1" pageNumber="146">Distribution:—</emphasis>
<collectingCountry box="[373,446,1886,1912]" name="Laos" pageId="1" pageNumber="146">LAOS</collectingCountry>
(so far known only from the Bolaven Plateau, 
<collectingRegion box="[972,1214,1885,1911]" country="Laos" name="Champasak" pageId="1" pageNumber="146">Champasak Province</collectingRegion>
).
</paragraph>
<paragraph blockId="1.[136,1452,1273,2092]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[189,467,1921,1948]" pageId="1" pageNumber="146">Habitat and Ecology:—</emphasis>
The species was found in wet evergreen forest at elev. 
<quantity box="[1053,1207,1922,1948]" metricMagnitude="3" metricUnit="m" metricValue="1.21" metricValueMax="1.28" metricValueMin="1.14" pageId="1" pageNumber="146" unit="m" value="1210.0" valueMax="1280.0" valueMin="1140.0">1140–1280 m</quantity>
. Flowering specimens were collected in December and February, and fruiting in December.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="148" pageId="1" pageNumber="146" type="etymology">
<paragraph blockId="1.[136,1452,1273,2092]" box="[189,1224,1993,2020]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[189,356,1994,2020]" pageId="1" pageNumber="146">Etymology:—</emphasis>
This specific epithet 
<taxonomicName box="[590,735,1993,2019]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[590,735,1993,2019]" italics="true" pageId="1" pageNumber="146">bolavenensis</emphasis>
</taxonomicName>
refers to the 
<typeStatus box="[882,930,1994,2020]" pageId="1" pageNumber="146">type</typeStatus>
locality, Bolaven Plateau.
</paragraph>
<paragraph blockId="1.[136,1452,1273,2092]" pageId="1" pageNumber="146">
<emphasis bold="true" box="[189,435,2030,2056]" pageId="1" pageNumber="146">Vernacular name:—</emphasis>
[(Xangsan Bolaven, “Xangsan” means 
<taxonomicName box="[1124,1247,2029,2055]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="146" phylum="Tracheophyta" rank="genus">
<emphasis box="[1124,1247,2029,2055]" italics="true" pageId="1" pageNumber="146">Lasianthus</emphasis>
</taxonomicName>
in general in Lao and “Bolaven” refers to the Bolaven Plateau where the species occurs), proposed here].
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7681612" ID-Zenodo-Dep="7681612" box="[136,1168,1744,1766]" httpUri="https://zenodo.org/record/7681612/files/figure.png" pageId="2" pageNumber="147" startId="2.[136,229,1744,1766]" targetBox="[258,1316,191,1711]" targetPageId="2">
<paragraph blockId="2.[136,1168,1744,1766]" box="[136,1168,1744,1766]" pageId="2" pageNumber="147">
<emphasis bold="true" box="[136,247,1744,1766]" pageId="2" pageNumber="147">FIGURE 1</emphasis>
. Isotype of 
<taxonomicName authority="Tagane &amp; Soulad." authorityName="Tagane &amp; Soulad." box="[362,776,1744,1766]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="147" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[362,597,1744,1765]" italics="true" pageId="2" pageNumber="147">Lasianthus bolavenensis</emphasis>
Tagane &amp; Soulad.
</taxonomicName>
(
<emphasis box="[789,1007,1744,1766]" italics="true" pageId="2" pageNumber="147">Souladeth et al. L3337</emphasis>
(KAG155702)).
</paragraph>
</caption>
<paragraph blockId="2.[136,1452,1812,2091]" pageId="2" pageNumber="147">
<emphasis bold="true" box="[189,681,1812,1839]" pageId="2" pageNumber="147">Preliminary conservation assessment:—</emphasis>
Critically Endangered (CR). From our observations by five field surveys in 2018–2020 (
<bibRefCitation author="Souladeth, P. &amp; Tagane, S. &amp; Newman, M. F. &amp; Prajaksood, A." box="[405,655,1849,1875]" pageId="2" pageNumber="147" pagination="56" refId="ref4852" refString="Souladeth, P., Tagane, S., Newman, M. F. &amp; Prajaksood, A. (2020) Two new species of Eriocaulon (Eriocaulaceae) from Laos. Kew Bulletin 75: 56. https: // doi. org / 10.1007 / s 12225 - 020 - 09909 - 0" type="journal article" year="2020">
Souladeth 
<emphasis box="[529,587,1849,1875]" italics="true" pageId="2" pageNumber="147">et al.</emphasis>
2020
</bibRefCitation>
, Tagane 
<emphasis box="[758,817,1849,1875]" italics="true" pageId="2" pageNumber="147">et al.</emphasis>
2020), 
<taxonomicName box="[908,1185,1849,1875]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="147" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[908,1185,1849,1875]" italics="true" pageId="2" pageNumber="147">Lasianthus bolavenensis</emphasis>
</taxonomicName>
is locally common but narrowly restricted in a small area on Bolaven Plateau: in wet evergreen forest at elevation between 
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, and we estimate the area of occupancy is less than 
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and mature individuals less than 1000. The most area of its habitat is located within the protected area of Dong Hua Sao National Park (formerly Dong Hua Sao National Protected Area), but some of the vicinities are being converted to coffee plantations, which might bring negative impact for the survival of this species. Given this situation, we propose Critically Endangered (CR) according to B2b(ii, iii)(
<bibRefCitation author="IUCN" pageId="2" pageNumber="147" refId="ref4467" refString="IUCN (2012) IUCN Red List Categories and Criteria, version 3.1. Gland and Cambridge. Available from: http: // www. iucnredlist. org (accessed 24 March 2021)" type="url" year="2012">IUCN 2012</bibRefCitation>
) for the first conservation assessment for this species.
</paragraph>
<paragraph blockId="3.[136,1452,158,329]" pageId="3" pageNumber="148">
<emphasis bold="true" box="[189,283,158,184]" pageId="3" pageNumber="148">Note:—</emphasis>
<taxonomicName box="[283,556,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="bolavenensis">
<emphasis box="[283,556,158,184]" italics="true" pageId="3" pageNumber="148">Lasianthus bolavenensis</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName authority="Blume (1826" authorityName="Blume" authorityYear="1826" box="[748,1128,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="inodorus">
<emphasis box="[748,977,158,184]" italics="true" pageId="3" pageNumber="148">Lasianthus inodorus</emphasis>
<bibRefCitation author="Blume, C. L." box="[982,1128,158,184]" pageId="3" pageNumber="148" pagination="994 - 1001" refId="ref4128" refString="Blume, C. L. (1826 - 1827) Bijdragen tot de flora van Nederlandsch Indie ¨, Part 16 - 17. Ter Lands Drukkerij, Batavia, pp. 994 - 1001. https: // doi. org / 10.5962 / bhl. title. 115427" type="book chapter" year="1826">Blume (1826</bibRefCitation>
</taxonomicName>
–1827: 998) 
<taxonomicName box="[1268,1445,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="subSpecies" species="inodorus" subSpecies="inodorus">
subsp. 
<emphasis box="[1345,1445,158,184]" italics="true" pageId="3" pageNumber="148">inodorus</emphasis>
</taxonomicName>
, widely distributed in 
<collectingCountry box="[387,521,194,220]" name="Bangladesh" pageId="3" pageNumber="148">Bangladesh</collectingCountry>
, 
<collectingCountry box="[536,654,194,220]" name="Cambodia" pageId="3" pageNumber="148">Cambodia</collectingCountry>
, 
<collectingCountry box="[669,739,194,220]" name="China" pageId="3" pageNumber="148">China</collectingCountry>
, 
<collectingCountry box="[755,815,194,220]" name="India" pageId="3" pageNumber="148">India</collectingCountry>
, 
<collectingCountry box="[830,939,194,220]" name="Indonesia" pageId="3" pageNumber="148">Indonesia</collectingCountry>
(
<collectingRegion box="[958,1056,194,220]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="148">Sumatra</collectingRegion>
, 
<collectingRegion box="[1071,1125,194,220]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="148">Java</collectingRegion>
), 
<collectingCountry box="[1148,1250,194,220]" name="Thailand" pageId="3" pageNumber="148">Thailand</collectingCountry>
, and 
<collectingCountry box="[1317,1415,194,220]" name="Vietnam" pageId="3" pageNumber="148">Vietnam</collectingCountry>
, in glabrous twigs and leaves (or sparsely puberulous on lower leaf surface) and sessile inflorescence, but different in having 4 or 5 pairs of secondary veins (vs. 6 or 7 pairs), smaller bracts 
<quantity box="[927,1041,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="3" pageNumber="148" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
long (vs. 
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long), and blueish oblong fruits (vs. dark orange-red turning black when mature and ovoid) (Table 1).
</paragraph>
</subSubSection>
</treatment>
</document>