treatments-xml/data/6D/31/8C/6D318C27FF995C6AFF1C38D4AEECFB02.xml
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<document id="FCDB3455A63579F1BF952E54B3DE491F" ID-CLB-Dataset="3907" ID-DOI="10.11646/zootaxa.5057.1.4" ID-GBIF-Dataset="4becfc46-f283-4a33-9f7a-2b4d041a7118" ID-ISSN="1175-5326" ID-Zenodo-Dep="5585534" ID-ZooBank="E20C7DF7-5EFC-40E1-BBE2-CFDAA05672AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634713448717" checkinUser="plazi" docAuthor="Nabozhenko, Maxim V., Keskin, Bekir, Keskin, Nurşen Alpagut, Gagarina, Ludmila V. &amp; Nabozhenko, Svetlana" docDate="2021" docId="6D318C27FF995C6AFF1C38D4AEECFB02" docLanguage="en" docName="zootaxa.5057.1.4.pdf" docOrigin="Zootaxa 5057 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Turkonalassus mavi M. Nabozhenko &amp; B. Keskin 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="73" masterDocId="9108F45FFF9B5C6EFF8B380BAD63FFF5" masterDocTitle="Two new species and new records of lichen-feeding darkling beetles (Coleoptera: Tenebrionidae: Helopini) from Turkey with notes on bionomics and trophic relations" masterLastPageNumber="86" masterPageNumber="69" pageNumber="71" updateTime="1699460421271" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="25D866AD0D9AFAB5B2B20E512A382028">Two new species and new records of lichen-feeding darkling beetles (Coleoptera: Tenebrionidae: Helopini) from Turkey with notes on bionomics and trophic relations</mods:title>
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<mods:namePart id="21D4B253C2BB49617C07962454A56BD6">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="272E8BD70C8F0FEBFF0B1F3D902C1B15">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. &amp; Dagestan State University, M. Gadzhiev str. 43 a, Makhachkala, Republic of Dagestan, 367000, Russia.</mods:affiliation>
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<mods:namePart id="EEB7B29899265D8E99124901B341236D">Keskin, Bekir</mods:namePart>
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<mods:affiliation id="F39DBEB0C642F34546D93B10A806EDB2">Ege University, Bornova-Izmir 35100 Turkey. &amp; bekir. keskin @ ege. edu. tr; https: // orcid. org / 0000 - 0002 - 6662 - 8736</mods:affiliation>
<mods:nameIdentifier id="1E5178D59E199E8C0FC1FFB15D16E26E" type="email">bekir.keskin@ege.edu.tr</mods:nameIdentifier>
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<mods:namePart id="EEA41CC99089DC7F67F947116B758792">Keskin, Nurşen Alpagut</mods:namePart>
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<mods:affiliation id="2AD1CA49A60EE23395F76BB598D6DA69">Ege University, Bornova-Izmir 35100 Turkey. &amp; nursen. alpagut @ ege. edu. tr; https: // orcid. org / 0000 - 0001 - 6614 - 8815</mods:affiliation>
<mods:nameIdentifier id="B6882B955A091735D972E90C8CA634D3" type="email">nursen.alpagut@ege.edu.tr</mods:nameIdentifier>
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<mods:namePart id="A7C913A587A7B0BF4AE9879AB9965635">Gagarina, Ludmila V.</mods:namePart>
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<mods:affiliation id="753D8FD36272AAC8F5866104923DF214">Komarov Botanical Institute of the Russian Academy of Sciences, Professor Popov str. 2, St Petersburg, 197376, Russia. gagarinalv @ binran. ru; https: // orcid. org / 0000 - 0003 - 3213 - 1673</mods:affiliation>
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<mods:namePart id="F6A62E16065912A61B05D0CC04717D96">Nabozhenko, Svetlana</mods:namePart>
<mods:affiliation id="7E00B81D52A1E65953AE01898F6A914F">Rostov Branch of the Russian Entomological Society, Rostov-on-Don, Russia.</mods:affiliation>
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<mods:date id="C739614767AE3301BA0F5079F9205096">2021</mods:date>
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<taxonomicName id="229846B2FF995C6CFF1C38D4AF9EFF0C" authority="M. Nabozhenko &amp; B. Keskin" authorityName="M. Nabozhenko &amp; B. Keskin" authorityYear="2021" box="[151,765,223,250]" class="Insecta" family="Tenebrionidae" genus="Turkonalassus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="mavi" status="sp. nov.">
<emphasis id="D7ECE123FF995C6CFF1C38D4ACEFFF0C" bold="true" box="[151,396,223,250]" italics="true" pageId="2" pageNumber="71">Turkonalassus mavi</emphasis>
M. Nabozhenko &amp; B. Keskin
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,
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(
<figureCitation id="7DA321B4FF995C6CFF2B3908AD83FEE8" box="[160,224,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Fig. 1</figureCitation>
)
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<emphasis id="D7ECE123FF995C6CFF1C3940AC65FE90" bold="true" box="[151,262,331,357]" pageId="2" pageNumber="71">Material.</emphasis>
<materialsCitation id="55F0376CFF995C6CFE863940AD94FE5B" ID-GBIF-Occurrence="3391261306" collectingDate="2021-04-18" collectionCode="ZDEU" collectorName="S. &amp; M. Nabozhenko &amp; Nabozhenko, B" country="Turkey" elevation="1300" latitude="37.57013" location="Baskonus Yaylasi" longLatPrecision="1" longitude="36.582996" pageId="2" pageNumber="71" specimenCount="7" specimenCount-female="4" specimenCount-male="2" stateProvince="Kahramanmaras" typeStatus="holotype">
<typeStatus id="3A238393FF995C6CFE863940AC1BFE90" box="[269,376,331,357]" pageId="2" pageNumber="71" type="holotype">Holotype</typeStatus>
,
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(
<collectionCode id="8389A5F4FF995C6CFE2E3940AC90FE90" box="[421,499,331,357]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="2" pageNumber="71">ZDEU</collectionCode>
) and
<specimenCount id="F39EF6B8FF995C6CFDA43940AFAFFE93" box="[559,716,331,358]" pageId="2" pageNumber="71" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
(
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and
<specimenCount id="F39EF6B8FF995C6CFCB83940AE35FE90" box="[819,854,331,357]" pageId="2" pageNumber="71" type="female">1♀</specimenCount>
in ZIN, and
<specimenCount id="F39EF6B8FF995C6CFC683940A965FE90" box="[995,1030,331,357]" pageId="2" pageNumber="71" type="female">3♀</specimenCount>
in
<collectionCode id="8389A5F4FF995C6CFBA33947A91AFE93" box="[1064,1145,332,358]" country="Turkey" httpUri="http://grbio.org/cool/s077-eqrn" name="Zoology Department, Ege University" pageId="2" pageNumber="71">ZDEU</collectionCode>
):
<collectingCountry id="9D8F7DA1FF995C6CFB023940A9B9FE90" box="[1161,1242,331,357]" name="Turkey" pageId="2" pageNumber="71">Turkey</collectingCountry>
,
<collectingRegion id="275CF3D3FF995C6CFB6E3940ADB6FE7F" country="Turkey" name="Kahramanmaras" pageId="2" pageNumber="71">Kahramanmaraş Prov.</collectingRegion>
,
<location id="E0476BEAFF995C6CFF6F3964ACD1FE7C" LSID="urn:lsid:plazi:treatment:6D318C27FF995C6AFF1C38D4AEECFB02:E0476BEAFF995C6CFF6F3964ACD1FE7C" box="[228,434,367,393]" country="Turkey" latitude="37.57013" longLatPrecision="1" longitude="36.582996" name="Baskonus Yaylasi" pageId="2" pageNumber="71" stateProvince="Kahramanmaras">Başkonuş Yaylası</location>
,
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/
<geoCoordinate id="80AC5BF6FF995C6CFD0C397AAE4DFE79" box="[647,814,369,396]" degrees="36" direction="east" minutes="34" orientation="longitude" pageId="2" pageNumber="71" precision="1" seconds="58.79" value="36.582996">36°34ʹ58.79ʺE</geoCoordinate>
,
<quantity id="226090D4FF995C6CFCB53964AEF9FE7F" box="[830,922,367,394]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="71" unit="m" value="1300.0">
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,
<date id="91261BF1FF995C6CFC213964A94BFE7C" box="[938,1064,367,393]" pageId="2" pageNumber="71" value="2021-04-18">
<collectingDate id="8162E219FF995C6CFC213964A94BFE7C" box="[938,1064,367,393]" pageId="2" pageNumber="71" value="2021-04-18">18.04.2021</collectingDate>
</date>
(leg.
<collectorName id="486D58E7FF995C6CFBF83964A811FE7C" box="[1139,1394,367,394]" pageId="2" pageNumber="71">S. &amp; M. Nabozhenko</collectorName>
, B. Keskin).
</materialsCitation>
</paragraph>
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<emphasis id="D7ECE123FF995C6CFF4C39BCAC3AFE24" bold="true" box="[199,345,439,465]" pageId="2" pageNumber="71">Description.</emphasis>
Male. Body robust, weakly shine dorsally; elytra dark-bluish, pronotum black, with bluish and greenish tint; head black, with very slight greenish tint; scutellar shield weakly bronze; legs, antennae, mouthparts, and ventral side of body brown (
<figureCitation id="7DA321B4FF995C6CFD8F39F4AFEBFDEF" box="[516,648,511,538]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Figs. 1A, B</figureCitation>
).
</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3A2FA997FDC8" blockId="2.[151,1437,331,2014]" box="[199,1268,547,580]" pageId="2" pageNumber="71">
Measurements: Y—1.57;
<collectingCountry id="9D8F7DA1FF995C6CFE633A28AF6DFDC8" box="[488,526,547,573]" name="Philippines" pageId="2" pageNumber="71">PH</collectingCountry>
—1.66; P
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P
<subScript id="791C3F74FF995C6CFD183A3EAFFBFDB6" attach="left" box="[659,664,565,579]" fontSize="6" pageId="2" pageNumber="71">l</subScript>
—1.25; E
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E
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—1.4; EH
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—2.1; EP
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—1.28; EP
<subScript id="791C3F74FF995C6CFB103A3EA9C3FDB6" attach="left" box="[1179,1184,565,579]" fontSize="6" pageId="2" pageNumber="71">l</subScript>
—2.26.
</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3A4CADA1FCAB" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">Head. Widest at eye level. Eyes moderately large, strongly convex. Lateral margin of genae weakly regularly rounded. Lateral margin of head with weak and short sinuation between gena and epistoma. Anterior margin of epistoma straight. Punctation of head coarse and dense, double (small punctures among large foveolate round punctation). Ventral side of head with transverse coarse wrinkles before gula. Antennae comparatively long, with 3 antennomeres extending beyond base of pronotum. Middle antennomeres not strongly thickened, antennae regularly widened from proximal to distal antennomeres, but ultimate antennomere 11 narrower than penultimate one. Ratio of length: width of antennomeres 211: 2: 1.1, 5.3: 2, 4.4: 2, 4: 2, 4: 2, 4: 2.2, 4.5: 2.4, 4: 2.7, 3.8: 2.6, 4: 2.5.</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3B6CAF4EFBAC" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">Prothorax. Pronotum transverse, widest before middle. Lateral margins of pronotum weakly rounded, but slightly sinuate in basal quarter; anterior margin rounded at middle and sinuated near lateral sides; base weakly bisinuate. Anterior corners widely rounded, weakly projected, posterior ones straight, pointed. All margins (except for middle of anterior one) narrowly beaded; base with wider bead at middle. Disc slightly widely flattened on lateral sides, punctation of disc the same as on head. Prothoracic hypomera with flattened lateral margins, coarsely longitudinally wrinkled. Prosternum with coarse and dense punctation, raduliform in middle. Prosternal process weakly convex, not beaded apically.</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3C68AECDFA6B" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">Pterothorax. Scutellar shield triangle, with slightly rounded lateral margins, finely and densely punctated, but with smooth midline. Elytra widest at middle. Lateral carina (dorsal surface of epipleura) and humeral angles very wide, completely visible dorsally. Punctures in striae elongate merged in interrupted deep furrows. Interstriae flat, with simple and weak transverse wrinkles coming from strial punctures, but not coriaceous, without dense irregular micro-wrinkles among punctation. Epipleura not reaching, and epipleural inner carina reaching elytra apex. Punctation of interstriae fine and sparse. Suture finely beaded at apical third of elytra. Eighth interstria convex, connected with lateral elytral margin, dorsal epipleural carina reaching only eighth interstria. Mesoventrite and its sclerites with coarse and dense rugose punctation; metepisterna with dense and coarse simple punctation; metaventrite with fine and sparse punctation and fine recumbent setae.</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3DA3AC2CF9FC" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">Legs. Trochanters with one long yellow seta. Femora with sparse recumbent setation; pro- and metatibiae straight, mesotibiae weakly bent; tibiae with denser setation of yellow setae. Tarsi long. Mesotarsi slightly shorter than mesotibiae.</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3E18AED7F948" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">
Abdomen. Abdominal ventrites 1 and 2 with simple fine and sparse punctation; ventrites 35 with the same, but double punctation (very small punctures among small ones); ventrite 5 beaded only laterally. Genitalia (
<figureCitation id="7DA321B4FF995C6CFAEF3E3CADB8F980" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Figs. 1DF</figureCitation>
). Aedeagus nalassoid (
<bibRefCitation id="810940C0FF995C6CFE673E50AFA3F983" author="Nabozhenko, M. V." box="[492,704,1627,1654]" pageId="2" pageNumber="71" pagination="37 - 48" refId="ref9513" refString="Nabozhenko, M. V. (2005) Interstructural correlations in evolution of darkling beetles of the tribe Helopini (Coleoptera: Tenebrionidae). Caucasian Entomological Bulletin, 1 (1), 37 - 48. [in Russian] https: // doi. org / 10.23885 / 1814 - 3326 - 2005 - 1 - 1 - 37 - 48" type="journal article" year="2005">Nabozhenko 2005</bibRefCitation>
); baculi of median lobe wide, connected (
<figureCitation id="7DA321B4FF995C6CFB133E50A992F980" box="[1176,1265,1627,1653]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Fig. 1D</figureCitation>
). Inner sternite VIII with rounded blades and dense setation of long setae. Spiculum gastrale with thin rods and long common stem; blades (derivates of tergite IX) triangle, with sclerotized outer margins.
</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3ECCAFECF917" blockId="2.[151,1437,331,2014]" box="[199,655,1735,1762]" pageId="2" pageNumber="71">
Body length
<quantity id="226090D4FF995C6CFED33ECCACA7F917" box="[344,452,1735,1762]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="2" pageNumber="71" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
, width
<quantity id="226090D4FF995C6CFD9D3ECCAFE8F917" box="[534,651,1735,1762]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="2" pageNumber="71" unit="mm" value="4.15" valueMax="4.3" valueMin="4.0">44.3 mm</quantity>
.
</paragraph>
<paragraph id="E5273D31FF995C6CFF4C3EE0A942F8BB" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">
Female (
<figureCitation id="7DA321B4FF995C6CFEA53EE0ACE9F8F0" box="[302,394,1771,1797]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Fig. 1B</figureCitation>
). Measurements:
<collectingCountry id="9D8F7DA1FF995C6CFDD73EE0AFE1F8F0" box="[604,642,1771,1797]" name="Philippines" pageId="2" pageNumber="71">PH</collectingCountry>
—1.67; P
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P
<subScript id="791C3F74FF995C6CFC803EF6AE73F8FE" attach="left" box="[779,784,1789,1803]" fontSize="6" pageId="2" pageNumber="71">l</subScript>
—1.15; E
<subScript id="791C3F74FF995C6CFC093EF6AEE4F8FE" attach="right" box="[898,903,1789,1803]" fontSize="6" pageId="2" pageNumber="71">l</subScript>
E
<subScript id="791C3F74FF995C6CFC1C3EF5AEC0F8F9" attach="left" box="[919,931,1790,1804]" fontSize="6" pageId="2" pageNumber="71">w</subScript>
—1.47; EH
<subScript id="791C3F74FF995C6CFBA23EF5A956F8F9" attach="left" box="[1065,1077,1790,1804]" fontSize="6" pageId="2" pageNumber="71">w</subScript>
—2.23; EP
<subScript id="791C3F74FF995C6CFB3D3EF5A9A1F8F9" attach="left" box="[1206,1218,1790,1804]" fontSize="6" pageId="2" pageNumber="71">w</subScript>
—1.33; EP
<subScript id="791C3F74FF995C6CFACF3EF6A82AF8FE" attach="left" box="[1348,1353,1789,1803]" fontSize="6" pageId="2" pageNumber="71">l</subScript>
—2.26. Body robuster, more bluish, pronotum with greenish-violet tint, disc of pronotum more flattened, elytra longer, protarsi slightly narrower than in male. Body length
<quantity id="226090D4FF995C6CFD6F3F38AE11F8BB" box="[740,882,1843,1870]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.1199999999999999" metricValueMin="1.1" pageId="2" pageNumber="71" unit="mm" value="11.1" valueMax="11.2" valueMin="11.0">1111.2 mm</quantity>
, width
<quantity id="226090D4FF995C6CFC4F3F38A97EF8BB" box="[964,1053,1843,1870]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="71" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="AD826EBAFF995C6CFF4C3F5CAC0EF860" pageId="2" pageNumber="71" type="etymology">
<paragraph id="E5273D31FF995C6CFF4C3F5CAC0EF860" blockId="2.[151,1437,331,2014]" pageId="2" pageNumber="71">
<emphasis id="D7ECE123FF995C6CFF4C3F5CAC2DF884" bold="true" box="[199,334,1879,1905]" pageId="2" pageNumber="71">Etymology.</emphasis>
The name derives from Turkish “mavi” (blue) and emphasizes the bluish colouration of the dorsum of the new species.
</paragraph>
</subSubSection>
<subSubSection id="AD826EBAFF995C6AFF4C3F94AEECFB02" lastPageId="4" lastPageNumber="73" pageId="2" pageNumber="71" type="diagnosis">
<paragraph id="E5273D31FF995C6AFF4C3F94AE3EFBB6" blockId="2.[151,1437,331,2014]" lastBlockId="4.[151,1437,848,1271]" lastPageId="4" lastPageNumber="73" pageId="2" pageNumber="71">
<emphasis id="D7ECE123FF995C6CFF4C3F94ACB3F84C" bold="true" box="[199,464,1951,1977]" pageId="2" pageNumber="71">Differential diagnosis.</emphasis>
This new species differs from all congeners by the bluish dorsal surface of the body (
<figureCitation id="7DA321B4FF995C6CFF143FC8AC7FF828" box="[159,284,1987,2013]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Figs. 1A,B</figureCitation>
), the pronotum widest at anterior half (
<figureCitation id="7DA321B4FF995C6CFD443FC8AE2AF828" box="[719,841,1987,2013]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="2" pageNumber="71">Figs. 1A,B</figureCitation>
) (at middle or behind the middle in other congeners), and wide merged baculi of the median lobe of the aedeagus (
<figureCitation id="7DA321B4FF9F5C6AFCB63B5BAEF1FC9E" box="[829,914,848,875]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[160,1429,185,1870]" captionTargetId="figure-17@3.[151,1436,179,1876]" captionTargetPageId="3" captionText="FIGURE 1. Turkonalassus mavi sp. n., habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="4" pageNumber="73">Fig. 1E</figureCitation>
). It is similar to
<emphasis id="D7ECE123FF9F5C6AFBC13B5AA9A1FC9F" box="[1098,1218,848,874]" italics="true" pageId="4" pageNumber="73">T. pentheri</emphasis>
(Reitter, 1905) and
<emphasis id="D7ECE123FF9F5C6AFF1C3B7EAC57FC7B" box="[151,308,884,910]" italics="true" pageId="4" pageNumber="73">T. petrophilus</emphasis>
<bibRefCitation id="810940C0FF9F5C6AFEB63B7FAE2CFC7A" author="Keskin, B. &amp; Nabozhenko, M. &amp; Alpagut Keskin, N." box="[317,847,884,911]" pageId="4" pageNumber="73" pagination="237 - 240" refId="ref9182" refString="Keskin, B., Nabozhenko, M. &amp; Alpagut Keskin, N. (2017 a) Eustenomacidius egeuniversitatis sp. n. - the first record of the tenebrionid genus in Turkey. Turkish Journal of Zoology, 41 (2), 237 - 240. https: // doi. org / 10.3906 / zoo- 1509 - 1" type="journal article" year="2017">Keskin, Nabozhenko &amp; Alpagut Keskin, 2017</bibRefCitation>
, which also have the punctation of elytral intervals fine and sparse, with the puncture diameter 35 times as less as interpuncture distance.
<taxonomicName id="229846B2FF9F5C6AFBFE3B93A837FC47" authority="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko, 2021" authorityName="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko" authorityYear="2021" box="[1141,1364,920,946]" class="Insecta" family="Tenebrionidae" genus="Turkonalassus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="mavi" status="sp. nov.">
<emphasis id="D7ECE123FF9F5C6AFBFE3B93A837FC47" box="[1141,1364,920,946]" italics="true" pageId="4" pageNumber="73">Turkonalassus mavi</emphasis>
</taxonomicName>
<emphasis id="D7ECE123FF9F5C6AFAD73B92A8FEFC46" bold="true" box="[1372,1437,921,947]" pageId="4" pageNumber="73">
<taxonomicNameLabel id="CCDF5C58FF9F5C6AFAD73B92A8FEFC46" box="[1372,1437,921,947]" pageId="4" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
additionally differs from the first species by not thickened male antennomeres 48 (which are thicker than three apical antennomeres in
<emphasis id="D7ECE123FF9F5C6AFE173BEAAF77FC0F" box="[412,532,992,1018]" italics="true" pageId="4" pageNumber="73">T. pentheri</emphasis>
) and strial punctures merged in furrows (separate in
<emphasis id="D7ECE123FF9F5C6AFBD23BEAA9B3FC0F" box="[1113,1232,992,1018]" italics="true" pageId="4" pageNumber="73">T. pentheri</emphasis>
). The new species additionally differs from the latter taxon by much denser and coarser punctation of round punctures (punctures smaller, sparse and elongated on lateral sides in
<emphasis id="D7ECE123FF9F5C6AFD393C22AE2EFBB7" box="[690,845,1064,1090]" italics="true" pageId="4" pageNumber="73">T. petrophilus</emphasis>
).
</paragraph>
<caption id="B1E76DB9FF985C6DFF1C3F65AC91F83B" ID-DOI="http://doi.org/10.5281/zenodo.5585536" ID-Zenodo-Dep="5585536" httpUri="https://zenodo.org/record/5585536/files/figure.png" pageId="3" pageNumber="72" startId="3.[151,250,1902,1926]" targetBox="[160,1429,185,1870]" targetPageId="3">
<paragraph id="E5273D31FF985C6DFF1C3F65AC91F83B" blockId="3.[151,1436,1902,1998]" pageId="3" pageNumber="72">
<emphasis id="D7ECE123FF985C6DFF1C3F65AC7BF873" bold="true" box="[151,280,1902,1926]" pageId="3" pageNumber="72">FIGURE 1.</emphasis>
<taxonomicName id="229846B2FF985C6DFEA93F65AC92F873" authority="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko, 2021" authorityName="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko" authorityYear="2021" box="[290,497,1902,1926]" class="Insecta" family="Tenebrionidae" genus="Turkonalassus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="species" species="mavi" status="sp. nov.">
<emphasis id="D7ECE123FF985C6DFEA93F65AC92F873" box="[290,497,1902,1926]" italics="true" pageId="3" pageNumber="72">Turkonalassus mavi</emphasis>
</taxonomicName>
<emphasis id="D7ECE123FF985C6DFE773F65AF5AF873" bold="true" box="[508,569,1902,1926]" pageId="3" pageNumber="72">
<taxonomicNameLabel id="CCDF5C58FF985C6DFE773F65AF5AF873" box="[508,569,1902,1926]" pageId="3" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, habitus and details of structure: A—habitus dorsally, male; B—habitus dorsally, female; C—habitus ventrally, male; D—male inner sternite VIII; E—aedeagus (bml—baculi of median lobe, apa—apical piece apophises); F—spiculum gastrale.
</paragraph>
</caption>
<caption id="B1E76DB9FF9F5C6AFF1C3AE6AFDAFCDC" ID-DOI="http://doi.org/10.5281/zenodo.5585538" ID-Zenodo-Dep="5585538" httpUri="https://zenodo.org/record/5585538/files/figure.png" pageId="4" pageNumber="73" startId="4.[151,250,749,773]" targetBox="[151,1436,181,725]" targetPageId="4">
<paragraph id="E5273D31FF9F5C6AFF1C3AE6AFDAFCDC" blockId="4.[151,1436,749,809]" pageId="4" pageNumber="73">
<emphasis id="D7ECE123FF9F5C6AFF1C3AE6AC77FCF0" bold="true" box="[151,276,749,773]" pageId="4" pageNumber="73">FIGURE 2.</emphasis>
<taxonomicName id="229846B2FF9F5C6AFE903AE6AF41FCF0" box="[283,546,749,773]" class="Lecanoromycetes" family="Parmeliaceae" genus="Pseudevernia" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="73" phylum="Ascomycota" rank="species" species="furfuraceae">
<emphasis id="D7ECE123FF9F5C6AFE903AE6AF41FCF0" box="[283,546,749,773]" italics="true" pageId="4" pageNumber="73">Pseudevernia furfuraceae</emphasis>
</taxonomicName>
, host lichen (on
<taxonomicName id="229846B2FF9F5C6AFD5B3AE5AE24FCF0" box="[720,839,750,773]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="D7ECE123FF9F5C6AFD5B3AE5AE24FCF0" box="[720,839,750,773]" italics="true" pageId="4" pageNumber="73">Pinus nigra</emphasis>
</taxonomicName>
) of
<taxonomicName id="229846B2FF9F5C6AFCF83AE6A95DFCF0" authority="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko, 2021" authorityName="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko" authorityYear="2021" box="[883,1086,749,773]" class="Insecta" family="Tenebrionidae" genus="Turkonalassus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="mavi" status="sp. nov.">
<emphasis id="D7ECE123FF9F5C6AFCF83AE6A95DFCF0" box="[883,1086,749,773]" italics="true" pageId="4" pageNumber="73">Turkonalassus mavi</emphasis>
</taxonomicName>
<emphasis id="D7ECE123FF9F5C6AFBCE3AE6A91CFCF0" bold="true" box="[1093,1151,749,773]" pageId="4" pageNumber="73">
<taxonomicNameLabel id="CCDF5C58FF9F5C6AFBCE3AE6A91CFCF0" box="[1093,1151,749,773]" pageId="4" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: A—general view of tallus; B—damage of isidia from beetles (shown by arrows).
</paragraph>
</caption>
<paragraph id="E5273D31FF9F5C6AFF4C3C47AEECFB02" blockId="4.[151,1437,848,1271]" pageId="4" pageNumber="73">
<emphasis id="D7ECE123FF9F5C6AFF4C3C47AF2DFB93" bold="true" box="[199,590,1100,1127]" pageId="4" pageNumber="73">Bionomics and trophic relations.</emphasis>
The species was collected in old
<taxonomicName id="229846B2FF9F5C6AFC473C46A9B7FB93" authority="J. F. Arnold" authorityName="J. F. Arnold" box="[972,1236,1100,1127]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="D7ECE123FF9F5C6AFC473C46A932FB93" box="[972,1105,1101,1126]" italics="true" pageId="4" pageNumber="73">Pinus nigra</emphasis>
J.F. Arnold
</taxonomicName>
forest with shrub undergrowth of
<taxonomicName id="229846B2FF9F5C6AFEDA3C7AAFF0FB7F" authority="L. The" authorityName="L. The" box="[337,659,1136,1163]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="D7ECE123FF9F5C6AFEDA3C7AAF5FFB7F" box="[337,572,1137,1162]" italics="true" pageId="4" pageNumber="73">Juniperus communis</emphasis>
L. The
</taxonomicName>
presence of this juniper shrub undergrowth is crucial factor for all species of
<taxonomicName id="229846B2FF9F5C6AFE843C9FACD2FB5B" authorityName="Keskin, Nabozhenko &amp; Alpagut-Keskin" authorityYear="2017" box="[271,433,1172,1198]" class="Insecta" family="Tenebrionidae" genus="Turkonalassus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="D7ECE123FF9F5C6AFE843C9FACD2FB5B" box="[271,433,1172,1198]" italics="true" pageId="4" pageNumber="73">Turkonalassus</emphasis>
</taxonomicName>
. Usually, beetles hide in thickets of
<taxonomicName id="229846B2FF9F5C6AFCC53C9EAEBFFB5B" box="[846,988,1173,1198]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="communis">
<emphasis id="D7ECE123FF9F5C6AFCC53C9EAEBFFB5B" box="[846,988,1173,1198]" italics="true" pageId="4" pageNumber="73">J. communis</emphasis>
</taxonomicName>
, and at night they go out on tree trunks or stones. Adult beetles of
<taxonomicName id="229846B2FF9F5C6AFE4C3CB2AF79FB27" authority="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko, 2021" authorityName="Nabozhenko &amp; Keskin &amp; Keskin &amp; Gagarina &amp; Nabozhenko" authorityYear="2021" box="[455,538,1209,1234]" class="Insecta" family="Tenebrionidae" genus="Turkonalassus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="mavi" status="sp. nov.">
<emphasis id="D7ECE123FF9F5C6AFE4C3CB2AF79FB27" box="[455,538,1209,1234]" italics="true" pageId="4" pageNumber="73">T. mavi</emphasis>
</taxonomicName>
<emphasis id="D7ECE123FF9F5C6AFDA93CB2AF00FB26" bold="true" box="[546,611,1209,1235]" pageId="4" pageNumber="73">
<taxonomicNameLabel id="CCDF5C58FF9F5C6AFDA93CB2AF00FB26" box="[546,611,1209,1235]" pageId="4" pageNumber="73" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were active at night since 20:30 on trunks of
<taxonomicName id="229846B2FF9F5C6AFBE63CB2A992FB27" box="[1133,1265,1209,1234]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="73" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="D7ECE123FF9F5C6AFBE63CB2A992FB27" box="[1133,1265,1209,1234]" italics="true" pageId="4" pageNumber="73">Pinus nigra</emphasis>
</taxonomicName>
. Host lichen is
<taxonomicName id="229846B2FF9F5C6AFF1C3CD7ACD9FB02" box="[151,442,1244,1271]" class="Lecanoromycetes" family="Parmeliaceae" genus="Pseudevernia" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="73" phylum="Ascomycota" rank="species" species="furfuraceae">
<emphasis id="D7ECE123FF9F5C6AFF1C3CD7ACD9FB02" box="[151,442,1244,1271]" italics="true" pageId="4" pageNumber="73">Pseudevernia furfuraceae</emphasis>
</taxonomicName>
(L.) Zopf (
<taxonomicName id="229846B2FF9F5C6AFDBC3CD7AE45FB02" authority="Zenker" authorityName="Zenker" authorityYear="1827" box="[567,806,1244,1271]" class="Lecanoromycetes" family="Parmeliaceae" kingdom="Fungi" order="Lecanorales" pageId="4" pageNumber="73" phylum="Ascomycota" rank="family">Parmeliaceae Zenker</taxonomicName>
) (
<figureCitation id="7DA321B4FF9F5C6AFCB63CD7AEE1FB02" box="[829,898,1244,1271]" captionStart="FIGURE 2" captionStartId="4.[151,250,749,773]" captionTargetBox="[151,1436,181,725]" captionTargetId="figure-18@4.[151,1436,181,725]" captionTargetPageId="4" captionText="FIGURE 2. Pseudevernia furfuraceae, host lichen (on Pinus nigra) of Turkonalassus mavi sp. n.: A—general view of tallus; B—damage of isidia from beetles (shown by arrows)." figureDoi="http://doi.org/10.5281/zenodo.5585538" httpUri="https://zenodo.org/record/5585538/files/figure.png" pageId="4" pageNumber="73">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>