treatments-xml/data/73/18/84/731884124924FF9DFF1D9F41FC3AF9D4.xml

<document ID-DOI="10.11646/phytotaxa.585.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="7681608" checkinTime="1677506995801" checkinUser="plazi" docAuthor="Tagane, Shuichiro, Souladeth, Phetlasy, Tanaka, Nobuyuki, Phengmala, Kachonesak &amp; Yahara, Tetsukazu" docDate="2023" docId="731884124924FF9DFF1D9F41FC3AF9D4" docLanguage="en" docName="phytotaxa.585.2.5.pdf" docOrigin="Phytotaxa 585 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Lasianthus laoticus Tagane, Soulad. &amp; Nob. Tanaka 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="151" masterDocId="8F21FC6A4927FF9BFF95992EFFA7FFE5" masterDocTitle="Two new species and a new record of Lasianthus (Rubiaceae) in Laos" masterLastPageNumber="153" masterPageNumber="145" pageNumber="148" updateTime="1677613052640" updateUser="diego" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo>
<mods:title>Two new species and a new record of Lasianthus (Rubiaceae) in Laos</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tagane, Shuichiro</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1974-7329</mods:nameIdentifier>
<mods:affiliation>The Kagoshima University Museum, Kagoshima University, Kagoshima, Japan &amp; stagane 29 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1974 - 7329</mods:affiliation>
<mods:nameIdentifier type="email">stagane29@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Souladeth, Phetlasy</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8563-1773</mods:nameIdentifier>
<mods:affiliation>Faculty of Forest Science, National University of Laos, Vientiane, Laos &amp; p. souladeth @ nuol. edu. la; https: // orcid. org / 0000 - 0001 - 8563 - 1773</mods:affiliation>
<mods:nameIdentifier type="email">p.souladeth@nuol.edu.la</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tanaka, Nobuyuki</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0229-8061</mods:nameIdentifier>
<mods:affiliation>Department of Botany, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan &amp; nobuyuki _ tanaka @ kahaku. go. jp; https: // orcid. org / 0000 - 0002 - 0229 - 8061</mods:affiliation>
<mods:nameIdentifier type="email">nobuyuki_tanaka@kahaku.go.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Phengmala, Kachonesak</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1508-0704</mods:nameIdentifier>
<mods:affiliation>Faculty of Forest Science, National University of Laos, Vientiane, Laos &amp; Kachonesak @ gmail. com; https: // orcid. org / 0000 - 0003 - 1508 - 0704</mods:affiliation>
<mods:nameIdentifier type="email">achonesak@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yahara, Tetsukazu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5105-7152</mods:nameIdentifier>
<mods:affiliation>Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan &amp; tet. yahara @ gmail. com; https: // orcid. org / 0000 - 0001 - 5105 - 7152</mods:affiliation>
<mods:nameIdentifier type="email">tet.yahara@gmail.com</mods:nameIdentifier>
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<mods:title>Phytotaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-27</mods:number>
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<mods:detail type="volume">
<mods:number>585</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>145</mods:start>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.585.2.5</mods:url>
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<mods:identifier type="DOI">10.11646/phytotaxa.585.2.5</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7681608</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:731884124924FF9DFF1D9F41FC3AF9D4" httpUri="http://treatment.plazi.org/id/731884124924FF9DFF1D9F41FC3AF9D4" lastPageId="6" lastPageNumber="151" pageId="3" pageNumber="148">
<subSubSection box="[136,985,1647,1674]" pageId="3" pageNumber="148" type="nomenclature">
<paragraph blockId="3.[136,985,1647,1674]" box="[136,985,1647,1674]" pageId="3" pageNumber="148">
<heading box="[136,985,1647,1674]" fontSize="11" level="2" pageId="3" pageNumber="148" reason="3">
<taxonomicName authority="Tagane, Soulad. &amp; Nob.Tanaka" authorityName="Tagane, Soulad. &amp; Nob. Tanaka" authorityYear="2023" box="[136,723,1647,1674]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="laoticus" status="sp. nov.">
<emphasis bold="true" box="[136,362,1647,1674]" italics="true" pageId="3" pageNumber="148">Lasianthus laoticus</emphasis>
Tagane, Soulad. &amp; Nob.Tanaka
</taxonomicName>
, 
<emphasis bold="true" box="[734,824,1648,1674]" pageId="3" pageNumber="148">
<taxonomicNameLabel box="[734,824,1648,1674]" pageId="3" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation box="[831,936,1647,1674]" captionStart="FIGURE 3" captionStartId="5.[136,229,2042,2064]" captionTargetBox="[199,1385,331,2014]" captionTargetId="figure-75@5.[195,1391,315,2018]" captionTargetPageId="5" captionText="FIGURE 3. Isotype of Lasianthus laoticus Tagane, Soulad. &amp; Nob.Tanaka. (Souladeth et al. L3024 (KAG129297))." figureDoi="http://doi.org/10.5281/zenodo.7681616" httpUri="https://zenodo.org/record/7681616/files/figure.png" pageId="3" pageNumber="148">Figures 3</figureCitation>
&amp; 
<figureCitation box="[971,985,1648,1674]" captionStart="FIGURE 4" captionStartId="6.[136,229,1310,1332]" captionTargetBox="[283,1303,190,1286]" captionTargetId="figure-25@6.[283,1303,190,1286]" captionTargetPageId="6" captionText="FIGURE 4. Lasianthus laoticus Tagane, Soulad. &amp; Nob.Tanaka. A. Flowering twig; B. Portion of abaxial leaf surface; C. Inflorescence; D. Calyx; E. Fruit.All from Souladeth et al. L3024." figureDoi="http://doi.org/10.5281/zenodo.7681618" httpUri="https://zenodo.org/record/7681618/files/figure.png" pageId="3" pageNumber="148">4</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="149" pageId="3" pageNumber="148" type="materials_examined">
<paragraph blockId="3.[136,1452,1686,1781]" pageId="3" pageNumber="148">
<emphasis bold="true" box="[136,232,1687,1708]" pageId="3" pageNumber="148">TYPE:—</emphasis>
<materialsCitation collectingDate="2019-07-06" collectionCode="FOF, TNS" collectorName="P. Souladeth &amp; S. Tagane &amp; N. Tanaka &amp; K. Souttakone &amp; Y. Suyama &amp; K. Phengmala &amp; N. Ishii" country="Laos" elevation="183" latitude="14.7883835" location="Sanamxai District" longLatPrecision="1" longitude="106.423584" pageId="3" pageNumber="148" specimenCode="KAG129297" specimenCount="1" stateProvince="Attapu" typeStatus="holotype">
<collectingCountry box="[232,298,1687,1708]" name="Laos" pageId="3" pageNumber="148">LAOS</collectingCountry>
. 
<collectingRegion box="[308,482,1687,1709]" country="Laos" name="Attapu" pageId="3" pageNumber="148">Attapeu Province</collectingRegion>
: 
<location LSID="urn:lsid:plazi:treatment:731884124924FF9DFF1D9F41FC3AF9D4:FE6E63DF4924FF98FE7B9FB9FD07F948" box="[494,672,1687,1709]" country="Laos" latitude="14.7883835" longLatPrecision="1" longitude="106.423584" name="Sanamxai District" pageId="3" pageNumber="148" stateProvince="Attapu">Sanamxai District</location>
, roadside, at edge of evergreen forest, 
<geoCoordinate box="[1061,1208,1686,1709]" degrees="14" direction="north" minutes="47" orientation="latitude" pageId="3" pageNumber="148" precision="1" seconds="18.18" value="14.7883835">14°47′18.18″N</geoCoordinate>
, 
<geoCoordinate box="[1220,1375,1686,1709]" degrees="106" direction="east" minutes="25" orientation="longitude" pageId="3" pageNumber="148" precision="1" seconds="24.90" value="106.423584">106°25′24.90″E</geoCoordinate>
, 
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<elevation box="[1388,1452,1687,1708]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="3" pageNumber="148" unit="m" value="183.0">183 m</elevation>
</quantity>
elev., 
<date box="[247,364,1723,1745]" pageId="3" pageNumber="148" value="2019-07-06">
<collectingDate box="[247,364,1723,1745]" pageId="3" pageNumber="148" value="2019-07-06">6 July 2019</collectingDate>
</date>
, 
<emphasis box="[375,1287,1723,1745]" italics="true" pageId="3" pageNumber="148">
<collectorName box="[375,498,1723,1745]" pageId="3" pageNumber="148">P. Souladeth</collectorName>
, 
<collectorName box="[510,607,1723,1745]" pageId="3" pageNumber="148">S. Tagane</collectorName>
, 
<collectorName box="[618,719,1723,1745]" pageId="3" pageNumber="148">N. Tanaka</collectorName>
, 
<collectorName box="[731,869,1723,1745]" pageId="3" pageNumber="148">K. Souttakone</collectorName>
, 
<collectorName box="[880,982,1723,1745]" pageId="3" pageNumber="148">Y. Suyama</collectorName>
, 
<collectorName box="[994,1135,1723,1745]" pageId="3" pageNumber="148">K. Phengmala</collectorName>
, 
<collectorName box="[1146,1218,1723,1745]" pageId="3" pageNumber="148">N. Ishii</collectorName>
L3024
</emphasis>
(
<typeStatus box="[1300,1386,1723,1745]" pageId="3" pageNumber="148">holotype</typeStatus>
<collectionCode box="[1393,1444,1723,1744]" pageId="3" pageNumber="148">FOF</collectionCode>
!, isotypes 
<specimenCode box="[274,405,1759,1780]" collectionCode="KAG" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="3" pageNumber="148" type="Herbarium">KAG129297</specimenCode>
!, 
<collectionCode box="[418,471,1759,1780]" country="Japan" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="3" pageNumber="148" type="Herbarium">TNS</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[136,1452,1827,2070]" lastBlockId="4.[136,1451,158,221]" lastPageId="4" lastPageNumber="149" pageId="3" pageNumber="148">
<taxonomicName box="[136,359,1827,1853]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[136,359,1827,1853]" italics="true" pageId="3" pageNumber="148">Lasianthus laoticus</emphasis>
</taxonomicName>
is similar to 
<taxonomicName authority="Poopath, Vajrodaya &amp; Napiroon" authorityName="Poopath, Vajrodaya &amp; Napiroon" box="[519,1053,1827,1854]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="kailarsenii">
<emphasis box="[519,672,1827,1853]" italics="true" pageId="3" pageNumber="148">L. kailarsenii</emphasis>
Poopath, Vajrodaya &amp; Napiroon
</taxonomicName>
in 
<bibRefCitation author="Napiroon, T. &amp; Balslev, H. &amp; Poopath, M. &amp; Chamchumroon, V. &amp; Vajrodaya, S. &amp; Chayamarit, K." box="[1096,1418,1827,1854]" pageId="3" pageNumber="148" pagination="207 - 216" refId="ref4591" refString="Napiroon, T., Balslev, H., Poopath, M., Chamchumroon, V., Vajrodaya, S. &amp; Chayamarit, K. (2018) Three new taxa and a new record of Lasianthus (Rubiaceae) in Thailand. Annales Botanici Fennici 55 (4 - 6): 207 - 216. https: // doi. org / 10.5735 / 085.055.0402" type="journal article" year="2018">
Napiroon 
<emphasis box="[1212,1272,1827,1853]" italics="true" pageId="3" pageNumber="148">et al.</emphasis>
(2018: 208)
</bibRefCitation>
of 
<collectingCountry box="[136,236,1863,1889]" name="Thailand" pageId="3" pageNumber="148">Thailand</collectingCountry>
and 
<taxonomicName authority="Parker (1925: 430)" authorityName="Parker" authorityPageNumber="430" authorityYear="1925" box="[291,748,1863,1890]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="longipedunculatus">
<emphasis box="[291,528,1863,1889]" italics="true" pageId="3" pageNumber="148">L. longipedunculatus</emphasis>
<bibRefCitation author="Parker, R." box="[535,748,1863,1890]" pageId="3" pageNumber="148" pagination="426 - 433" refId="ref4728" refString="Parker, R. (1925) Decades Kewenses. Plantarum Novarum in Herbario Hortii Regii Conservatorum. Kew Bulletin 10: 426 - 433. https: // doi. org / 10.2307 / 4115104" type="journal article" year="1925">Parker (1925: 430)</bibRefCitation>
</taxonomicName>
of 
<collectingCountry box="[786,895,1864,1890]" name="Myanmar" pageId="3" pageNumber="148">Myanmar</collectingCountry>
and 
<collectingCountry box="[950,1050,1863,1889]" name="Thailand" pageId="3" pageNumber="148">Thailand</collectingCountry>
in having conspicuous pedunculate cymes, but distinguished by appressed hairy twigs and petiole (vs. glabrous), reticulate tertiary veins (vs. parallel), and smaller triangle stipule ca. 
<quantity box="[496,584,1936,1962]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="148" unit="mm" value="1.8">1.8 mm</quantity>
long (vs. lanceolate, 
<quantity box="[831,926,1936,1962]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="148" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long in 
<taxonomicName box="[1024,1175,1935,1961]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="kailarsenii">
<emphasis box="[1024,1175,1935,1961]" italics="true" pageId="3" pageNumber="148">L. kailarsenii</emphasis>
</taxonomicName>
and triangular, 
<quantity box="[1357,1452,1935,1962]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="3" pageNumber="148" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long in 
<taxonomicName authorityName="Parker" authorityYear="1925" box="[225,463,1971,1997]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="longipedunculatus">
<emphasis box="[225,463,1971,1997]" italics="true" pageId="3" pageNumber="148">L. longipedunculatus</emphasis>
</taxonomicName>
). In addition to these vegetative traits, 
<taxonomicName box="[912,1032,1971,1997]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[912,1032,1971,1997]" italics="true" pageId="3" pageNumber="148">L. laoticus</emphasis>
</taxonomicName>
differs from 
<taxonomicName box="[1185,1336,1971,1997]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="kailarsenii">
<emphasis box="[1185,1336,1971,1997]" italics="true" pageId="3" pageNumber="148">L. kailarsenii</emphasis>
</taxonomicName>
in having cylindrical-campanulate corolla with triangular lobes 
<quantity box="[740,805,2008,2034]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="148" unit="mm" value="3.0">3 mm</quantity>
long (vs. tuber, lobes oblong-lanceolate 
<quantity box="[1264,1378,2007,2034]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="148" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long), longer style 
<quantity box="[275,341,2044,2070]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="148" unit="mm" value="4.0">4 mm</quantity>
long (vs. 
<quantity box="[453,572,2043,2070]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="148" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
), and from 
<taxonomicName authorityName="Parker" authorityYear="1925" box="[701,938,2043,2069]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="148" phylum="Tracheophyta" rank="species" species="longipedunculatus">
<emphasis box="[701,938,2043,2069]" italics="true" pageId="3" pageNumber="148">L. longipedunculatus</emphasis>
</taxonomicName>
in having shorter peduncles 
<quantity box="[1262,1345,2044,2070]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="148" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long (vs. 
<quantity box="[136,234,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="4" pageNumber="149" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 cm</quantity>
long), smaller linear bracts ca. 
<quantity box="[593,680,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="149" unit="mm" value="2.2">2.2 mm</quantity>
long (vs. leaf-like, linear-lanceolate, 
<quantity box="[1104,1212,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="4" pageNumber="149" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
long), and triangular calyx lobes ca. 
<quantity box="[309,374,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="149" unit="mm" value="1.0">1 mm</quantity>
long (vs. oblong-lanceolate, 
<quantity box="[700,814,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="4" pageNumber="149" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
long) (
<tableCitation box="[894,982,194,220]" captionStart="Table 2" captionStartId="4.[136,199,268,292]" captionTargetBox="[144,1442,377,954]" captionTargetPageId="4" captionText="Table 2. Comparison of Lasianthus laoticus with three morphologically similar species. Data on L. kailarsenii, and L. longipedunculatus and L. lancilobus were retrieved from the protologue Napiroon et al. (2018), Parker (1925) and Zhu (2012), as well as their type specimens, respectively." pageId="4" pageNumber="149">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
<caption pageId="4" pageNumber="149" startId="4.[136,199,268,292]" targetBox="[144,1442,377,954]" targetIsTable="true" targetPageId="4">
<paragraph blockId="4.[136,1452,268,364]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[136,229,268,292]" pageId="4" pageNumber="149">Table 2.</emphasis>
Comparison of 
<taxonomicName box="[413,624,268,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[413,624,268,292]" italics="true" pageId="4" pageNumber="149">Lasianthus laoticus</emphasis>
</taxonomicName>
with three morphologically similar species. Data on 
<taxonomicName box="[1218,1364,268,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="kailarsenii">
<emphasis box="[1218,1364,268,292]" italics="true" pageId="4" pageNumber="149">L. kailarsenii</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[1430,1452,269,292]" pageId="4" pageNumber="149">
<emphasis box="[1430,1452,269,292]" italics="true" pageId="4" pageNumber="149">L.</emphasis>
</taxonomicName>
<emphasis box="[136,333,304,328]" italics="true" pageId="4" pageNumber="149">longipedunculatus</emphasis>
and 
<taxonomicName box="[391,533,304,328]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="lancilobus">
<emphasis box="[391,533,304,328]" italics="true" pageId="4" pageNumber="149">L. lancilobus</emphasis>
</taxonomicName>
were retrieved from the protologue 
<bibRefCitation author="Napiroon, T. &amp; Balslev, H. &amp; Poopath, M. &amp; Chamchumroon, V. &amp; Vajrodaya, S. &amp; Chayamarit, K." box="[937,1187,304,328]" pageId="4" pageNumber="149" pagination="207 - 216" refId="ref4591" refString="Napiroon, T., Balslev, H., Poopath, M., Chamchumroon, V., Vajrodaya, S. &amp; Chayamarit, K. (2018) Three new taxa and a new record of Lasianthus (Rubiaceae) in Thailand. Annales Botanici Fennici 55 (4 - 6): 207 - 216. https: // doi. org / 10.5735 / 085.055.0402" type="journal article" year="2018">
Napiroon 
<emphasis box="[1047,1104,304,328]" italics="true" pageId="4" pageNumber="149">et al.</emphasis>
(2018)
</bibRefCitation>
, 
<bibRefCitation author="Parker, R." box="[1200,1351,304,328]" pageId="4" pageNumber="149" pagination="426 - 433" refId="ref4728" refString="Parker, R. (1925) Decades Kewenses. Plantarum Novarum in Herbario Hortii Regii Conservatorum. Kew Bulletin 10: 426 - 433. https: // doi. org / 10.2307 / 4115104" type="journal article" year="1925">Parker (1925)</bibRefCitation>
and 
<bibRefCitation author="Zhu, H. &amp; Roos, M. C. &amp; Ridsdale, C. E." box="[1408,1451,304,328]" pageId="4" pageNumber="149" pagination="1 - 102" refId="ref5152" refString="Zhu, H., Roos, M. C. &amp; Ridsdale, C. E. (2012) A taxonomic revision of the Malesian species of Lasianthus (Rubiaceae). Blumea 57: 1 - 102. https: // doi. org / 10.3767 / 000651912 X 652012" type="journal article" year="2012">Zhu</bibRefCitation>
(2012), as well as their 
<typeStatus box="[386,432,340,364]" pageId="4" pageNumber="149">type</typeStatus>
specimens, respectively.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="149">
<table box="[144,1442,377,954]" gridcols="5" gridrows="19" pageId="4" pageNumber="149">
<tr box="[144,1442,377,399]" gridrow="0" pageId="4" pageNumber="149">
<th box="[144,311,377,399]" gridcol="0" gridrow="0" pageId="4" pageNumber="149">
<emphasis bold="true" box="[144,259,377,399]" pageId="4" pageNumber="149">Characters</emphasis>
</th>
<th box="[353,613,377,399]" gridcol="1" gridrow="0" pageId="4" pageNumber="149">
<taxonomicName box="[353,547,377,399]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis bold="true" box="[353,547,377,399]" italics="true" pageId="4" pageNumber="149">Lasianthus laoticus</emphasis>
</taxonomicName>
</th>
<th box="[630,880,377,399]" gridcol="2" gridrow="0" pageId="4" pageNumber="149">
<taxonomicName box="[630,761,377,399]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="kailarsenii">
<emphasis bold="true" box="[630,761,377,399]" italics="true" pageId="4" pageNumber="149">L. kailarsenii</emphasis>
</taxonomicName>
</th>
<th box="[906,1164,377,399]" gridcol="3" gridrow="0" pageId="4" pageNumber="149">
<taxonomicName authorityName="Parker" authorityYear="1925" box="[906,1117,377,399]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="longipedunculatus">
<emphasis bold="true" box="[906,1117,377,399]" italics="true" pageId="4" pageNumber="149">L. longipedunculatus</emphasis>
</taxonomicName>
</th>
<th box="[1183,1442,377,399]" gridcol="4" gridrow="0" pageId="4" pageNumber="149">
<emphasis bold="true" box="[1183,1312,377,399]" italics="true" pageId="4" pageNumber="149">L. lancilobus</emphasis>
</th>
</tr>
<tr box="[144,1442,409,431]" gridrow="1" pageId="4" pageNumber="149">
<th box="[144,311,409,431]" gridcol="0" gridrow="1" pageId="4" pageNumber="149">Lamina</th>
<td box="[353,613,409,431]" gridcol="1" gridrow="1" pageId="4" pageNumber="149">ovate-oblong, 3.5–13 ×</td>
<td box="[630,880,409,431]" gridcol="2" gridrow="1" pageId="4" pageNumber="149">lanceolate to lanceolate-</td>
<td box="[906,1164,409,431]" gridcol="3" gridrow="1" pageId="4" pageNumber="149">oblong, 15–20 × 4–6 cm</td>
<td box="[1183,1442,409,431]" gridcol="4" gridrow="1" pageId="4" pageNumber="149">oblong, 6–13 × 2.5–3.5 cm</td>
</tr>
<tr box="[144,1442,438,460]" gridrow="2" pageId="4" pageNumber="149" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<td box="[353,613,438,460]" gridcol="1" gridrow="2" pageId="4" pageNumber="149">1.4–4.1 cm</td>
<td box="[630,880,438,460]" gridcol="2" gridrow="2" pageId="4" pageNumber="149">elliptic, 5–7 × 2.5–3 cm,</td>
</tr>
<tr box="[144,1442,470,492]" gridrow="3" pageId="4" pageNumber="149">
<th box="[144,311,470,492]" gridcol="0" gridrow="3" pageId="4" pageNumber="149">Petiole length</th>
<td box="[353,613,470,492]" gridcol="1" gridrow="3" pageId="4" pageNumber="149">1–3 mm</td>
<td box="[630,880,470,492]" gridcol="2" gridrow="3" pageId="4" pageNumber="149">1 mm</td>
<td box="[906,1164,470,492]" gridcol="3" gridrow="3" pageId="4" pageNumber="149">1–2 mm</td>
<td box="[1183,1442,470,492]" gridcol="4" gridrow="3" pageId="4" pageNumber="149">6–8 mm</td>
</tr>
<tr box="[144,1442,502,524]" gridrow="4" pageId="4" pageNumber="149">
<th box="[144,311,502,524]" gridcol="0" gridrow="4" pageId="4" pageNumber="149">Hairlines on twig</th>
<td box="[353,613,502,524]" gridcol="1" gridrow="4" pageId="4" pageNumber="149">appressed hairy</td>
<td box="[630,880,502,524]" gridcol="2" gridrow="4" pageId="4" pageNumber="149">glabrous except base</td>
<td box="[906,1164,502,524]" gridcol="3" gridrow="4" pageId="4" pageNumber="149">glabrous</td>
<td box="[1183,1442,502,524]" gridcol="4" gridrow="4" pageId="4" pageNumber="149">(sparsely) hirsute</td>
</tr>
<tr box="[144,1442,531,553]" gridrow="5" pageId="4" pageNumber="149" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<th box="[144,311,531,553]" gridcol="0" gridrow="5" pageId="4" pageNumber="149">and petiole</th>
<td box="[630,880,531,553]" gridcol="2" gridrow="5" pageId="4" pageNumber="149">which is sparsely hirsute</td>
</tr>
<tr box="[144,1442,563,585]" gridrow="6" pageId="4" pageNumber="149">
<th box="[144,311,563,585]" gridcol="0" gridrow="6" pageId="4" pageNumber="149">Number of</th>
<td box="[353,613,563,585]" gridcol="1" gridrow="6" pageId="4" pageNumber="149">8–11 pairs</td>
<td box="[630,880,563,585]" gridcol="2" gridrow="6" pageId="4" pageNumber="149">6–8 pairs</td>
<td box="[906,1164,563,585]" gridcol="3" gridrow="6" pageId="4" pageNumber="149">6–8 pairs</td>
<td box="[1183,1442,563,585]" gridcol="4" gridrow="6" pageId="4" pageNumber="149">5–6 pairs</td>
</tr>
<tr box="[144,1442,592,614]" gridrow="7" pageId="4" pageNumber="149" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th box="[144,311,592,614]" gridcol="0" gridrow="7" pageId="4" pageNumber="149">secondary veins</th>
</tr>
<tr box="[144,1442,624,646]" gridrow="8" pageId="4" pageNumber="149">
<th box="[144,311,624,646]" gridcol="0" gridrow="8" pageId="4" pageNumber="149">Tertiary veins</th>
<td box="[353,613,624,646]" gridcol="1" gridrow="8" pageId="4" pageNumber="149">reticulate</td>
<td box="[630,880,624,646]" gridcol="2" gridrow="8" pageId="4" pageNumber="149">parallel</td>
<td box="[906,1164,624,646]" gridcol="3" gridrow="8" pageId="4" pageNumber="149">parallel</td>
<td box="[1183,1442,624,646]" gridcol="4" gridrow="8" pageId="4" pageNumber="149">subparallel</td>
</tr>
<tr box="[144,1442,655,677]" gridrow="9" pageId="4" pageNumber="149">
<th box="[144,311,655,677]" gridcol="0" gridrow="9" pageId="4" pageNumber="149">Stipules</th>
<td box="[353,613,655,677]" gridcol="1" gridrow="9" pageId="4" pageNumber="149">triangular, ca. 1.8 mm long</td>
<td box="[630,880,655,677]" gridcol="2" gridrow="9" pageId="4" pageNumber="149">lanceolate, 3–4 mm long</td>
<td box="[906,1164,655,677]" gridcol="3" gridrow="9" pageId="4" pageNumber="149">triangular, 2–3(–5) mm</td>
<td box="[1183,1442,655,677]" gridcol="4" gridrow="9" pageId="4" pageNumber="149">Lanceolate, ca. 3 mm long</td>
</tr>
<tr box="[144,1442,685,707]" gridrow="10" pageId="4" pageNumber="149" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td box="[906,1164,685,707]" gridcol="3" gridrow="10" pageId="4" pageNumber="149">long</td>
</tr>
<tr box="[144,1442,716,739]" gridrow="11" pageId="4" pageNumber="149">
<th box="[144,311,716,739]" gridcol="0" gridrow="11" pageId="4" pageNumber="149">Peduncle length</th>
<td box="[353,613,716,739]" gridcol="1" gridrow="11" pageId="4" pageNumber="149">1–3 cm</td>
<td box="[630,880,716,739]" gridcol="2" gridrow="11" pageId="4" pageNumber="149">2.5–4 cm</td>
<td box="[906,1164,716,739]" gridcol="3" gridrow="11" pageId="4" pageNumber="149">4–10 cm</td>
<td box="[1183,1442,716,739]" gridcol="4" gridrow="11" pageId="4" pageNumber="149">2.5–3 cm</td>
</tr>
<tr box="[144,1442,748,770]" gridrow="12" pageId="4" pageNumber="149">
<th box="[144,311,748,770]" gridcol="0" gridrow="12" pageId="4" pageNumber="149">Bracts</th>
<td box="[353,613,748,770]" gridcol="1" gridrow="12" pageId="4" pageNumber="149">linear, ca. 2.2 mm long</td>
<td box="[630,880,748,770]" gridcol="2" gridrow="12" pageId="4" pageNumber="149">lanceolate, 2–3 mm long</td>
<td box="[906,1164,748,770]" gridcol="3" gridrow="12" pageId="4" pageNumber="149">leaf-like, linear-lanceolate,</td>
<td box="[1183,1442,748,770]" gridcol="4" gridrow="12" pageId="4" pageNumber="149">linear, up to 15 mm long</td>
</tr>
<tr box="[144,1442,778,800]" gridrow="13" pageId="4" pageNumber="149" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td box="[906,1164,778,800]" gridcol="3" gridrow="13" pageId="4" pageNumber="149">8–12 mm long</td>
</tr>
<tr box="[144,1442,809,831]" gridrow="14" pageId="4" pageNumber="149">
<th box="[144,311,809,831]" gridcol="0" gridrow="14" pageId="4" pageNumber="149">Calyx lobes</th>
<td box="[353,613,809,831]" gridcol="1" gridrow="14" pageId="4" pageNumber="149">triangular, ca. 1 mm long</td>
<td box="[630,880,809,831]" gridcol="2" gridrow="14" pageId="4" pageNumber="149">triangular, 1–1.5 mm long</td>
<td box="[906,1164,809,831]" gridcol="3" gridrow="14" pageId="4" pageNumber="149">oblong-lanceolate, 4–4.5</td>
<td box="[1183,1442,809,831]" gridcol="4" gridrow="14" pageId="4" pageNumber="149">lanceolate, 5 mm long</td>
</tr>
<tr box="[144,1442,839,861]" gridrow="15" pageId="4" pageNumber="149" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td box="[906,1164,839,861]" gridcol="3" gridrow="15" pageId="4" pageNumber="149">mm long</td>
</tr>
<tr box="[144,1442,871,893]" gridrow="16" pageId="4" pageNumber="149">
<th box="[144,311,871,893]" gridcol="0" gridrow="16" pageId="4" pageNumber="149">Corolla lobes</th>
<td box="[353,613,871,893]" gridcol="1" gridrow="16" pageId="4" pageNumber="149">triangular, ca. 3 mm long</td>
<td box="[630,880,871,893]" gridcol="2" gridrow="16" pageId="4" pageNumber="149">oblong-lanceolate, with</td>
<td box="[906,1164,871,893]" gridcol="3" gridrow="16" pageId="4" pageNumber="149">oblong, 2 mm long</td>
<td box="[1183,1442,871,893]" gridcol="4" gridrow="16" pageId="4" pageNumber="149">-</td>
</tr>
<tr box="[144,1442,900,922]" gridrow="17" pageId="4" pageNumber="149" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1">
<td box="[630,880,900,922]" gridcol="2" gridrow="17" pageId="4" pageNumber="149">lobes 1.5–2 mm long</td>
</tr>
<tr box="[144,1442,931,954]" gridrow="18" pageId="4" pageNumber="149">
<th box="[144,311,931,954]" gridcol="0" gridrow="18" pageId="4" pageNumber="149">Style length</th>
<td box="[353,613,931,954]" gridcol="1" gridrow="18" pageId="4" pageNumber="149">4 mm</td>
<td box="[630,880,931,954]" gridcol="2" gridrow="18" pageId="4" pageNumber="149">1.5–2 mm</td>
<td box="[906,1164,931,954]" gridcol="3" gridrow="18" pageId="4" pageNumber="149">-</td>
<td box="[1183,1442,931,954]" gridcol="4" gridrow="18" pageId="4" pageNumber="149">-</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="4" pageNumber="149" type="description">
<paragraph blockId="4.[136,1452,997,2068]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,327,997,1023]" pageId="4" pageNumber="149">Description</emphasis>
:—Shrub, to 
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high. Current year twig 
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in diam., flattened, appressed brownish hairy, glabrescent; internode 
<quantity box="[395,526,1034,1060]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.2" metricValueMin="1.3" pageId="4" pageNumber="149" unit="cm" value="2.75" valueMax="4.2" valueMin="1.3">1.3–4.2 cm</quantity>
. Leaves opposite, petiolate; petiole 
<quantity box="[934,1029,1034,1060]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="149" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
long, densely appressed hairy; blade ovate-oblong, 3.5–13 × 
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, subcoriaceous, adaxially dark yellow green, glossy, glabrous, abaxially gray green, densely hairy on midrib to tertiary veins, apex acuminate to aristate, tip pointed, 
<quantity box="[1103,1237,1106,1132]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="4" pageNumber="149" unit="mm" value="1.1" valueMax="1.4" valueMin="0.8">0.8–1.4 mm</quantity>
long, base cordate, margin finely undulate, midrib sunken adaxially, prominent abaxially, secondary veins 8–11 pairs, prominent abaxially, tertiary veins reticulate. Stipules triangular, ca. 
<quantity box="[676,764,1178,1204]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="149" unit="mm" value="1.8">1.8 mm</quantity>
long, brownish hairy outside, especially denser near margin and apex. Inflorescence axillary, cymose, solitary to 7-flowered; peduncle 
<quantity box="[959,1061,1214,1240]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="3.0" metricValueMin="1.3" pageId="4" pageNumber="149" unit="cm" value="2.15" valueMax="3.0" valueMin="1.3">1.3–3 cm</quantity>
long, pubescent, glabrescent; bract linear, ca. 
<quantity box="[253,339,1250,1276]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="149" unit="mm" value="2.2">2.2 mm</quantity>
long, pubescent. Flowers subsessile to short-stalked; pedicels up to 
<quantity box="[1108,1195,1250,1276]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="149" unit="mm" value="2.2">2.2 mm</quantity>
long; calyx obconical, sparsely pubescent, tube ca. 
<quantity box="[456,522,1286,1312]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="149" unit="mm" value="2.0">2 mm</quantity>
long, lobes 4, triangular, ca. 
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long; corolla white with a faint tinge of yellow, cylindrical-campanulate, tube 
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long, outside glabrous, inside villous in upper 1/2, lobes triangular, ca. 
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long, apex apiculate, outside glabrous, inside villous, stamens 4, exerted from corolla, filaments basally connate to corolla, free part ca. 
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long, glabrous, anthers ca. 
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long, ciliolate; ovary 4-locular, style ca. 
<quantity box="[1321,1387,1394,1420]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="149" unit="mm" value="4.0">4 mm</quantity>
long, glabrous, stigma 4-lobed, ca. 
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long. Fruits drupaceous, subglobose or ovoid, 
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long, ca. 
<quantity box="[1281,1346,1429,1456]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="149" unit="mm" value="5.0">5 mm</quantity>
in diam., crowned by the persistent calyx, blackish brown when ripe, sparsely appressed hairy on the surface; the persistent calyx lobes 4, linear to linear-lanceolate, ca. 
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long, apex acute, hirsute; pyrenes 4, grooved on the dorsal curved face, obovoid, ca. 
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long, 
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wide.
</paragraph>
<paragraph blockId="4.[136,1452,997,2068]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,589,1573,1599]" pageId="4" pageNumber="149">Additional specimen examined:—</emphasis>
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, 
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, 
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, 
<geoCoordinate box="[136,304,1611,1638]" degrees="106" direction="east" minutes="01" orientation="longitude" pageId="4" pageNumber="149" precision="1" seconds="2.36" value="106.01733">106°01′2.36″E</geoCoordinate>
, 
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<elevation box="[315,386,1609,1636]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="4" pageNumber="149" unit="m" value="158.0">158 m</elevation>
</quantity>
elev., 
<date box="[459,675,1609,1636]" pageId="4" pageNumber="149" value="2018-12-13">
<collectingDate box="[459,675,1609,1636]" pageId="4" pageNumber="149" value="2018-12-13">13 December 2018</collectingDate>
</date>
, 
<emphasis box="[686,936,1609,1636]" italics="true" pageId="4" pageNumber="149">
<collectorName box="[686,795,1610,1636]" pageId="4" pageNumber="149">S. Tagane</collectorName>
et al. L2222
</emphasis>
[young infl.] (
<collectionCode box="[1100,1154,1610,1636]" pageId="4" pageNumber="149">FOF</collectionCode>
, 
<specimenCode box="[1165,1315,1610,1636]" collectionCode="KAG" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="4" pageNumber="149" type="Herbarium">KAG128322</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="149" type="distribution">
<paragraph blockId="4.[136,1452,997,2068]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,373,1645,1671]" pageId="4" pageNumber="149">Distribution:—</emphasis>
<collectingCountry box="[373,427,1646,1672]" name="Laos" pageId="4" pageNumber="149">Laos</collectingCountry>
(so far known only from the two provinces, 
<collectingRegion box="[934,1023,1646,1672]" country="Laos" name="Attapu" pageId="4" pageNumber="149">Attapeu</collectingRegion>
and 
<collectingRegion box="[1079,1210,1645,1671]" country="Laos" name="Champasak" pageId="4" pageNumber="149">Champasak</collectingRegion>
Provinces, located at the southern basin of Bolaven Plateau).
</paragraph>
<paragraph blockId="4.[136,1452,997,2068]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,475,1717,1744]" pageId="4" pageNumber="149">Habitat and Ecology:—</emphasis>
The species was found in lowland evergreen forest at 
<quantity box="[1090,1219,1717,1744]" metricMagnitude="2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="4" pageNumber="149" unit="m" value="170.0" valueMax="190.0" valueMin="150.0">150–190 m</quantity>
elev. Flowering and fruiting specimens were collected in July.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="151" pageId="4" pageNumber="149" type="etymology">
<paragraph blockId="4.[136,1452,997,2068]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,356,1790,1816]" pageId="4" pageNumber="149">Etymology:—</emphasis>
This specific epithet 
<taxonomicName box="[620,709,1789,1815]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[620,709,1789,1815]" italics="true" pageId="4" pageNumber="149">laoticus</emphasis>
</taxonomicName>
is derived from the country name where this species was discovered.
</paragraph>
<paragraph blockId="4.[136,1452,997,2068]" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,437,1862,1888]" pageId="4" pageNumber="149">Vernacular name:—</emphasis>
[(Xangsan Lao, “Xangsan” means 
<taxonomicName box="[1064,1187,1861,1887]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis box="[1064,1187,1861,1887]" italics="true" pageId="4" pageNumber="149">Lasianthus</emphasis>
</taxonomicName>
in general in Lao and “Lao” refers to the country where the species is discovered), proposed here].
</paragraph>
<paragraph blockId="4.[136,1452,997,2068]" lastBlockId="5.[136,1451,158,292]" lastPageId="5" lastPageNumber="150" pageId="4" pageNumber="149">
<emphasis bold="true" box="[189,674,1933,1960]" pageId="4" pageNumber="149">Preliminary conservation assessment:—</emphasis>
Endangered (EN). To date, 
<taxonomicName box="[992,1213,1933,1959]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="149" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[992,1213,1933,1959]" italics="true" pageId="4" pageNumber="149">Lasianthus laoticus</emphasis>
</taxonomicName>
is known only from two populations with each ca. 30 mature individuals in the lowland evergreen forest in lower elevations at the southern basin of Bolaven Plateau, at elevation of 158 and 
<specimenCount box="[709,789,2006,2032]" pageId="4" pageNumber="149" type="generic">
<quantity box="[709,785,2006,2032]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="149" unit="m" value="183.0">183 m</quantity>
.
</specimenCount>
Considering the distance between the two localities is ca. 
<quantity box="[136,208,2041,2068]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="149" unit="km" value="18.0">18 km</quantity>
and same environment is found in the area, the estimated extent of occupancy for this species is 
<quantity box="[1300,1386,2041,2068]" metricMagnitude="5" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="149" unit="km" value="900.0">900 km</quantity>
<superScript attach="left" box="[1386,1394,2041,2055]" fontSize="6" pageId="4" pageNumber="149">2</superScript>
. The localities we collected the above specimens are along a recently constructed logging road, and are being gradually deforested. Therefore, we suggest the category of this species as Endangered according to the IUCN criteria B1ab(i, ii, iii) (
<bibRefCitation author="IUCN" box="[215,354,230,256]" pageId="5" pageNumber="150" refId="ref4467" refString="IUCN (2012) IUCN Red List Categories and Criteria, version 3.1. Gland and Cambridge. Available from: http: // www. iucnredlist. org (accessed 24 March 2021)" type="url" year="2012">IUCN 2012</bibRefCitation>
). More accurate data on its population size and number of individuals is needed for developing a conservation plan.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7681616" ID-Zenodo-Dep="7681616" box="[136,1254,2042,2064]" httpUri="https://zenodo.org/record/7681616/files/figure.png" pageId="5" pageNumber="150" startId="5.[136,229,2042,2064]" targetBox="[199,1385,331,2014]" targetPageId="5">
<paragraph blockId="5.[136,1254,2042,2064]" box="[136,1254,2042,2064]" pageId="5" pageNumber="150">
<emphasis bold="true" box="[136,247,2042,2064]" pageId="5" pageNumber="150">FIGURE 3</emphasis>
. Isotype of 
<taxonomicName authority="Tagane, Soulad. &amp; Nob. Tanaka." authorityName="Tagane, Soulad. &amp; Nob. Tanaka." box="[362,863,2042,2064]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[362,549,2042,2063]" italics="true" pageId="5" pageNumber="150">Lasianthus laoticus</emphasis>
Tagane, Soulad. &amp; Nob.Tanaka.
</taxonomicName>
(
<emphasis box="[876,1093,2042,2064]" italics="true" pageId="5" pageNumber="150">Souladeth et al. L3024</emphasis>
(KAG129297)).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7681618" ID-Zenodo-Dep="7681618" httpUri="https://zenodo.org/record/7681618/files/figure.png" pageId="6" pageNumber="151" startId="6.[136,229,1310,1332]" targetBox="[283,1303,190,1286]" targetPageId="6">
<paragraph blockId="6.[136,1452,1310,1368]" pageId="6" pageNumber="151">
<emphasis bold="true" box="[136,248,1310,1332]" pageId="6" pageNumber="151">FIGURE 4</emphasis>
. 
<taxonomicName authority="Tagane, Soulad. &amp; Nob. Tanaka. A. Flowering" box="[260,894,1310,1332]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[260,448,1310,1331]" italics="true" pageId="6" pageNumber="151">Lasianthus laoticus</emphasis>
Tagane, Soulad. &amp; Nob.Tanaka. A. Flowering
</taxonomicName>
twig; B. Portion of abaxial leaf surface; C. Inflorescence; D. Calyx; E. Fruit. All from 
<emphasis box="[409,626,1346,1368]" italics="true" pageId="6" pageNumber="151">Souladeth et al. L3024</emphasis>
.
</paragraph>
</caption>
<paragraph blockId="6.[136,1452,1415,1585]" pageId="6" pageNumber="151">
<emphasis bold="true" box="[189,294,1415,1441]" pageId="6" pageNumber="151">Notes:—</emphasis>
Among the species of 
<taxonomicName box="[553,676,1415,1441]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="genus">
<emphasis box="[553,676,1415,1441]" italics="true" pageId="6" pageNumber="151">Lasianthus</emphasis>
</taxonomicName>
in 
<collectingCountry box="[716,773,1415,1441]" name="Laos" pageId="6" pageNumber="151">Laos</collectingCountry>
, 
<taxonomicName box="[786,907,1415,1441]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="species" species="laoticus">
<emphasis box="[786,907,1415,1441]" italics="true" pageId="6" pageNumber="151">L. laoticus</emphasis>
</taxonomicName>
is easily distinguished by its slender peduncles (vs. inflorescence sessile or subsessile in the other species). It is also similar to 
<taxonomicName box="[1040,1188,1451,1477]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="species" species="lancilobus">
<emphasis box="[1040,1188,1451,1477]" italics="true" pageId="6" pageNumber="151">L. lancilobus</emphasis>
</taxonomicName>
of West Kalimantan in lamina size and slender peduncles but different from 
<taxonomicName box="[727,873,1487,1513]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="species" species="lancilobus">
<emphasis box="[727,873,1487,1513]" italics="true" pageId="6" pageNumber="151">L. lancilobus</emphasis>
</taxonomicName>
in having shorter petiole 
<quantity box="[1157,1249,1487,1513]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="6" pageNumber="151" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
long (vs. 
<quantity box="[1359,1452,1486,1513]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="151" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long in 
<taxonomicName box="[221,367,1523,1549]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="species" species="lancilobus">
<emphasis box="[221,367,1523,1549]" italics="true" pageId="6" pageNumber="151">L. lancilobus</emphasis>
</taxonomicName>
), cordate leaf base (vs. acute to cuneate), 8–11 pairs of secondary veins (vs. 5–6 pairs), and small triangular calyx lobe 
<quantity box="[375,440,1559,1585]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="151" unit="mm" value="1.0">1 mm</quantity>
long (vs. lanceolate, 
<quantity box="[679,744,1558,1585]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="151" unit="mm" value="5.0">5 mm</quantity>
long) (
<tableCitation box="[824,912,1559,1585]" captionStart="Table 2" captionStartId="4.[136,199,268,292]" captionTargetBox="[144,1442,377,954]" captionTargetPageId="4" captionText="Table 2. Comparison of Lasianthus laoticus with three morphologically similar species. Data on L. kailarsenii, and L. longipedunculatus and L. lancilobus were retrieved from the protologue Napiroon et al. (2018), Parker (1925) and Zhu (2012), as well as their type specimens, respectively." pageId="6" pageNumber="151">Table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>