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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1053.67326" ID-GBIF-Dataset="6ab5131b-fa67-412c-ad75-6e75f8eac2f8" ID-PMC="PMC8352857" ID-Pensoft-Pub="1313-2970-1053-185" ID-Pensoft-UUID="4A8522D8A609541581BC3FB89AFFFCFE" ID-PubMed="34393559" ID-ZooBank="6C907A4B587440D1881E18BA380D9C21" ModsDocID="1313-2970-1053-185" checkinTime="1627926266140" checkinUser="pensoft" docAuthor="Ng, Peter K. L. &amp; Chan, Tin-Yam" docDate="2021" docId="6539059668145134B5E473D5D99F14CA" docLanguage="en" docName="ZooKeys 1053: 185-195" docOrigin="ZooKeys 1053" docPubDate="2021-08-02" docSource="http://dx.doi.org/10.3897/zookeys.1053.67326" docTitle="Orientotlos Sakai 1980" docType="treatment" docVersion="4" id="4A8522D8A609541581BC3FB89AFFFCFE" lastPageNumber="185" masterDocId="4A8522D8A609541581BC3FB89AFFFCFE" masterDocTitle="Rediscovery of Orientotlos iishibai Sakai, 1980 (Crustacea, Decapoda, Brachyura, Leucosiidae) in Taiwan" masterLastPageNumber="195" masterPageNumber="185" pageNumber="185" updateTime="1668150789084" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rediscovery of Orientotlos iishibai Sakai, 1980 (Crustacea, Decapoda, Brachyura, Leucosiidae) in Taiwan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chan, Tin-Yam</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8143-0007</mods:nameIdentifier>
<mods:affiliation>Institute of Marine Biology and Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung 202301, Taiwan</mods:affiliation>
<mods:nameIdentifier type="email">tychan@mail.ntou.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1053</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>185</mods:start>
<mods:end>195</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1053.67326</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1053.67326</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1053-185</mods:identifier>
<mods:identifier type="ZooBank">6C907A4B587440D1881E18BA380D9C21</mods:identifier>
<mods:identifier type="Pensoft-UUID">4A8522D8A609541581BC3FB89AFFFCFE</mods:identifier>
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<treatment ID-GBIF-Taxon="183907164" LSID="urn:lsid:plazi:treatment:6539059668145134B5E473D5D99F14CA" httpUri="http://treatment.plazi.org/id/6539059668145134B5E473D5D99F14CA" lastPageNumber="185" pageId="0" pageNumber="185">
<subSubSection pageId="0" pageNumber="185" type="nomenclature">
<paragraph pageId="0" pageNumber="185">
Genus
<taxonomicName LSID="65390596-6814-5134-B5E4-73D5D99F14CA" authority="Sakai, 1980" authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">Orientotlos Sakai, 1980</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="type species">
<paragraph pageId="0" pageNumber="185">Type species.</paragraph>
<paragraph pageId="0" pageNumber="185">
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
Sakai, 1980, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="diagnosis">
<paragraph pageId="0" pageNumber="185">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="185">Carapace subhexagonal in outline; dorsal surfaces between plates and bosses with numerous, well-spaced boletiform and rounded tubercles; subhepatic region forming a distinct obtuse angle visible in dorsal view; hepatic plate distinct, separated from first anterolateral tooth by wide cleft; anterolateral margin with 3 large lobiform teeth; posterolateral margin concave, with median triangular tooth; posterior carapace margin with 2 large lozenge-shaped bosses; postfrontal median keel prominent, high, extending posteriorly to cardiac region as raised row of rounded tubercles; postorbital region without deep depression; large boletiform plates on protogastric, epibranchial and metabranchial regions; cardiac region with raised, vaguely T-shaped ridge formed of fused granules; intestinal region inflated, with a large subtriangular boletiform plate; antennule with basal segment occupying lower two-thirds of fossa; basal antennal article large, subquadrate, fused with epistome, forming most of suborbital margin; third maxilliped with merus, ischium and exopod paved with numerous flattened rounded tubercles, basal parts with boletiform tubercles, exopod stout, broad, reaching to about half length of merus; palm of cheliped short, stout, without ridges, lobes or teeth, fingers shorter than palm; ambulatory legs short, merus, carpus and propodus covered with slender and boletiform tubercles along upper and lower margins, dactylo-propodal lock present; anterior thoracic sternites (1-4) strongly compressed, surface of sternite 3 with numerous boletiform tubercles; female thoracic sternite 4 forming keel around distal part of sternopleonal cavity; vulvae relatively small, round, positioned distinctly apart; female pleon ovate, shield-like, covered with rounded tubercles, somites 1 and 2 free, somites 3-6 fused, telson narrowly triangular with distal part linguiform.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="185" type="remarks">
<paragraph pageId="0" pageNumber="185">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="185">
The concept of
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Oreophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oreophorus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Oreophorus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Rüppell">Rueppell</normalizedToken>
, 1830 has changed substantially since 1980, with several revisions clarifying the identities of allied genera (
<taxonomicName class="Malacostraca" family="Leucosiidae" genus="Tlos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Tlos</emphasis>
</taxonomicName>
Adams &amp; White, 1849, and
<taxonomicName authorityName="Ihle" authorityYear="1918" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Oreotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Oreotlos</emphasis>
</taxonomicName>
Ihle, 1918) and the establishment of several new ones:
<taxonomicName authorityName="C.G.S.Tan &amp; Richer de Forges" authorityYear="1993" class="Malacostraca" family="Leucosiidae" genus="Dolos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dolos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Dolos</emphasis>
</taxonomicName>
Tan &amp; Richer de Forges, 1993,
<taxonomicName class="Malacostraca" family="Leucosiidae" genus="Alox" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alox" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Alox</emphasis>
</taxonomicName>
Tan &amp; Ng, 1996, and
<taxonomicName class="Malacostraca" family="Leucosiidae" genus="Cateios" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cateios" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Cateios</emphasis>
</taxonomicName>
Tan &amp; Ng, 1996.
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos</emphasis>
</taxonomicName>
can nevertheless be easily distinguished from these genera by the anterolateral margin of carapace not expanded posteriorly and sometimes reaching level of the posterior carapace margin, the anterolateral margin is distinctly lobiform or dentiform, dorsal carapace surface without regions distinctly raised to form bosses, with depressed areas never eroded and no obvious postocular depression or groove, cardiac region not large or strongly inflated, and the palm of cheliped is short ovate, with fingers short and relatively slender and cutting edges lined with low teeth (cf.
<bibRefCitation author="Tan, CGS" journalOrPublisher="Raffles Bulletin of Zoology" pageId="0" pageNumber="185" pagination="119 - 132" refId="B23" refString="Tan, CGS, Richer de Forges, B, 1993. On the systematics and ecology of two species of mimetic crabs belonging to the family Leucosiidae (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology 41 (1): 119 - 132" title="On the systematics and ecology of two species of mimetic crabs belonging to the family Leucosiidae (Crustacea: Decapoda: Brachyura)." volume="41" year="1993">Tan and Richer de Forges 1993</bibRefCitation>
;
<bibRefCitation author="Tan, CGS" editor="Richer de Forges, B" journalOrPublisher="2. ORSTOM, 1995" pageId="0" pageNumber="185" pagination="101 - 205" refId="B22" refString="Tan, CGS, Ng, PKL, 1996. A revision of the IndoPacific genus Oreophorus Rueppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: Richer de Forges, B, Ed., Les fonds meubles des lagons de NouvelleCaledonie (Sedimentologie, Benthos), Etudes &amp; Theses Vol. 2. ORSTOM, 1995: 101 - 205" title="A revision of the IndoPacific genus Oreophorus Rueppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae)." volumeTitle="Les fonds meubles des lagons de NouvelleCaledonie (Sedimentologie, Benthos), Etudes &amp; Theses Vol." year="1996">Tan and Ng 1996</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="185">
The carapace of
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos</emphasis>
</taxonomicName>
only superficially resembles that of
<taxonomicName authorityName="Doflein" authorityYear="1904" class="Malacostraca" family="Leucosiidae" genus="Atlantotlos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Atlantotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Atlantotlos</emphasis>
</taxonomicName>
(type and only species
<taxonomicName authorityName="Doflein" authorityYear="1904" class="Malacostraca" family="Leucosiidae" genus="Atlantotlos" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Atlantotlos rhombifer" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="rhombifer">
<emphasis italics="true" pageId="0" pageNumber="185">Atlantotlos rhombifer</emphasis>
</taxonomicName>
Doflein, 1904), described from off the Congo in West Africa, in general shape (
<bibRefCitation author="Doflein, F" journalOrPublisher="The Crustacea (Vol. 9 C-I): Decapoda: Brachyura (Part 1). Brill, Leiden" pageId="0" pageNumber="185" refId="B5" refString="Doflein, F, 1904. Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Verlag von Gustav Fischer, Jena 6, [1-6 + i-xiv +] 1-314. [Atlas (pls 1-58)]" title="Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Verlag von Gustav Fischer, Jena 6, [1 - 6 + i-xiv +] 1 - 314. [Atlas (pls 1 - 58)]" year="1904">Doflein 1904</bibRefCitation>
: 51). The latter genus differs markedly from
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos</emphasis>
</taxonomicName>
in possessing a smooth carapace, without inflated bosses, boletiform, or rounded tubercles, an entire posterior carapace margin, and less prominently armed pereopods (cf.
<bibRefCitation author="Doflein, F" journalOrPublisher="The Crustacea (Vol. 9 C-I): Decapoda: Brachyura (Part 1). Brill, Leiden" pageId="0" pageNumber="185" refId="B5" refString="Doflein, F, 1904. Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot;Valdivia&quot; 1898-1899. Verlag von Gustav Fischer, Jena 6, [1-6 + i-xiv +] 1-314. [Atlas (pls 1-58)]" title="Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Verlag von Gustav Fischer, Jena 6, [1 - 6 + i-xiv +] 1 - 314. [Atlas (pls 1 - 58)]" year="1904">Doflein 1904</bibRefCitation>
: pl. 15, figs 7, 8).
</paragraph>
<paragraph pageId="0" pageNumber="185">
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos</emphasis>
</taxonomicName>
is actually most similar to
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
A. Milne-Edwards, 1873 in having the anterolateral margin of the carapace with three large lobiform teeth (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
) (versus margin with low lobes or spines but never clearly dentiform); a prominent lobe on posterolateral margin of carapace (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
) (versus lobe absent or undiscernible in
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
, although margin may be uneven or spiniform); median keel on gastric, cardiac and intestinal regions high and strongly inflated (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
) (versus much lower in
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
); intestinal region formed by a pair of distinct, fused subtriangular bosses (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, C, D</figureCitation>
) (versus more coniform or evenly rounded in
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
); posterior carapace margin with two broad truncate, lozenge-shaped bosses (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2C, D</figureCitation>
) (versus entire or with dentiform projections in
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
); female cheliped palm and fingers proportionately shorter and stouter (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientotlos iishibai Sakai, 1980 A holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after Sakai 1980: text-fig. 1] B color in life, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan." figureDoi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A overall dorsal view B overall ventral view C, D dorsal view of carapace from different angles." figureDoi="10.3897/zookeys.1053.67326.figure2" httpUri="https://binary.pensoft.net/fig/572660" pageId="0" pageNumber="185">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A frontal view of cephalothorax B frontal view showing antennules, antennae and orbits C lateral view of cephalothorax D pterygostomial region, buccal cavity, third maxillipeds and anterior part of sternopleonal cavity E left third maxilliped F outer view of left chela G right fourth ambulatory leg." figureDoi="10.3897/zookeys.1053.67326.figure3" httpUri="https://binary.pensoft.net/fig/572661" pageId="0" pageNumber="185">3F</figureCitation>
) (versus more elongate and slender in
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
); and vulvae widely spaced (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Orientotlos iishibai Sakai, 1980, female (7.8 x 6.5 mm) (NTOU B 00125), Taiwan A intestinal region, posterior carapace lobes and pleonal somites 2 - 6 B posterior carapace lobes, pleonal somites 2 - 6, and telson C pleonal somites 4 - 6 and telson D anterior thoracic sternites and sternopleonal cavity E sternopleonal cavity and vulvae." figureDoi="10.3897/zookeys.1053.67326.figure4" httpUri="https://binary.pensoft.net/fig/572662" pageId="0" pageNumber="185">4E</figureCitation>
) (versus proportionately larger and closer to the midline in
<taxonomicName class="Crustacea" family="Leucosiidae" genus="Merocryptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Merocryptus" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="185">Merocryptus</emphasis>
</taxonomicName>
) (cf. A.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10644" author="Milne-Edwards, A" journalOrPublisher="Journal du Museum Godeffroy" pageId="0" pageNumber="185" pagination="77 - 88" refId="B14" refString="Milne-Edwards, A, 1873. Descriptions des quelques crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal du Museum Godeffroy 1: 77 - 88, DOI: https://doi.org/10.5962/bhl.title.10644" title="Descriptions des quelques crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy." url="https://doi.org/10.5962/bhl.title.10644" volume="1" year="1873">Milne-Edwards 1873</bibRefCitation>
;
<bibRefCitation author="Yokoya, Y" journalOrPublisher="Journal of the College of Agriculture, Tokyo Imperial University" pageId="0" pageNumber="185" pagination="1 - 226" refId="B24" refString="Yokoya, Y, 1933. On the distribution of decapod crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by S. S. Soyo-Maru, during the year [sic] 1923-1930. Journal of the College of Agriculture, Tokyo Imperial University 12 (1): 1 - 226" title="On the distribution of decapod crustaceans inhabiting the continental shelf around Japan, chiefly based upon the materials collected by S. S. Soyo-Maru, during the year [sic] 1923 - 1930." volume="12" year="1933">Yokoya 1933</bibRefCitation>
;
<bibRefCitation author="Serene, R" journalOrPublisher="Treubia" pageId="0" pageNumber="185" pagination="137 - 218" refId="B20" refString="Serene, R, 1955. Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2e partie). Treubia 23 (1): 137 - 218" title="Sur quelques especes rares de Brachyures (Leucosidae) de l'Indo-Pacifique (2 e partie)." volume="23" year="1955">
<normalizedToken originalValue="Serène">Serene</normalizedToken>
1955
</bibRefCitation>
;
<bibRefCitation author="Sakai, T" journalOrPublisher=", Japanese text, 461 pp., Plates volume" pageId="0" pageNumber="185" refId="B18" refString="Sakai, T, 1976. Crabs of Japan and the Adjacent Seas. In three volumes: English text, [xxix +] 773 pp., Japanese text, 461 pp., Plates volume" title="Crabs of Japan and the Adjacent Seas. In three volumes: English text, [xxix +] 773 pp." year="1976">Sakai 1976</bibRefCitation>
;
<bibRefCitation author="Zarenkov, NA" editor="Kuznetsov, AP" journalOrPublisher="Shirshov Institute of Oceanology" pageId="0" pageNumber="185" pagination="97 - 125" refId="B25" refString="Zarenkov, NA, 1994. [= Crabs from seamounts of the Western part of the Indian Ocean]. In: Kuznetsov, AP, Mironov, AN, Eds., Bottom Fauna of Seamounts. Transactions of the P. P. Shirshov Institute of Oceanology 129: 97 - 125" title="[= Crabs from seamounts of the Western part of the Indian Ocean]." volume="129" volumeTitle="Bottom Fauna of Seamounts. Transactions of the P. P." year="1994">Zarenkov 1994</bibRefCitation>
;
<bibRefCitation author="Chen, H" journalOrPublisher="Invertebrata 30, Science Press, Beijing" pageId="0" pageNumber="185" refId="B3" refString="Chen, H, Sun, H, 2002. ArthropodaCrustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata 30, Science Press, Beijing" title="ArthropodaCrustacea. Brachyura. Marine primitive crabs. Fauna Sinica." year="2002">Chen and Sun 2002</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.4314/wiojms.v18i2.11" author="Galil, BS" journalOrPublisher="Western Indian Ocean Journal of Marine Science" pageId="0" pageNumber="185" pagination="93 - 97" refId="B6" refString="Galil, BS, 2019. Re-description and new records of a poorly known leucosiid crab, Merocryptus boletisculpta Zarenkov, 1994 (Crustacea, Decapoda, Brachyura). Western Indian Ocean Journal of Marine Science 18 (2): 93 - 97, DOI: https://doi.org/10.4314/wiojms.v18i2.11" title="Re-description and new records of a poorly known leucosiid crab, Merocryptus boletisculpta Zarenkov, 1994 (Crustacea, Decapoda, Brachyura)." url="https://doi.org/10.4314/wiojms.v18i2.11" volume="18" year="2019">Galil 2019</bibRefCitation>
;
<bibRefCitation author="Galil, BS" journalOrPublisher="Tropical Deep-Sea Benthos, Paris" pageId="0" pageNumber="185" refId="B9" refString="Galil, BS, Ng, PKL, in press. New and rare leucosiid crabs (Crustacea: Decapoda: Brachyura) from Madagascar. In: Richer de Forges B, Corbari L (Eds) Deep-Sea Crustaceans from Madagascar. Tropical Deep-Sea Benthos, in press." title="New and rare leucosiid crabs (Crustacea: Decapoda: Brachyura) from Madagascar. In: Richer de Forges B, Corbari L (Eds) Deep-Sea Crustaceans from Madagascar. Tropical Deep-Sea Benthos, in press." year="in press">Galil and Ng in press</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1053.67326.figure1" httpUri="https://binary.pensoft.net/fig/572659" pageId="0" pageNumber="185" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="185">
<emphasis bold="true" pageId="0" pageNumber="185">Figure 1.</emphasis>
<taxonomicName authorityName="Sakai" authorityYear="1980" class="Malacostraca" family="Leucosiidae" genus="Orientotlos" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orientotlos iishibai" order="Decapoda" pageId="0" pageNumber="185" phylum="Arthropoda" rank="species" species="iishibai">
<emphasis italics="true" pageId="0" pageNumber="185">Orientotlos iishibai</emphasis>
</taxonomicName>
Sakai, 1980
<emphasis bold="true" pageId="0" pageNumber="185">A</emphasis>
holotype female (7.0 by 5.5 mm) (repository unknown), Japan [after
<bibRefCitation DOI="https://doi.org/10.18353/rcustacea.10.0_73" author="Sakai, T" journalOrPublisher="Researches on Crustacea" pageId="0" pageNumber="185" pagination="73 - 84" refId="B19" refString="Sakai, T, 1980. On new and rare crabs taken from Japanese and Central Pacific waters. Researches on Crustacea 10: 73 - 84, DOI: https://doi.org/10.18353/rcustacea.10.0_73" title="On new and rare crabs taken from Japanese and Central Pacific waters." url="https://doi.org/10.18353/rcustacea.10.0_73" volume="10" year="1980">Sakai 1980</bibRefCitation>
: text-fig. 1]
<emphasis bold="true" pageId="0" pageNumber="185">B</emphasis>
color in life, female (7.8
<normalizedToken originalValue="×">x</normalizedToken>
6.5 mm) (NTOU B00125), Taiwan.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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