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<document ID-DOI="10.11646/phytotaxa.585.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="7681608" checkinTime="1677506995801" checkinUser="plazi" docAuthor="Tagane, Shuichiro, Souladeth, Phetlasy, Tanaka, Nobuyuki, Phengmala, Kachonesak &amp; Yahara, Tetsukazu" docDate="2023" docId="731884124921FF9CFF1D9F88FC99FEE5" docLanguage="en" docName="phytotaxa.585.2.5.pdf" docOrigin="Phytotaxa 585 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.585.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Lasianthus repoeuensis Pierre ex Pitard 1924" docType="treatment" docVersion="1" lastPageNumber="152" masterDocId="8F21FC6A4927FF9BFF95992EFFA7FFE5" masterDocTitle="Two new species and a new record of Lasianthus (Rubiaceae) in Laos" masterLastPageNumber="153" masterPageNumber="145" pageNumber="151" updateTime="1677613052640" updateUser="diego" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species and a new record of Lasianthus (Rubiaceae) in Laos</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tagane, Shuichiro</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-1974-7329</mods:nameIdentifier>
<mods:affiliation>The Kagoshima University Museum, Kagoshima University, Kagoshima, Japan &amp; stagane 29 @ gmail. com; https: // orcid. org / 0000 - 0002 - 1974 - 7329</mods:affiliation>
<mods:nameIdentifier type="email">stagane29@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Souladeth, Phetlasy</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8563-1773</mods:nameIdentifier>
<mods:affiliation>Faculty of Forest Science, National University of Laos, Vientiane, Laos &amp; p. souladeth @ nuol. edu. la; https: // orcid. org / 0000 - 0001 - 8563 - 1773</mods:affiliation>
<mods:nameIdentifier type="email">p.souladeth@nuol.edu.la</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tanaka, Nobuyuki</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0229-8061</mods:nameIdentifier>
<mods:affiliation>Department of Botany, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan &amp; nobuyuki _ tanaka @ kahaku. go. jp; https: // orcid. org / 0000 - 0002 - 0229 - 8061</mods:affiliation>
<mods:nameIdentifier type="email">nobuyuki_tanaka@kahaku.go.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Phengmala, Kachonesak</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1508-0704</mods:nameIdentifier>
<mods:affiliation>Faculty of Forest Science, National University of Laos, Vientiane, Laos &amp; Kachonesak @ gmail. com; https: // orcid. org / 0000 - 0003 - 1508 - 0704</mods:affiliation>
<mods:nameIdentifier type="email">achonesak@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yahara, Tetsukazu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-5105-7152</mods:nameIdentifier>
<mods:affiliation>Kyushu Open University, 744 Motooka, Fukuoka, 819 - 0395, Japan &amp; tet. yahara @ gmail. com; https: // orcid. org / 0000 - 0001 - 5105 - 7152</mods:affiliation>
<mods:nameIdentifier type="email">tet.yahara@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>585</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>153</mods:end>
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<mods:url>http://dx.doi.org/10.11646/phytotaxa.585.2.5</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.585.2.5</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7681608</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:731884124921FF9CFF1D9F88FC99FEE5" httpUri="http://treatment.plazi.org/id/731884124921FF9CFF1D9F88FC99FEE5" lastPageId="7" lastPageNumber="152" pageId="6" pageNumber="151">
<subSubSection box="[136,733,1702,1729]" pageId="6" pageNumber="151" type="nomenclature">
<paragraph blockId="6.[136,1452,1702,1765]" box="[136,733,1702,1729]" pageId="6" pageNumber="151">
<treatmentCitationGroup box="[136,733,1702,1729]" pageId="6" pageNumber="151">
<treatmentCitation author="Pitard, J." box="[136,729,1702,1729]" page=")" pageId="6" pageNumber="151" year="1924">
<taxonomicName ID-CoL="3SCRM" authority="Pierre ex Pitard (1924: 387)" authorityName="Pierre ex Pitard" authorityPageNumber="387" authorityYear="1924" box="[136,729,1702,1729]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="species" species="repoeuensis">
<emphasis bold="true" box="[136,409,1702,1729]" italics="true" pageId="6" pageNumber="151">Lasianthus repoeuensis</emphasis>
Pierre ex
<bibRefCitation author="Pitard, J." box="[522,729,1703,1729]" pageId="6" pageNumber="151" pagination="371 - 399" refId="ref4804" refString="Pitard, J. (1924) Lasianthus Jack. In: Lecomte, P. H., Humbert, H. &amp; Gagnepain, F. (Eds.) Flore generale de l'Indo-Chine 3. Masson, Paris, pp. 371 - 399." type="book chapter" year="1924">Pitard (1924: 387)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="151" type="materials_examined">
<paragraph blockId="6.[136,1452,1702,1765]" pageId="6" pageNumber="151">
<location LSID="urn:lsid:plazi:treatment:731884124921FF9CFF1D9F88FC99FEE5:FE6E63DF4921FF9DFD769F89FCBAF924" box="[739,797,1703,1729]" country="Cambodia" municipality="Montibus Knang Krepoeu" name="Type" pageId="6" pageNumber="151">Type</location>
:—
<collectingCountry box="[830,987,1703,1729]" name="Cambodia" pageId="6" pageNumber="151">CAMBODIA</collectingCountry>
.
<collectingMunicipality box="[996,1290,1703,1729]" pageId="6" pageNumber="151">Montibus Knang Krepoeu</collectingMunicipality>
,
<date box="[1301,1418,1703,1729]" pageId="6" pageNumber="151" value="1870-05">
<collectingDate box="[1301,1418,1703,1729]" pageId="6" pageNumber="151" value="1870-05">May 1870</collectingDate>
</date>
,
<emphasis italics="true" pageId="6" pageNumber="151">
<collectorName pageId="6" pageNumber="151">L. Pierre</collectorName>
1257
</emphasis>
(
<typeStatus box="[284,384,1739,1765]" pageId="6" pageNumber="151">holotype</typeStatus>
A00096531 image!, isotypes P02285238, P3981160, images!).
<figureCitation box="[1099,1193,1738,1765]" captionStart="FIGURE 5" captionStartId="7.[136,229,850,872]" captionTargetBox="[283,1303,281,825]" captionTargetId="figure-59@7.[283,1303,281,825]" captionTargetPageId="7" captionText="FIGURE 5. Lasianthus repoeuensis Pierre ex Pit. A. Leafy twig; B. Calyx; C. Portion of abaxial leaf surface. All from Souladeth et al. L3067." figureDoi="http://doi.org/10.5281/zenodo.7681620" httpUri="https://zenodo.org/record/7681620/files/figure.png" pageId="6" pageNumber="151">Figure 5</figureCitation>
</paragraph>
<paragraph blockId="6.[136,1451,1810,2017]" pageId="6" pageNumber="151">
<emphasis bold="true" box="[136,411,1810,1836]" pageId="6" pageNumber="151">Specimen examined:—</emphasis>
<materialsCitation collectingDate="2019-07-06" collectionCode="FOF, KAG, TNS" collectorName="P. Souladeth" country="Laos" elevation="223" latitude="14.805278" location="Bolaven Plateau" longLatPrecision="21" longitude="106.44417" municipality="Sanamxai Diostrict" pageId="6" pageNumber="151" specimenCount="1" stateProvince="Attapu">
<collectingCountry box="[411,488,1811,1837]" name="Laos" pageId="6" pageNumber="151">LAOS</collectingCountry>
.
<collectingRegion box="[497,696,1811,1837]" country="Laos" name="Attapu" pageId="6" pageNumber="151">Attapeu Province</collectingRegion>
,
<collectingMunicipality box="[708,927,1811,1837]" pageId="6" pageNumber="151">Sanamxai Diostrict</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:731884124921FF9CFF1D9F88FC99FEE5:FE6E63DF4921FF9DFC3F9E3DFBC3F8C8" box="[938,1124,1811,1837]" country="Laos" latitude="14.805278" longLatPrecision="21" longitude="106.44417" municipality="Sanamxai Diostrict" name="Bolaven Plateau" pageId="6" pageNumber="151" stateProvince="Attapu">Bolaven Plateau</location>
, in lowland evergreen forest,
<geoCoordinate box="[136,271,1848,1875]" degrees="14" direction="north" minutes="48" orientation="latitude" pageId="6" pageNumber="151" precision="15" seconds="19" value="14.805278">14°4819″N</geoCoordinate>
,
<geoCoordinate box="[283,430,1848,1875]" degrees="106" direction="east" minutes="26" orientation="longitude" pageId="6" pageNumber="151" precision="15" seconds="39" value="106.44417">106°2639″E</geoCoordinate>
,
<quantity box="[441,512,1847,1873]" metricMagnitude="2" metricUnit="m" metricValue="2.23" pageId="6" pageNumber="151" unit="m" value="223.0">
<elevation box="[441,512,1847,1873]" metricMagnitude="2" metricUnit="m" metricValue="2.23" pageId="6" pageNumber="151" unit="m" value="223.0">223 m</elevation>
</quantity>
elev.,
<date box="[585,719,1846,1873]" pageId="6" pageNumber="151" value="2019-07-06">
<collectingDate box="[585,719,1846,1873]" pageId="6" pageNumber="151" value="2019-07-06">6 July 2019</collectingDate>
</date>
,
<emphasis box="[730,1011,1846,1873]" italics="true" pageId="6" pageNumber="151">
<collectorName box="[730,869,1847,1873]" pageId="6" pageNumber="151">P. Souladeth</collectorName>
et al. L3067
</emphasis>
(
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,
<collectionCode box="[1090,1153,1847,1873]" country="Japan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15116" lsid="urn:lsid:biocol.org:col:15116" name="Kagoshima University Museum - Herbarium" pageId="6" pageNumber="151" type="Herbarium">KAG</collectionCode>
,
<collectionCode box="[1164,1221,1847,1873]" country="Japan" lsid="urn:lsid:biocol.org:col:15622" name="National Museum of Nature and Science" pageId="6" pageNumber="151" type="Herbarium">TNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[189,786,1882,1909]" pageId="6" pageNumber="151" type="distribution">
<paragraph blockId="6.[136,1451,1810,2017]" box="[189,786,1882,1909]" pageId="6" pageNumber="151">
<emphasis bold="true" box="[189,373,1882,1908]" pageId="6" pageNumber="151">Distribution:—</emphasis>
<collectingCountry box="[373,491,1883,1909]" name="Cambodia" pageId="6" pageNumber="151">Cambodia</collectingCountry>
,
<collectingCountry box="[502,559,1883,1909]" name="Laos" pageId="6" pageNumber="151">Laos</collectingCountry>
,
<collectingCountry box="[570,672,1883,1909]" name="Thailand" pageId="6" pageNumber="151">Thailand</collectingCountry>
,
<collectingCountry box="[683,782,1883,1909]" name="Vietnam" pageId="6" pageNumber="151">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="151" type="vernacular_names">
<paragraph blockId="6.[136,1451,1810,2017]" pageId="6" pageNumber="151">
<emphasis bold="true" box="[189,437,1919,1945]" pageId="6" pageNumber="151">Vernacular name:—</emphasis>
[(Xangsan Speu, “Xangsan” means
<taxonomicName box="[1062,1185,1919,1945]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="151" phylum="Tracheophyta" rank="genus">
<emphasis box="[1062,1185,1919,1945]" italics="true" pageId="6" pageNumber="151">Lasianthus</emphasis>
</taxonomicName>
in general in Lao and “Speu” refers to “Knang Krepoeu”, the
<typeStatus box="[601,649,1955,1981]" pageId="6" pageNumber="151">type</typeStatus>
locality of this species, which is currently known as
<collectingRegion country="Cambodia" name="Kampong Speu" pageId="6" pageNumber="151">Kampong Speu Province</collectingRegion>
), proposed here].
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="152" type="discussion">
<paragraph blockId="7.[136,1451,158,257]" pageId="7" pageNumber="152">
<emphasis bold="true" box="[189,294,158,184]" pageId="7" pageNumber="152">Notes:—</emphasis>
<taxonomicName authorityName="Pierre ex Pitard" authorityYear="1924" box="[294,559,158,184]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="152" phylum="Tracheophyta" rank="species" species="repoeuensis">
<emphasis box="[294,559,158,184]" italics="true" pageId="7" pageNumber="152">Lasianthus repoeuensis</emphasis>
</taxonomicName>
is distinguished from the other
<taxonomicName box="[932,1146,158,184]" pageId="7" pageNumber="152">
<emphasis box="[932,1055,158,184]" italics="true" pageId="7" pageNumber="152">Lasianthus</emphasis>
species
</taxonomicName>
in
<collectingCountry box="[1187,1241,159,185]" name="Laos" pageId="7" pageNumber="152">Laos</collectingCountry>
by having lamina oblong-elliptic, elliptic-obovate, 917 ×
<quantity box="[587,714,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" metricValueMax="4.3" metricValueMin="2.8" pageId="7" pageNumber="152" unit="cm" value="3.55" valueMax="4.3" valueMin="2.8">2.84.3 cm</quantity>
, glabrous, adaxially blackish brown when dry, with 1416 pairs of secondary veins, and stipule broadly triangular,
<quantity box="[673,766,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="152" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long.
</paragraph>
</subSubSection>
</treatment>
</document>