treatments-xml/data/61/70/87/61708795E7081017BEB704A2FC26F910.xml

<document id="AC756A5723BB32BE72385DF2A639C988" ID-DOI="10.11646/zootaxa.3637.1.7" ID-GBIF-Dataset="16a61a70-3efa-4954-ace3-3a9cbf7213d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="283602" ID-ZooBank="0872BF7E-6BD2-465C-869D-A68CB87909D6" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460590683478" checkinUser="plazi" docAuthor="Belousov, Igor A. &amp; Nyundo, Bruno A." docDate="2013" docId="61708795E7081017BEB704A2FC26F910" docLanguage="en" docName="zt03637p073.pdf" docOrigin="Zootaxa 3637 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachytrechodes brevis Belousov &amp; Nyundo, sp. n." docType="treatment" docVersion="6" lastPageNumber="72" masterDocId="9D49FFEDE7091010BE200616FF90FFFE" masterDocTitle="A new species of Pachytrechodes Jeannel, 1960 (Coleoptera: Carabidae: Trechinae) from Tanzania, with a key to species" masterLastPageNumber="73" masterPageNumber="65" pageNumber="66" updateTime="1698325865938" updateUser="plazi">
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<mods:title id="88F506E6E1984A88A6DE1CD0F4DE6D5D">A new species of Pachytrechodes Jeannel, 1960 (Coleoptera: Carabidae: Trechinae) from Tanzania, with a key to species</mods:title>
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<mods:namePart id="46AACD4BEF48002961D32DA33B7E8CE8">Belousov, Igor A.</mods:namePart>
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<mods:namePart id="BC2BDEB722ADE6176CB89F36A894E501">Nyundo, Bruno A.</mods:namePart>
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<mods:date id="8192A5FC92955B36723CDF94A7D0D7C4">2013</mods:date>
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<treatment id="61708795E7081017BEB704A2FC26F910" ID-DOI="http://doi.org/10.5281/zenodo.6151245" ID-GBIF-Taxon="127663778" ID-Zenodo-Dep="6151245" LSID="urn:lsid:plazi:treatment:61708795E7081017BEB704A2FC26F910" httpUri="http://treatment.plazi.org/id/61708795E7081017BEB704A2FC26F910" lastPageId="7" lastPageNumber="72" pageId="1" pageNumber="66">
<subSubSection id="A1C36508E7081011BEB704A2FD67FD33" box="[151,759,691,718]" pageId="1" pageNumber="66" type="nomenclature">
<paragraph id="E9663683E7081011BEB704A2FD67FD33" blockId="1.[151,759,691,718]" box="[151,759,691,718]" pageId="1" pageNumber="66">
<heading id="B22E81EFE7081011BEB704A2FD67FD33" bold="true" box="[151,759,691,718]" fontSize="11" level="1" pageId="1" pageNumber="66" reason="1">
<emphasis id="DBADEA91E7081011BEB704A2FD67FD33" bold="true" box="[151,759,691,718]" pageId="1" pageNumber="66">
<taxonomicName id="2ED94D00E7081011BEB704A2FD38FD33" ID-CoL="4BVT9" authority="Belousov &amp; Nyundo" authorityName="Belousov &amp; Nyundo" box="[151,680,691,718]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E7081011BEB704A2FE35FD30" bold="true" box="[151,421,692,718]" italics="true" pageId="1" pageNumber="66">Pachytrechodes brevis</emphasis>
Belousov &amp; Nyundo
</taxonomicName>
, 
<taxonomicNameLabel id="C09E57EAE7081011BC9504A5FD67FD33" box="[693,759,691,717]" pageId="1" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1C36508E7081011BEB704EDFE16FC38" pageId="1" pageNumber="66" type="materials_examined">
<paragraph id="E9663683E7081011BEB704EDFE16FC38" blockId="1.[151,1437,762,1974]" pageId="1" pageNumber="66">
<emphasis id="DBADEA91E7081011BEB704EDFDFFFCED" bold="true" box="[151,623,762,787]" pageId="1" pageNumber="66">
<typeStatus id="36628821E7081011BEB704EDFF42FCED" box="[151,210,763,787]" pageId="1" pageNumber="66">Type</typeStatus>
material: 
<typeStatus id="36628821E7081011BF7304EDFE54FCED" box="[339,452,763,787]" pageId="1" pageNumber="66" type="holotype">Holotype</typeStatus>
: 
<collectingCountry id="91CE7613E7081011BFF304ECFDFAFCED" box="[467,618,762,787]" name="Tanzania" pageId="1" pageNumber="66">TANZANIA</collectingCountry>
:
</emphasis>
1(1)3, “ 
<collectingCountry id="91CE7613E7081011BCFA04ECFCD2FCEC" box="[730,834,762,786]" name="Tanzania" pageId="1" pageNumber="66">Tanzania</collectingCountry>
, Udzungwa Mts Nat. Park; forest floor, 
<quantity id="2E219B66E7081011BB3404ECFAFCFCEC" box="[1300,1388,762,786]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="66" unit="m" value="1500.0">1500 m</quantity>
asl; 
<geoCoordinate id="8CED5044E7081011BEB70508FEB5FCC8" box="[151,293,798,822]" direction="east" orientation="longitude" pageId="1" pageNumber="66" precision="15" value="36.866665">E36º52´00´´</geoCoordinate>
<geoCoordinate id="8CED5044E7081011BF050508FE0EFCC8" box="[293,414,798,822]" direction="south" orientation="latitude" pageId="1" pageNumber="66" precision="15" value="-7.758333">S7º45´30´´</geoCoordinate>
; 
<date id="9D671043E7081011BF8B0508FDA2FCC8" box="[427,562,798,822]" pageId="1" pageNumber="66" value="1999-08-03">03.viii.1999</date>
Coll. B.A. Nyundo; searching on ground, daytime; B.A. Nyundo Expedition (
<date id="9D671043E7081011BEBF0554FEB1FCA4" box="[159,289,834,858]" pageId="1" pageNumber="66" value="1998" valueMax="2001">1998–2001</date>
)”(ZMUN). 
<emphasis id="DBADEA91E7081011BF8D0555FD48FCA5" bold="true" box="[429,728,834,859]" pageId="1" pageNumber="66">
<typeStatus id="36628821E7081011BF8D0555FDB9FCA5" box="[429,553,835,859]" pageId="1" pageNumber="66" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="91CE7613E7081011BC1C0554FD44FCA5" box="[572,724,834,859]" name="Tanzania" pageId="1" pageNumber="66">TANZANIA</collectingCountry>
:
</emphasis>
10(2)3, 14Ƥ, collected with the 
<typeStatus id="36628821E7081011BAA00554FB72FCA4" box="[1152,1250,834,858]" pageId="1" pageNumber="66" type="holotype">holotype</typeStatus>
(UDSM, ZISP, ZMUN); 1(1)3, 17Ƥ, same data but 
<date id="9D671043E7081011BC1A0570FD23FC80" box="[570,691,870,894]" pageId="1" pageNumber="66" value="1999-08-04">4.viii.1999</date>
Coll. Hashim Rashid (cBK, ZISP, ZMUN); 7(5)3, same data but 
<date id="9D671043E7081011BEB7059CFE80FC5C" box="[151,272,906,930]" pageId="1" pageNumber="66" value="1999-08-05">5.viii.1999</date>
(cBK, UDSM; ZISP, ZMUN); 2Ƥ, same data but 
<date id="9D671043E7081011BD7B059CFC41FC5C" box="[859,977,906,930]" pageId="1" pageNumber="66" value="1999-04-19">19.iv.1999</date>
Coll. Hezron Mwasomola (UDSM). 
<specimenCount id="FFDFFD0AE7081011BBA0059CFE9CFC38" pageId="1" pageNumber="66" type="generic">20 specimens</specimenCount>
measured.
</paragraph>
</subSubSection>
<subSubSection id="A1C36508E7081016BEE705C4FEFCFEE2" lastPageId="6" lastPageNumber="71" pageId="1" pageNumber="66" type="description">
<paragraph id="E9663683E7081011BEE705C4FF4EFB84" blockId="1.[151,1437,762,1974]" pageId="1" pageNumber="66">
<emphasis id="DBADEA91E7081011BEE705C4FEC9FC15" bold="true" box="[199,345,978,1003]" pageId="1" pageNumber="66">Description.</emphasis>
Very small-sized apterous species. Body length 
<quantity id="2E219B66E7081011BDAC05C4FBA4FC14" box="[908,1076,978,1002]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.18" metricValueMin="2.02" pageId="1" pageNumber="66" unit="mm" value="2.1" valueMax="2.18" valueMin="2.02">2.02–2.18 mm</quantity>
. Body robust and briefly ovate (
<figureCitation id="71E22A06E7081011BEBF05E0FF71FBF0" box="[159,225,1014,1038]" captionStart="FIGURE 1" captionStartId="2.[151,250,1991,2014]" captionText="FIGURE 1. Pachytrechodes brevis sp. n., habitus, female paratype." pageId="1" pageNumber="66">Fig. 1</figureCitation>
), weakly convex, with fairly long legs and antennae. Dorsal side unicolorous reddish brown to dark brown, with paler brown yellowish legs. First antennomere yellowish, antennomeres 2–7 more or less distinctly obscured, distal antennomeres beginning with 8–9 becoming gradually paler. Head and pronotum occasionally darker than elytra.
</paragraph>
<paragraph id="E9663683E7081011BEE70290FAB6F968" blockId="1.[151,1437,762,1974]" pageId="1" pageNumber="66">
Head rather small, much narrower than pronotum. Frontal furrows well and evenly impressed, arcuate, complete; frons convex. Eyes small, moderately protruding, more strongly in anterior third, slightly longer than tempora (eye length to temple length ratio 1.14–1.35), tempora strongly convex behind their mid-length, smooth, without any trace of pubescence. Labrum relatively narrow, its anterior margin deeply notched, bearing six setae, lateral lobes narrowly rounded. Mandibles (
<figureCitation id="71E22A06E7081011BCAB0300FD41FAD0" box="[651,721,1302,1326]" captionStart="FIGURE 2" captionStartId="3.[151,250,1005,1028]" captionText="FIGURE 2. Pachytrechodes brevis sp. n., mandibles: a—left, b—right." pageId="1" pageNumber="66">Fig. 2</figureCitation>
) rather short and thin, curved in apical quarter. Tooth on right mandible tridentate, with long base (i.e. proximal and distal denticles widely spaced). Last segment of maxillary palpi only little shorter than penultimate segment but much more narrow, subulate. Maxillary palpi completely glabrous. Penultimate segment of labial palpi quadrisetose. Mentum and submentum completely fused, without any trace of labial suture. Labial tooth simple, with anterior margin widely rounded to truncate. Four submental setae of which two are subangular. Glossum clearly notched at apex, with two large median setae and a few much smaller lateral setae. Paraglossae thin and long, faintly curved, protruding far beyond anterior margin of glossum. Two supraorbital setiferous pores on each side of head located on lines divergent posteriad. Anterior supraorbital setiferous pore strongly foveolate, posterior one without oblique fovea. Parietal transverse impression very shallow. Antennae rather long and thin, antennomeres 2 and 3 subequal in length, the latter being only marginally longer. Viewed from above, antennomere 2 nearly cylindrical, without distinct constriction in basal part.
</paragraph>
<paragraph id="E9663683E7081013BEE700B5FD8DFEC1" blockId="1.[151,1437,762,1974]" lastBlockId="3.[151,1436,151,319]" lastPageId="3" lastPageNumber="68" pageId="1" pageNumber="66">
Pronotum strongly transverse and convex, with basal lobe rather narrow and feebly produced; sides broadly rounded in anterior and middle part, nearly straight in posterior part, without distinct sinuation before hind angles. Lateral gutter very narrow throughout, only marginally widened near hind angles; lateral border complete, continued to basal lobe and more or less reduced only in most median part. Basal margin of pronotum weakly concave medially, obliquely emarginated laterally, most strongly sinuate near basal foveae, the latter extraordinarily small but deep, pore-like, attached to lateral border. 
<collectingCountry id="91CE7613E7081011BDAB0140FC53F890" box="[907,963,1878,1902]" name="India" pageId="1" pageNumber="66">Hind</collectingCountry>
angles of pronotum obtusangular, blunt at apices or even completely rounded off. Front angles broadly rounded but detectable, weakly produced anteriad. Anterior margin of pronotum moderately and evenly concave. Apical transverse impression shallow, normally distinct laterally, reduced medially. Prebasal transverse impression very shallow, consisting of a median triangular depression and occasionally irregular lateral foveae. Median line finely engraved, normally developed only in median part. Discal foveae missing. Base of pronoum smooth, or, at most, with a few very shallow wrinkles. Anterior lateral seta of pronotum located markedly before mid-length of pronotum, slightly before its broadest point, posterior one in hind angles.
</paragraph>
<caption id="BDA6660BE70B1012BEB701D1FCD8F823" box="[151,840,1991,2014]" pageId="2" pageNumber="67">
<paragraph id="E9663683E70B1012BEB701D1FCD8F823" blockId="2.[151,840,1991,2014]" box="[151,840,1991,2014]" pageId="2" pageNumber="67">
<emphasis id="DBADEA91E70B1012BEB701D1FE83F823" bold="true" box="[151,275,1991,2014]" pageId="2" pageNumber="67">FIGURE 1.</emphasis>
<taxonomicName id="2ED94D00E70B1012BF3901D1FE6DF823" box="[281,509,1991,2013]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70B1012BF3901D1FE6DF823" box="[281,509,1991,2013]" italics="true" pageId="2" pageNumber="67">Pachytrechodes brevis</emphasis>
</taxonomicName>
<emphasis id="DBADEA91E70B1012BC2301DEFDACF823" bold="true" box="[515,572,1992,2013]" pageId="2" pageNumber="67">
<taxonomicNameLabel id="C09E57EAE70B1012BC2301DEFDACF823" box="[515,572,1992,2013]" pageId="2" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, habitus, female paratype.
</paragraph>
</caption>
<caption id="BDA6660BE70A1013BEB705FBFCFEFBFD" box="[151,878,1005,1028]" pageId="3" pageNumber="68">
<paragraph id="E9663683E70A1013BEB705FBFCFEFBFD" blockId="3.[151,878,1005,1028]" box="[151,878,1005,1028]" pageId="3" pageNumber="68">
<emphasis id="DBADEA91E70A1013BEB705FBFE83FBFD" bold="true" box="[151,275,1005,1028]" pageId="3" pageNumber="68">FIGURE 2.</emphasis>
<taxonomicName id="2ED94D00E70A1013BF3905F8FE6DFBFA" box="[281,509,1006,1028]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70A1013BF3905F8FE6DFBFA" box="[281,509,1006,1028]" italics="true" pageId="3" pageNumber="68">Pachytrechodes brevis</emphasis>
</taxonomicName>
<emphasis id="DBADEA91E70A1013BC2305F8FDACFBFD" bold="true" box="[515,572,1006,1027]" pageId="3" pageNumber="68">
<taxonomicNameLabel id="C09E57EAE70A1013BC2305F8FDACFBFD" box="[515,572,1006,1027]" pageId="3" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, mandibles: a—left, b—right.
</paragraph>
</caption>
<paragraph id="E9663683E70A1013BEE7023DFE57F90D" blockId="3.[151,1437,1067,1995]" pageId="3" pageNumber="68">Elytra briefly ovate, more or less flattened on disk, steeply sloped laterally, with maximum width near midlength, sides broadly rounded at humeral area, nearly parallel in median part (though varying in this respect), broadly rounded at apex, with distinct preapical sinuation posteriorly. Lateral gutter evenly wide throughout its length, lateral margins moderately to weakly deflexed, without serration in humeral area. Basal border of elytra complete, with distinct sinuation at level of dorsal stria 4. Elytral striae rather well-impressed, at least five inner striae distinct in their median part, though becoming increasingly shallower and shorter toward sides, sixth stria distinguishable only fragmentary. First stria much more sharply engraved as compared to other striae, smooth, rectilinear, without distinct sinuation in anterior third while all other dorsal striae impressed more shallowly, waved and faintly punctured. Third stria angularly curved at anterior discal setiferous pore. Eighth stria distinct only in posterior half of elytra (only marginally extended anteriad beyond umbilicate pore 5) and some fragments may be distinguishable in humeral area. Interspaces flat. Parascutellar pore well developed, located anteriorly of basal border of elytra. Parascutellar striole sharply impressed, partially overlapped with basal portion of first discal stria. Apical striole clearly sinuate, moderately impressed posteriorly and very shallow anteriorly. Two discal setiferous pores in third stria (normally anterior one at level between umbilicate pores 3 and 4, and posterior one slightly before umbilicate pore 5) and one preapical pore located close to apex, in apical cross of third and second elytral striae at level slightly anteriorly of umbilicate pore 8. Both discal setiferous pores clearly foveolate, their setae very short and thin, shorter than parascutellar seta. Both apical pores present, located very closely to each other, anguloapical one being much less developed and located rather far from elytral suture. Umbilicate setiferous pores wellaggregated in longitudinal direction, but some pores (pores 3 and 4 weakly, pore 5 very strongly) translocated inward, onto disk of elytra.</paragraph>
<paragraph id="E9663683E70A1013BEE700E9FBDDF8E9" blockId="3.[151,1437,1067,1995]" box="[199,1101,1791,1815]" pageId="3" pageNumber="68">Metathoracical wings reduced to small vestiges shorter than one fourth of elytra.</paragraph>
<paragraph id="E9663683E70A1013BEE70132FB0BF8A1" blockId="3.[151,1437,1067,1995]" pageId="3" pageNumber="68">Legs rather thin and long, tibiae weakly curved. Protibiae without distinct pubescence on their anterior surface, sharply grooved exteriorly. Two basal segments in male protarsi dilated and dentate inward.</paragraph>
<paragraph id="E9663683E70A1013BEE7017DFC30F835" blockId="3.[151,1437,1067,1995]" pageId="3" pageNumber="68">Ventral side of abdomen rather evenly pubescent except for lateral parts, two paramedian setae on visible abdominal ventrites 4–6. Two setae along posterior margin of anal ventrite in male and four setae in female. Posterior margin of anal ventrite concave in male, rounded in female.</paragraph>
<paragraph id="E9663683E70D1014BEB7068EFD86FE77" blockId="4.[151,1436,152,393]" pageId="4" pageNumber="69">
<emphasis id="DBADEA91E70D1014BEB7068EFE96FF53" bold="true" box="[151,262,152,173]" pageId="4" pageNumber="69">TABLE 1.</emphasis>
Morphometric characters of 
<taxonomicName id="2ED94D00E70D1014BC14068EFC8AFF50" box="[564,794,152,174]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70D1014BC14068EFC8AFF50" box="[564,794,152,174]" italics="true" pageId="4" pageNumber="69">Pachytrechodes brevis</emphasis>
</taxonomicName>
, 
<emphasis id="DBADEA91E70D1014BD09068EFCF4FF53" bold="true" box="[809,868,152,173]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="C09E57EAE70D1014BD09068EFCF4FF53" box="[809,868,152,173]" pageId="4" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
The following abbreviations are used: 
<emphasis id="DBADEA91E70D1014BADC068FFABBFF50" box="[1276,1323,153,174]" italics="true" pageId="4" pageNumber="69">a_pr</emphasis>
—width of pronotum at apical margin; 
<emphasis id="DBADEA91E70D1014BF9206AEFE4DFF33" box="[434,477,184,205]" italics="true" pageId="4" pageNumber="69">ante</emphasis>
—length of antenna; 
<emphasis id="DBADEA91E70D1014BC9606A1FD76FF33" box="[694,742,183,205]" italics="true" pageId="4" pageNumber="69">b_pr</emphasis>
—width of pronotum at basal margin; 
<emphasis id="DBADEA91E70D1014BA4F06A1FB0CFF33" box="[1135,1180,182,205]" italics="true" pageId="4" pageNumber="69">
d_ 
<emphasis id="DBADEA91E70D1014BAA806A0FB1FFF32" bold="true" box="[1160,1167,182,204]" italics="true" pageId="4" pageNumber="69">i</emphasis>
p
</emphasis>
—distance between level of 
<emphasis id="DBADEA91E70D1014BE9306C1FF5DFF1F" box="[179,205,213,238]" italics="true" pageId="4" pageNumber="69">
<emphasis id="DBADEA91E70D1014BE9306C1FF2AFF13" bold="true" box="[179,186,215,237]" italics="true" pageId="4" pageNumber="69">i</emphasis>
-th
</emphasis>
discal setiferous pore and basal border of elytra; 
<emphasis id="DBADEA91E70D1014BCE506CFFD77FF10" box="[709,743,217,238]" italics="true" pageId="4" pageNumber="69">eye</emphasis>
—length of eye in dorsal view; 
<emphasis id="DBADEA91E70D1014BA0B06C1FBC8FF13" box="[1067,1112,215,237]" italics="true" pageId="4" pageNumber="69">h_ta</emphasis>
—length of tarsus; 
<emphasis id="DBADEA91E70D1014BB3C06C1FAD3FF13" box="[1308,1347,215,237]" italics="true" pageId="4" pageNumber="69">h_ti</emphasis>
—length of tibia; 
<emphasis id="DBADEA91E70D1014BED406E0FEA3FEF2" box="[244,307,246,268]" italics="true" pageId="4" pageNumber="69">height</emphasis>
—maximum height of body across elytra; 
<emphasis id="DBADEA91E70D1014BCD806E0FCB5FEF2" box="[760,805,246,268]" italics="true" pageId="4" pageNumber="69">l_2a</emphasis>
—length of second antennal segment; 
<emphasis id="DBADEA91E70D1014BA9D06E0FB7AFEF2" box="[1213,1258,246,268]" italics="true" pageId="4" pageNumber="69">l_3a</emphasis>
—length of third antennal segment; 
<emphasis id="DBADEA91E70D1014BF7F0703FE15FED5" box="[351,389,277,299]" italics="true" pageId="4" pageNumber="69">l_el</emphasis>
—length of elytra; 
<emphasis id="DBADEA91E70D1014BC760703FD10FED5" box="[598,640,277,299]" italics="true" pageId="4" pageNumber="69">l_pr</emphasis>
—length of pronotum along median line; 
<emphasis id="DBADEA91E70D1014BA650700FBE2FED5" box="[1093,1138,278,299]" italics="true" pageId="4" pageNumber="69">s_pr</emphasis>
—distance between level of anterior lateral seta of pronotum and anterior margin of pronotum; 
<emphasis id="DBADEA91E70D1014BD1A0723FCFAFEB4" box="[826,874,309,330]" italics="true" pageId="4" pageNumber="69">tmpr</emphasis>
—length of tempora; 
<emphasis id="DBADEA91E70D1014BA640723FBE6FEB7" box="[1092,1142,307,330]" italics="true" pageId="4" pageNumber="69">
um_ 
<emphasis id="DBADEA91E70D1014BA4F0725FBE6FEB7" bold="true" box="[1135,1142,307,329]" italics="true" pageId="4" pageNumber="69">i</emphasis>
</emphasis>
—distance from basal border of elytra to level of 
<emphasis id="DBADEA91E70D1014BF450743FEFCFE95" bold="true" box="[357,364,341,363]" italics="true" pageId="4" pageNumber="69">i</emphasis>
th umbilicate pore; 
<emphasis id="DBADEA91E70D1014BC090743FDCFFE95" box="[553,607,341,363]" italics="true" pageId="4" pageNumber="69">w_3a</emphasis>
—width of third antennal segment; 
<emphasis id="DBADEA91E70D1014BDEE0743FB95FE95" box="[974,1029,341,363]" italics="true" pageId="4" pageNumber="69">w_hd</emphasis>
—maximum width of head across eyes; 
<emphasis id="DBADEA91E70D1014BEB70762FF5BFE77" box="[151,203,372,393]" italics="true" pageId="4" pageNumber="69">w_pr</emphasis>
—maximum width of pronotum.
</paragraph>
<paragraph id="E9663683E70D1014BEBF07B3FACFF942" pageId="4" pageNumber="69">
<table id="9BD9C423E70DEFEFBEBF07B3FACFF942" box="[159,1375,421,1724]" gridcols="6" gridrows="31" pageId="4" pageNumber="69">
<tr id="57E934C1E70DEFEFBEBF07B3FACFFE45" box="[159,1375,421,443]" gridrow="0" pageId="4" pageNumber="69" rowspan-1="1" rowspan-3="1" rowspan-5="1">
<th id="14385DBDE70DEFEFBEBF07B3FDBCFE45" box="[159,556,421,443]" gridcol="0" gridrow="0" pageId="4" pageNumber="69">Indices Males</th>
<th id="14385DBDE70DEFEFBCD407B3FC0BFE45" box="[756,923,421,443]" gridcol="2" gridrow="0" pageId="4" pageNumber="69">Females</th>
<th id="14385DBDE70DEFEFBA6C07B3FB63FE45" box="[1100,1267,421,443]" gridcol="4" gridrow="0" pageId="4" pageNumber="69">Total</th>
</tr>
<tr id="57E934C1E70DEFEFBEBF07D9FACFFDF3" box="[159,1375,463,525]" gridrow="1" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF07D9FDBCFDF3" box="[159,556,463,525]" gridcol="0" gridrow="1" pageId="4" pageNumber="69">
Range (average) eye/tmpr 1.16–1.35(
<emphasis id="DBADEA91E70D1014BFD307EEFD8FFDF3" bold="true" box="[499,543,504,525]" pageId="4" pageNumber="69">1.28</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB407D9FD3EFDF3" box="[660,686,463,525]" gridcol="1" gridrow="1" pageId="4" pageNumber="69">N 10</td>
<td id="14385DBDE70DEFEFBCD407D9FC0BFDF3" box="[756,923,463,525]" gridcol="2" gridrow="1" pageId="4" pageNumber="69">
Range (average) 1.14–1.28(
<emphasis id="DBADEA91E70D1014BD4207EEFC1EFDF3" bold="true" box="[866,910,504,525]" pageId="4" pageNumber="69">1.21</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD07D9FB97FDF3" box="[1005,1031,463,525]" gridcol="3" gridrow="1" pageId="4" pageNumber="69">N 10</td>
<td id="14385DBDE70DEFEFBA6C07D9FB63FDF3" box="[1100,1267,463,525]" gridcol="4" gridrow="1" pageId="4" pageNumber="69">
Range (average) 1.14–1.35(
<emphasis id="DBADEA91E70D1014BA9A07EEFB76FDF3" bold="true" box="[1210,1254,504,525]" pageId="4" pageNumber="69">1.24</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6507D9FACFFDF3" box="[1349,1375,463,525]" gridcol="5" gridrow="1" pageId="4" pageNumber="69">N 20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0437FACFFDC9" box="[159,1375,545,567]" gridrow="2" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0437FDBCFDC9" box="[159,556,545,567]" gridcol="0" gridrow="2" pageId="4" pageNumber="69">
l_3a/l_2a 1.00–1.07(
<emphasis id="DBADEA91E70D1014BFD30437FD8FFDC8" bold="true" box="[499,543,545,566]" pageId="4" pageNumber="69">1.03</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40437FD3EFDC9" box="[660,686,545,567]" gridcol="1" gridrow="2" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40437FC0BFDC9" box="[756,923,545,567]" gridcol="2" gridrow="2" pageId="4" pageNumber="69">
1.00–1.07(
<emphasis id="DBADEA91E70D1014BD420437FC1EFDC8" bold="true" box="[866,910,545,566]" pageId="4" pageNumber="69">1.03</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0437FB97FDC9" box="[1005,1031,545,567]" gridcol="3" gridrow="2" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0437FB63FDC9" box="[1100,1267,545,567]" gridcol="4" gridrow="2" pageId="4" pageNumber="69">
1.00–1.07(
<emphasis id="DBADEA91E70D1014BA9A0437FB76FDC8" bold="true" box="[1210,1254,545,566]" pageId="4" pageNumber="69">1.03</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650437FACFFDC9" box="[1349,1375,545,567]" gridcol="5" gridrow="2" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF045CFACFFD9E" box="[159,1375,586,608]" gridrow="3" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF045CFDBCFD9E" box="[159,556,586,608]" gridcol="0" gridrow="3" pageId="4" pageNumber="69">
l_3a/w_3a 2.48–2.67(
<emphasis id="DBADEA91E70D1014BFD3045CFD8FFDA1" bold="true" box="[499,543,586,607]" pageId="4" pageNumber="69">2.56</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4045CFD3EFD9E" box="[660,686,586,608]" gridcol="1" gridrow="3" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4045CFC0BFD9E" box="[756,923,586,608]" gridcol="2" gridrow="3" pageId="4" pageNumber="69">
2.33–2.54(
<emphasis id="DBADEA91E70D1014BD42045CFC1EFDA1" bold="true" box="[866,910,586,607]" pageId="4" pageNumber="69">2.45</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD045CFB97FD9E" box="[1005,1031,586,608]" gridcol="3" gridrow="3" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C045CFB63FD9E" box="[1100,1267,586,608]" gridcol="4" gridrow="3" pageId="4" pageNumber="69">
2.33–2.67(
<emphasis id="DBADEA91E70D1014BA9A045CFB76FDA1" bold="true" box="[1210,1254,586,607]" pageId="4" pageNumber="69">2.51</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65045CFACFFD9E" box="[1349,1375,586,608]" gridcol="5" gridrow="3" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0465FACFFD77" box="[159,1375,627,649]" gridrow="4" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0465FDBCFD77" box="[159,556,627,649]" gridcol="0" gridrow="4" pageId="4" pageNumber="69">
eye/l_3a 1.00–1.26(
<emphasis id="DBADEA91E70D1014BFD30465FD8FFD76" bold="true" box="[499,543,627,648]" pageId="4" pageNumber="69">1.09</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40465FD3EFD77" box="[660,686,627,649]" gridcol="1" gridrow="4" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40465FC0BFD77" box="[756,923,627,649]" gridcol="2" gridrow="4" pageId="4" pageNumber="69">
1.07–1.19(
<emphasis id="DBADEA91E70D1014BD420465FC1EFD76" bold="true" box="[866,910,627,648]" pageId="4" pageNumber="69">1.13</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0465FB97FD77" box="[1005,1031,627,649]" gridcol="3" gridrow="4" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0465FB63FD77" box="[1100,1267,627,649]" gridcol="4" gridrow="4" pageId="4" pageNumber="69">
1.00–1.26(
<emphasis id="DBADEA91E70D1014BA9A0465FB75FD76" bold="true" box="[1210,1253,627,648]" pageId="4" pageNumber="69">1.11</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650465FACFFD77" box="[1349,1375,627,649]" gridcol="5" gridrow="4" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF048BFACFFD4D" box="[159,1375,669,691]" gridrow="5" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF048BFDBCFD4D" box="[159,556,669,691]" gridcol="0" gridrow="5" pageId="4" pageNumber="69">
(s_pr/l_pr) x 100 22.7–28.6(
<emphasis id="DBADEA91E70D1014BFD3048BFD8FFD4C" bold="true" box="[499,543,669,690]" pageId="4" pageNumber="69">26.1</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4048BFD3EFD4D" box="[660,686,669,691]" gridcol="1" gridrow="5" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4048BFC0BFD4D" box="[756,923,669,691]" gridcol="2" gridrow="5" pageId="4" pageNumber="69">
23.9–28.6(
<emphasis id="DBADEA91E70D1014BD42048BFC1EFD4C" bold="true" box="[866,910,669,690]" pageId="4" pageNumber="69">26.4</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD048BFB97FD4D" box="[1005,1031,669,691]" gridcol="3" gridrow="5" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C048BFB63FD4D" box="[1100,1267,669,691]" gridcol="4" gridrow="5" pageId="4" pageNumber="69">
22.7–28.6(
<emphasis id="DBADEA91E70D1014BA9A048BFB76FD4C" bold="true" box="[1210,1254,669,690]" pageId="4" pageNumber="69">26.3</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65048BFACFFD4D" box="[1349,1375,669,691]" gridcol="5" gridrow="5" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF04D0FACFFD22" box="[159,1375,710,732]" gridrow="6" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF04D0FDBCFD22" box="[159,556,710,732]" gridcol="0" gridrow="6" pageId="4" pageNumber="69">
length 2.02–2.17(
<emphasis id="DBADEA91E70D1014BFD304D0FD8FFD25" bold="true" box="[499,543,710,731]" pageId="4" pageNumber="69">2.09</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB404D0FD3EFD22" box="[660,686,710,732]" gridcol="1" gridrow="6" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD404D0FC0BFD22" box="[756,923,710,732]" gridcol="2" gridrow="6" pageId="4" pageNumber="69">
2.02–2.18(
<emphasis id="DBADEA91E70D1014BD4204D0FC1EFD25" bold="true" box="[866,910,710,731]" pageId="4" pageNumber="69">2.13</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD04D0FB97FD22" box="[1005,1031,710,732]" gridcol="3" gridrow="6" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C04D0FB63FD22" box="[1100,1267,710,732]" gridcol="4" gridrow="6" pageId="4" pageNumber="69">
2.02–2.18(
<emphasis id="DBADEA91E70D1014BA9A04D0FB75FD25" bold="true" box="[1210,1253,710,731]" pageId="4" pageNumber="69">2.11</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6504D0FACFFD22" box="[1349,1375,710,732]" gridcol="5" gridrow="6" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF04F9FACFFCFB" box="[159,1375,751,773]" gridrow="7" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF04F9FDBCFCFB" box="[159,556,751,773]" gridcol="0" gridrow="7" pageId="4" pageNumber="69">
l_el/height 2.57–2.80(
<emphasis id="DBADEA91E70D1014BFD304E6FD8FFCFB" bold="true" box="[499,543,752,773]" pageId="4" pageNumber="69">2.68</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB404F9FD3EFCFB" box="[660,686,751,773]" gridcol="1" gridrow="7" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD404F9FC0BFCFB" box="[756,923,751,773]" gridcol="2" gridrow="7" pageId="4" pageNumber="69">
2.43–2.74(
<emphasis id="DBADEA91E70D1014BD4204E6FC1EFCFB" bold="true" box="[866,910,752,773]" pageId="4" pageNumber="69">2.59</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD04F9FB97FCFB" box="[1005,1031,751,773]" gridcol="3" gridrow="7" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C04F9FB63FCFB" box="[1100,1267,751,773]" gridcol="4" gridrow="7" pageId="4" pageNumber="69">
2.43–2.80(
<emphasis id="DBADEA91E70D1014BA9A04E6FB76FCFB" bold="true" box="[1210,1254,752,773]" pageId="4" pageNumber="69">2.63</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6504F9FACFFCFB" box="[1349,1375,751,773]" gridcol="5" gridrow="7" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF050FFACFFCD1" box="[159,1375,793,815]" gridrow="8" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF050FFDBCFCD1" box="[159,556,793,815]" gridcol="0" gridrow="8" pageId="4" pageNumber="69">
l_el/h_ti 2.43–2.59(
<emphasis id="DBADEA91E70D1014BFD3050FFD8FFCD0" bold="true" box="[499,543,793,814]" pageId="4" pageNumber="69">2.50</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4050FFD3EFCD1" box="[660,686,793,815]" gridcol="1" gridrow="8" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4050FFC0BFCD1" box="[756,923,793,815]" gridcol="2" gridrow="8" pageId="4" pageNumber="69">
2.42–2.56(
<emphasis id="DBADEA91E70D1014BD42050FFC1EFCD0" bold="true" box="[866,910,793,814]" pageId="4" pageNumber="69">2.48</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD050FFB97FCD1" box="[1005,1031,793,815]" gridcol="3" gridrow="8" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C050FFB63FCD1" box="[1100,1267,793,815]" gridcol="4" gridrow="8" pageId="4" pageNumber="69">
2.42–2.59(
<emphasis id="DBADEA91E70D1014BA9A050FFB76FCD0" bold="true" box="[1210,1254,793,814]" pageId="4" pageNumber="69">2.49</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65050FFACFFCD1" box="[1349,1375,793,815]" gridcol="5" gridrow="8" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0554FACFFCA6" box="[159,1375,834,856]" gridrow="9" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0554FDBCFCA6" box="[159,556,834,856]" gridcol="0" gridrow="9" pageId="4" pageNumber="69">
l_el/l_pr 2.63–2.79(
<emphasis id="DBADEA91E70D1014BFD30554FD8FFCA9" bold="true" box="[499,543,834,855]" pageId="4" pageNumber="69">2.73</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40554FD3EFCA6" box="[660,686,834,856]" gridcol="1" gridrow="9" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40554FC0BFCA6" box="[756,923,834,856]" gridcol="2" gridrow="9" pageId="4" pageNumber="69">
2.64–2.79(
<emphasis id="DBADEA91E70D1014BD420554FC1EFCA9" bold="true" box="[866,910,834,855]" pageId="4" pageNumber="69">2.70</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0554FB97FCA6" box="[1005,1031,834,856]" gridcol="3" gridrow="9" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0554FB63FCA6" box="[1100,1267,834,856]" gridcol="4" gridrow="9" pageId="4" pageNumber="69">
2.63–2.79(
<emphasis id="DBADEA91E70D1014BA9A0554FB76FCA9" bold="true" box="[1210,1254,834,855]" pageId="4" pageNumber="69">2.72</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650554FACFFCA6" box="[1349,1375,834,856]" gridcol="5" gridrow="9" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF057DFACFFC7F" box="[159,1375,875,897]" gridrow="10" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF057DFDBCFC7F" box="[159,556,875,897]" gridcol="0" gridrow="10" pageId="4" pageNumber="69">
h_ti/h_ta 1.42–1.50(
<emphasis id="DBADEA91E70D1014BFD3057DFD8FFC7E" bold="true" box="[499,543,875,896]" pageId="4" pageNumber="69">1.46</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4057DFD3EFC7F" box="[660,686,875,897]" gridcol="1" gridrow="10" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4057DFC0BFC7F" box="[756,923,875,897]" gridcol="2" gridrow="10" pageId="4" pageNumber="69">
1.45–1.51(
<emphasis id="DBADEA91E70D1014BD42057DFC1EFC7E" bold="true" box="[866,910,875,896]" pageId="4" pageNumber="69">1.48</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD057DFB97FC7F" box="[1005,1031,875,897]" gridcol="3" gridrow="10" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C057DFB63FC7F" box="[1100,1267,875,897]" gridcol="4" gridrow="10" pageId="4" pageNumber="69">
1.42–1.51(
<emphasis id="DBADEA91E70D1014BA9A057DFB76FC7E" bold="true" box="[1210,1254,875,896]" pageId="4" pageNumber="69">1.47</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65057DFACFFC7F" box="[1349,1375,875,897]" gridcol="5" gridrow="10" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0583FACFFC55" box="[159,1375,917,939]" gridrow="11" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0583FDBCFC55" box="[159,556,917,939]" gridcol="0" gridrow="11" pageId="4" pageNumber="69">
w_pr/w_hd 1.45–1.53(
<emphasis id="DBADEA91E70D1014BFD30583FD8FFC54" bold="true" box="[499,543,917,938]" pageId="4" pageNumber="69">1.49</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40583FD3EFC55" box="[660,686,917,939]" gridcol="1" gridrow="11" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40583FC0BFC55" box="[756,923,917,939]" gridcol="2" gridrow="11" pageId="4" pageNumber="69">
1.41–1.49(
<emphasis id="DBADEA91E70D1014BD420583FC1EFC54" bold="true" box="[866,910,917,938]" pageId="4" pageNumber="69">1.46</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0583FB97FC55" box="[1005,1031,917,939]" gridcol="3" gridrow="11" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0583FB63FC55" box="[1100,1267,917,939]" gridcol="4" gridrow="11" pageId="4" pageNumber="69">
1.41–1.53(
<emphasis id="DBADEA91E70D1014BA9A0583FB76FC54" bold="true" box="[1210,1254,917,938]" pageId="4" pageNumber="69">1.47</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650583FACFFC55" box="[1349,1375,917,939]" gridcol="5" gridrow="11" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF05A8FACFFC2A" box="[159,1375,958,980]" gridrow="12" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF05A8FDBCFC2A" box="[159,556,958,980]" gridcol="0" gridrow="12" pageId="4" pageNumber="69">
l_el/ante 0.98–1.01(
<emphasis id="DBADEA91E70D1014BFD305A8FD8FFC2D" bold="true" box="[499,543,958,979]" pageId="4" pageNumber="69">1.00</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB405A8FD3EFC2A" box="[660,686,958,980]" gridcol="1" gridrow="12" pageId="4" pageNumber="69">9</td>
<td id="14385DBDE70DEFEFBCD405A8FC0BFC2A" box="[756,923,958,980]" gridcol="2" gridrow="12" pageId="4" pageNumber="69">
0.99–1.03(
<emphasis id="DBADEA91E70D1014BD4205A8FC1EFC2D" bold="true" box="[866,910,958,979]" pageId="4" pageNumber="69">1.00</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD05A8FB97FC2A" box="[1005,1031,958,980]" gridcol="3" gridrow="12" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C05A8FB63FC2A" box="[1100,1267,958,980]" gridcol="4" gridrow="12" pageId="4" pageNumber="69">
0.98–1.03(
<emphasis id="DBADEA91E70D1014BA9A05A8FB76FC2D" bold="true" box="[1210,1254,958,979]" pageId="4" pageNumber="69">1.00</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6505A8FACFFC2A" box="[1349,1375,958,980]" gridcol="5" gridrow="12" pageId="4" pageNumber="69">19</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF05F1FACFFC03" box="[159,1375,999,1021]" gridrow="13" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF05F1FDBCFC03" box="[159,556,999,1021]" gridcol="0" gridrow="13" pageId="4" pageNumber="69">
w_pr/l_pr 1.40–1.49(
<emphasis id="DBADEA91E70D1014BFD305FEFD8FFC03" bold="true" box="[499,543,1000,1021]" pageId="4" pageNumber="69">1.45</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB405F1FD3EFC03" box="[660,686,999,1021]" gridcol="1" gridrow="13" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD405F1FC0BFC03" box="[756,923,999,1021]" gridcol="2" gridrow="13" pageId="4" pageNumber="69">
1.43–1.49(
<emphasis id="DBADEA91E70D1014BD4205FEFC1EFC03" bold="true" box="[866,910,1000,1021]" pageId="4" pageNumber="69">1.46</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD05F1FB97FC03" box="[1005,1031,999,1021]" gridcol="3" gridrow="13" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C05F1FB63FC03" box="[1100,1267,999,1021]" gridcol="4" gridrow="13" pageId="4" pageNumber="69">
1.40–1.49(
<emphasis id="DBADEA91E70D1014BA9A05FEFB76FC03" bold="true" box="[1210,1254,1000,1021]" pageId="4" pageNumber="69">1.45</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6505F1FACFFC03" box="[1349,1375,999,1021]" gridcol="5" gridrow="13" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0207FACFFBD9" box="[159,1375,1041,1063]" gridrow="14" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0207FDBCFBD9" box="[159,556,1041,1063]" gridcol="0" gridrow="14" pageId="4" pageNumber="69">
w_pr/b_pr 1.22–1.31(
<emphasis id="DBADEA91E70D1014BFD30207FD8FFBD8" bold="true" box="[499,543,1041,1062]" pageId="4" pageNumber="69">1.26</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40207FD3EFBD9" box="[660,686,1041,1063]" gridcol="1" gridrow="14" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40207FC0BFBD9" box="[756,923,1041,1063]" gridcol="2" gridrow="14" pageId="4" pageNumber="69">
1.26–1.31(
<emphasis id="DBADEA91E70D1014BD420207FC1EFBD8" bold="true" box="[866,910,1041,1062]" pageId="4" pageNumber="69">1.28</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0207FB97FBD9" box="[1005,1031,1041,1063]" gridcol="3" gridrow="14" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0207FB63FBD9" box="[1100,1267,1041,1063]" gridcol="4" gridrow="14" pageId="4" pageNumber="69">
1.22–1.31(
<emphasis id="DBADEA91E70D1014BA9A0207FB76FBD8" bold="true" box="[1210,1254,1041,1062]" pageId="4" pageNumber="69">1.27</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650207FACFFBD9" box="[1349,1375,1041,1063]" gridcol="5" gridrow="14" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF022CFACFFBAE" box="[159,1375,1082,1104]" gridrow="15" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF022CFDBCFBAE" box="[159,556,1082,1104]" gridcol="0" gridrow="15" pageId="4" pageNumber="69">
a_pr/b_pr 0.81–0.85(
<emphasis id="DBADEA91E70D1014BFD3022CFD8FFBB1" bold="true" box="[499,543,1082,1103]" pageId="4" pageNumber="69">0.83</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4022CFD3EFBAE" box="[660,686,1082,1104]" gridcol="1" gridrow="15" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4022CFC0BFBAE" box="[756,923,1082,1104]" gridcol="2" gridrow="15" pageId="4" pageNumber="69">
0.83–0.86(
<emphasis id="DBADEA91E70D1014BD42022CFC1EFBB1" bold="true" box="[866,910,1082,1103]" pageId="4" pageNumber="69">0.84</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD022CFB97FBAE" box="[1005,1031,1082,1104]" gridcol="3" gridrow="15" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C022CFB63FBAE" box="[1100,1267,1082,1104]" gridcol="4" gridrow="15" pageId="4" pageNumber="69">
0.81–0.86(
<emphasis id="DBADEA91E70D1014BA9A022CFB76FBB1" bold="true" box="[1210,1254,1082,1103]" pageId="4" pageNumber="69">0.83</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65022CFACFFBAE" box="[1349,1375,1082,1104]" gridcol="5" gridrow="15" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0275FACFFB84" box="[159,1375,1123,1146]" gridrow="16" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0275FDBCFB84" box="[159,556,1123,1146]" gridcol="0" gridrow="16" pageId="4" pageNumber="69">
b_pr/a_pr 1.17–1.23(
<emphasis id="DBADEA91E70D1014BFD30272FD8FFB87" bold="true" box="[499,543,1124,1145]" pageId="4" pageNumber="69">1.21</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40275FD3EFB84" box="[660,686,1123,1146]" gridcol="1" gridrow="16" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40275FC0BFB84" box="[756,923,1123,1146]" gridcol="2" gridrow="16" pageId="4" pageNumber="69">
1.16–1.21(
<emphasis id="DBADEA91E70D1014BD420272FC1EFB87" bold="true" box="[866,910,1124,1145]" pageId="4" pageNumber="69">1.18</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0275FB97FB84" box="[1005,1031,1123,1146]" gridcol="3" gridrow="16" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0275FB63FB84" box="[1100,1267,1123,1146]" gridcol="4" gridrow="16" pageId="4" pageNumber="69">
1.16–1.23(
<emphasis id="DBADEA91E70D1014BA9A0272FB76FB87" bold="true" box="[1210,1254,1124,1145]" pageId="4" pageNumber="69">1.20</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650275FACFFB84" box="[1349,1375,1123,1146]" gridcol="5" gridrow="16" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF029BFACFFB5D" box="[159,1375,1165,1187]" gridrow="17" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF029BFDBCFB5D" box="[159,556,1165,1187]" gridcol="0" gridrow="17" pageId="4" pageNumber="69">
l_el/w_el 1.27–1.32(
<emphasis id="DBADEA91E70D1014BFD3029BFD8FFB5C" bold="true" box="[499,543,1165,1186]" pageId="4" pageNumber="69">1.30</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4029BFD3EFB5D" box="[660,686,1165,1187]" gridcol="1" gridrow="17" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4029BFC0BFB5D" box="[756,923,1165,1187]" gridcol="2" gridrow="17" pageId="4" pageNumber="69">
1.27–1.31(
<emphasis id="DBADEA91E70D1014BD42029BFC1EFB5C" bold="true" box="[866,910,1165,1186]" pageId="4" pageNumber="69">1.28</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD029BFB97FB5D" box="[1005,1031,1165,1187]" gridcol="3" gridrow="17" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C029BFB63FB5D" box="[1100,1267,1165,1187]" gridcol="4" gridrow="17" pageId="4" pageNumber="69">
1.27–1.32(
<emphasis id="DBADEA91E70D1014BA9A029BFB76FB5C" bold="true" box="[1210,1254,1165,1186]" pageId="4" pageNumber="69">1.29</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65029BFACFFB5D" box="[1349,1375,1165,1187]" gridcol="5" gridrow="17" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF02A0FACFFB32" box="[159,1375,1206,1228]" gridrow="18" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF02A0FDBCFB32" box="[159,556,1206,1228]" gridcol="0" gridrow="18" pageId="4" pageNumber="69">
w_el/w_hd 2.11–2.22(
<emphasis id="DBADEA91E70D1014BFD202A0FD8EFB35" bold="true" box="[498,542,1206,1227]" pageId="4" pageNumber="69">2.15</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB402A0FD3EFB32" box="[660,686,1206,1228]" gridcol="1" gridrow="18" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD402A0FC0BFB32" box="[756,923,1206,1228]" gridcol="2" gridrow="18" pageId="4" pageNumber="69">
2.07–2.16(
<emphasis id="DBADEA91E70D1014BD4202A0FC1DFB35" bold="true" box="[866,909,1206,1227]" pageId="4" pageNumber="69">2.11</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD02A0FB97FB32" box="[1005,1031,1206,1228]" gridcol="3" gridrow="18" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C02A0FB63FB32" box="[1100,1267,1206,1228]" gridcol="4" gridrow="18" pageId="4" pageNumber="69">
2.07–2.22(
<emphasis id="DBADEA91E70D1014BA9A02A0FB76FB35" bold="true" box="[1210,1254,1206,1227]" pageId="4" pageNumber="69">2.13</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6502A0FACFFB32" box="[1349,1375,1206,1228]" gridcol="5" gridrow="18" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF02C9FACFFB0B" box="[159,1375,1247,1269]" gridrow="19" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF02C9FDBCFB0B" box="[159,556,1247,1269]" gridcol="0" gridrow="19" pageId="4" pageNumber="69">
w_el/w_pr 1.42–1.48(
<emphasis id="DBADEA91E70D1014BFD302F6FD8FFB0B" bold="true" box="[499,543,1248,1269]" pageId="4" pageNumber="69">1.45</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB402C9FD3EFB0B" box="[660,686,1247,1269]" gridcol="1" gridrow="19" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD402C9FC0BFB0B" box="[756,923,1247,1269]" gridcol="2" gridrow="19" pageId="4" pageNumber="69">
1.41–1.48(
<emphasis id="DBADEA91E70D1014BD4202F6FC1EFB0B" bold="true" box="[866,910,1248,1269]" pageId="4" pageNumber="69">1.45</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD02C9FB97FB0B" box="[1005,1031,1247,1269]" gridcol="3" gridrow="19" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C02C9FB63FB0B" box="[1100,1267,1247,1269]" gridcol="4" gridrow="19" pageId="4" pageNumber="69">
1.41–1.48(
<emphasis id="DBADEA91E70D1014BA9A02F6FB76FB0B" bold="true" box="[1210,1254,1248,1269]" pageId="4" pageNumber="69">1.45</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6502C9FACFFB0B" box="[1349,1375,1247,1269]" gridcol="5" gridrow="19" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF031FFACFFAE1" box="[159,1375,1289,1311]" gridrow="20" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF031FFDBCFAE1" box="[159,556,1289,1311]" gridcol="0" gridrow="20" pageId="4" pageNumber="69">
(d_1p/l_el) x 100 21.1–24.4(
<emphasis id="DBADEA91E70D1014BFD3031FFD8FFAE0" bold="true" box="[499,543,1289,1310]" pageId="4" pageNumber="69">22.8</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4031FFD3EFAE1" box="[660,686,1289,1311]" gridcol="1" gridrow="20" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4031FFC0BFAE1" box="[756,923,1289,1311]" gridcol="2" gridrow="20" pageId="4" pageNumber="69">
21.6–23.9(
<emphasis id="DBADEA91E70D1014BD42031FFC1EFAE0" bold="true" box="[866,910,1289,1310]" pageId="4" pageNumber="69">22.9</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD031FFB97FAE1" box="[1005,1031,1289,1311]" gridcol="3" gridrow="20" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C031FFB63FAE1" box="[1100,1267,1289,1311]" gridcol="4" gridrow="20" pageId="4" pageNumber="69">
21.1–24.4(
<emphasis id="DBADEA91E70D1014BA9A031FFB76FAE0" bold="true" box="[1210,1254,1289,1310]" pageId="4" pageNumber="69">22.8</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65031FFACFFAE1" box="[1349,1375,1289,1311]" gridcol="5" gridrow="20" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0324FACFFAB6" box="[159,1375,1330,1352]" gridrow="21" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0324FDBCFAB6" box="[159,556,1330,1352]" gridcol="0" gridrow="21" pageId="4" pageNumber="69">
(d_2p/l_el) x100 54.0–60.0(
<emphasis id="DBADEA91E70D1014BFD30324FD8FFAB9" bold="true" box="[499,543,1330,1351]" pageId="4" pageNumber="69">56.5</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40324FD3EFAB6" box="[660,686,1330,1352]" gridcol="1" gridrow="21" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40324FC0BFAB6" box="[756,923,1330,1352]" gridcol="2" gridrow="21" pageId="4" pageNumber="69">
54.7–61.0(
<emphasis id="DBADEA91E70D1014BD420324FC1EFAB9" bold="true" box="[866,910,1330,1351]" pageId="4" pageNumber="69">58.0</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0324FB97FAB6" box="[1005,1031,1330,1352]" gridcol="3" gridrow="21" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0324FB63FAB6" box="[1100,1267,1330,1352]" gridcol="4" gridrow="21" pageId="4" pageNumber="69">
54.0–61.0(
<emphasis id="DBADEA91E70D1014BA9A0324FB76FAB9" bold="true" box="[1210,1254,1330,1351]" pageId="4" pageNumber="69">57.3</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650324FACFFAB6" box="[1349,1375,1330,1352]" gridcol="5" gridrow="21" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF034DFACFFA8C" box="[159,1375,1371,1394]" gridrow="22" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF034DFDBCFA8C" box="[159,556,1371,1394]" gridcol="0" gridrow="22" pageId="4" pageNumber="69">
(d_3p/l_el) x100 87.0–91.8(
<emphasis id="DBADEA91E70D1014BFD3034AFD8FFA8F" bold="true" box="[499,543,1372,1393]" pageId="4" pageNumber="69">88.7</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4034DFD3EFA8C" box="[660,686,1371,1394]" gridcol="1" gridrow="22" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4034DFC0BFA8C" box="[756,923,1371,1394]" gridcol="2" gridrow="22" pageId="4" pageNumber="69">
86.4–90.9(
<emphasis id="DBADEA91E70D1014BD42034AFC1EFA8F" bold="true" box="[866,910,1372,1393]" pageId="4" pageNumber="69">89.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD034DFB97FA8C" box="[1005,1031,1371,1394]" gridcol="3" gridrow="22" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C034DFB63FA8C" box="[1100,1267,1371,1394]" gridcol="4" gridrow="22" pageId="4" pageNumber="69">
86.4–91.8(
<emphasis id="DBADEA91E70D1014BA9A034AFB76FA8F" bold="true" box="[1210,1254,1372,1393]" pageId="4" pageNumber="69">88.9</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65034DFACFFA8C" box="[1349,1375,1371,1394]" gridcol="5" gridrow="22" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0393FACFFA65" box="[159,1375,1413,1435]" gridrow="23" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0393FDBCFA65" box="[159,556,1413,1435]" gridcol="0" gridrow="23" pageId="4" pageNumber="69">
(um_1/l_el) x 100 7.71–8.99(
<emphasis id="DBADEA91E70D1014BFD30393FD8FFA64" bold="true" box="[499,543,1413,1434]" pageId="4" pageNumber="69">8.23</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40393FD3EFA65" box="[660,686,1413,1435]" gridcol="1" gridrow="23" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40393FC0BFA65" box="[756,923,1413,1435]" gridcol="2" gridrow="23" pageId="4" pageNumber="69">
7.73–9.18(
<emphasis id="DBADEA91E70D1014BD420393FC1EFA64" bold="true" box="[866,910,1413,1434]" pageId="4" pageNumber="69">8.47</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0393FB97FA65" box="[1005,1031,1413,1435]" gridcol="3" gridrow="23" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0393FB63FA65" box="[1100,1267,1413,1435]" gridcol="4" gridrow="23" pageId="4" pageNumber="69">
7.71–9.18(
<emphasis id="DBADEA91E70D1014BA9A0393FB76FA64" bold="true" box="[1210,1254,1413,1434]" pageId="4" pageNumber="69">8.35</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650393FACFFA65" box="[1349,1375,1413,1435]" gridcol="5" gridrow="23" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF03B8FACFFA3A" box="[159,1375,1454,1476]" gridrow="24" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF03B8FDBCFA3A" box="[159,556,1454,1476]" gridcol="0" gridrow="24" pageId="4" pageNumber="69">
(um_2/l_el) x 100 13.2–15.7(
<emphasis id="DBADEA91E70D1014BFD303B8FD8FFA3D" bold="true" box="[499,543,1454,1475]" pageId="4" pageNumber="69">14.1</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB403B8FD3EFA3A" box="[660,686,1454,1476]" gridcol="1" gridrow="24" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD403B8FC0BFA3A" box="[756,923,1454,1476]" gridcol="2" gridrow="24" pageId="4" pageNumber="69">
13.3–15.1(
<emphasis id="DBADEA91E70D1014BD4203B8FC1EFA3D" bold="true" box="[866,910,1454,1475]" pageId="4" pageNumber="69">14.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD03B8FB97FA3A" box="[1005,1031,1454,1476]" gridcol="3" gridrow="24" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C03B8FB63FA3A" box="[1100,1267,1454,1476]" gridcol="4" gridrow="24" pageId="4" pageNumber="69">
13.2–15.7(
<emphasis id="DBADEA91E70D1014BA9A03B8FB76FA3D" bold="true" box="[1210,1254,1454,1475]" pageId="4" pageNumber="69">14.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6503B8FACFFA3A" box="[1349,1375,1454,1476]" gridcol="5" gridrow="24" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF03C1FACFFA13" box="[159,1375,1495,1517]" gridrow="25" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF03C1FDBCFA13" box="[159,556,1495,1517]" gridcol="0" gridrow="25" pageId="4" pageNumber="69">
(um_3/l_el) x 100 18.7–21.3(
<emphasis id="DBADEA91E70D1014BFD303CEFD8FFA13" bold="true" box="[499,543,1496,1517]" pageId="4" pageNumber="69">19.5</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB403C1FD3EFA13" box="[660,686,1495,1517]" gridcol="1" gridrow="25" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD403C1FC0BFA13" box="[756,923,1495,1517]" gridcol="2" gridrow="25" pageId="4" pageNumber="69">
19.0–20.8(
<emphasis id="DBADEA91E70D1014BD4203CEFC1EFA13" bold="true" box="[866,910,1496,1517]" pageId="4" pageNumber="69">20.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD03C1FB97FA13" box="[1005,1031,1495,1517]" gridcol="3" gridrow="25" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C03C1FB63FA13" box="[1100,1267,1495,1517]" gridcol="4" gridrow="25" pageId="4" pageNumber="69">
18.7–21.3(
<emphasis id="DBADEA91E70D1014BA9A03CEFB76FA13" bold="true" box="[1210,1254,1496,1517]" pageId="4" pageNumber="69">19.8</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6503C1FACFFA13" box="[1349,1375,1495,1517]" gridcol="5" gridrow="25" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0017FACFF9E9" box="[159,1375,1537,1559]" gridrow="26" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0017FDBCF9E9" box="[159,556,1537,1559]" gridcol="0" gridrow="26" pageId="4" pageNumber="69">
(um_4/l_el) x 100 24.1–27.0(
<emphasis id="DBADEA91E70D1014BFD30017FD8FF9E8" bold="true" box="[499,543,1537,1558]" pageId="4" pageNumber="69">25.1</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40017FD3EF9E9" box="[660,686,1537,1559]" gridcol="1" gridrow="26" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40017FC0BF9E9" box="[756,923,1537,1559]" gridcol="2" gridrow="26" pageId="4" pageNumber="69">
23.9–26.1(
<emphasis id="DBADEA91E70D1014BD420017FC1EF9E8" bold="true" box="[866,910,1537,1558]" pageId="4" pageNumber="69">25.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0017FB97F9E9" box="[1005,1031,1537,1559]" gridcol="3" gridrow="26" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0017FB63F9E9" box="[1100,1267,1537,1559]" gridcol="4" gridrow="26" pageId="4" pageNumber="69">
23.9–27.0(
<emphasis id="DBADEA91E70D1014BA9A0017FB76F9E8" bold="true" box="[1210,1254,1537,1558]" pageId="4" pageNumber="69">25.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650017FACFF9E9" box="[1349,1375,1537,1559]" gridcol="5" gridrow="26" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF003CFACFF9BE" box="[159,1375,1578,1600]" gridrow="27" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF003CFDBCF9BE" box="[159,556,1578,1600]" gridcol="0" gridrow="27" pageId="4" pageNumber="69">
(um_5/l_el) x 100 57.5–61.8(
<emphasis id="DBADEA91E70D1014BFD3003CFD8FF9C1" bold="true" box="[499,543,1578,1599]" pageId="4" pageNumber="69">59.3</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4003CFD3EF9BE" box="[660,686,1578,1600]" gridcol="1" gridrow="27" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4003CFC0BF9BE" box="[756,923,1578,1600]" gridcol="2" gridrow="27" pageId="4" pageNumber="69">
58.0–61.0(
<emphasis id="DBADEA91E70D1014BD42003CFC1EF9C1" bold="true" box="[866,910,1578,1599]" pageId="4" pageNumber="69">60.0</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD003CFB97F9BE" box="[1005,1031,1578,1600]" gridcol="3" gridrow="27" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C003CFB63F9BE" box="[1100,1267,1578,1600]" gridcol="4" gridrow="27" pageId="4" pageNumber="69">
57.5–61.8(
<emphasis id="DBADEA91E70D1014BA9A003CFB76F9C1" bold="true" box="[1210,1254,1578,1599]" pageId="4" pageNumber="69">59.6</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65003CFACFF9BE" box="[1349,1375,1578,1600]" gridcol="5" gridrow="27" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF0045FACFF994" box="[159,1375,1619,1642]" gridrow="28" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF0045FDBCF994" box="[159,556,1619,1642]" gridcol="0" gridrow="28" pageId="4" pageNumber="69">
(um_6/l_el) x 100 62.2–66.5(
<emphasis id="DBADEA91E70D1014BFD30042FD8FF997" bold="true" box="[499,543,1620,1641]" pageId="4" pageNumber="69">64.0</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB40045FD3EF994" box="[660,686,1619,1642]" gridcol="1" gridrow="28" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD40045FC0BF994" box="[756,923,1619,1642]" gridcol="2" gridrow="28" pageId="4" pageNumber="69">
62.4–65.5(
<emphasis id="DBADEA91E70D1014BD420042FC1EF997" bold="true" box="[866,910,1620,1641]" pageId="4" pageNumber="69">64.4</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD0045FB97F994" box="[1005,1031,1619,1642]" gridcol="3" gridrow="28" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C0045FB63F994" box="[1100,1267,1619,1642]" gridcol="4" gridrow="28" pageId="4" pageNumber="69">
62.2–66.5(
<emphasis id="DBADEA91E70D1014BA9A0042FB76F997" bold="true" box="[1210,1254,1620,1641]" pageId="4" pageNumber="69">64.2</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB650045FACFF994" box="[1349,1375,1619,1642]" gridcol="5" gridrow="28" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF006BFACFF96D" box="[159,1375,1661,1683]" gridrow="29" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF006BFDBCF96D" box="[159,556,1661,1683]" gridcol="0" gridrow="29" pageId="4" pageNumber="69">
(um_7/l_el) x 100 80.5–84.3(
<emphasis id="DBADEA91E70D1014BFD3006BFD8FF96C" bold="true" box="[499,543,1661,1682]" pageId="4" pageNumber="69">82.1</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB4006BFD3EF96D" box="[660,686,1661,1683]" gridcol="1" gridrow="29" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD4006BFC0BF96D" box="[756,923,1661,1683]" gridcol="2" gridrow="29" pageId="4" pageNumber="69">
80.7–83.6(
<emphasis id="DBADEA91E70D1014BD42006BFC1EF96C" bold="true" box="[866,910,1661,1682]" pageId="4" pageNumber="69">82.0</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD006BFB97F96D" box="[1005,1031,1661,1683]" gridcol="3" gridrow="29" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C006BFB63F96D" box="[1100,1267,1661,1683]" gridcol="4" gridrow="29" pageId="4" pageNumber="69">
80.5–84.3(
<emphasis id="DBADEA91E70D1014BA9A006BFB76F96C" bold="true" box="[1210,1254,1661,1682]" pageId="4" pageNumber="69">82.1</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB65006BFACFF96D" box="[1349,1375,1661,1683]" gridcol="5" gridrow="29" pageId="4" pageNumber="69">20</td>
</tr>
<tr id="57E934C1E70DEFEFBEBF00B0FACFF942" box="[159,1375,1702,1724]" gridrow="30" pageId="4" pageNumber="69">
<th id="14385DBDE70DEFEFBEBF00B0FDBCF942" box="[159,556,1702,1724]" gridcol="0" gridrow="30" pageId="4" pageNumber="69">
(um_8/l_el) x 100 89.2–93.3(
<emphasis id="DBADEA91E70D1014BFD300B0FD8FF945" bold="true" box="[499,543,1702,1723]" pageId="4" pageNumber="69">91.0</emphasis>
)
</th>
<td id="14385DBDE70DEFEFBCB400B0FD3EF942" box="[660,686,1702,1724]" gridcol="1" gridrow="30" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBCD400B0FC0BF942" box="[756,923,1702,1724]" gridcol="2" gridrow="30" pageId="4" pageNumber="69">
89.8–92.3(
<emphasis id="DBADEA91E70D1014BD4200B0FC1EF945" bold="true" box="[866,910,1702,1723]" pageId="4" pageNumber="69">90.7</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBDCD00B0FB97F942" box="[1005,1031,1702,1724]" gridcol="3" gridrow="30" pageId="4" pageNumber="69">10</td>
<td id="14385DBDE70DEFEFBA6C00B0FB63F942" box="[1100,1267,1702,1724]" gridcol="4" gridrow="30" pageId="4" pageNumber="69">
89.2–93.3(
<emphasis id="DBADEA91E70D1014BA9A00B0FB76F945" bold="true" box="[1210,1254,1702,1723]" pageId="4" pageNumber="69">90.9</emphasis>
)
</td>
<td id="14385DBDE70DEFEFBB6500B0FACFF942" box="[1349,1375,1702,1724]" gridcol="5" gridrow="30" pageId="4" pageNumber="69">20</td>
</tr>
</table>
</paragraph>
<paragraph id="E9663683E70D1014BEE70117FC34F8C3" blockId="4.[151,1437,1793,2033]" pageId="4" pageNumber="69">Microsculpture strongly reduced, making surface of body shining, only disc of pronotum with distinct microsculpture consisting of finely impressed transverse serrate lines.</paragraph>
<paragraph id="E9663683E70D1014BEE7015FFC9BF80F" blockId="4.[151,1437,1793,2033]" pageId="4" pageNumber="69">
Male genitalia (
<figureCitation id="71E22A06E70D1014BF58015FFE76F89F" box="[376,486,1865,1889]" captionStart="FIGURES 3 – 4" captionStartId="5.[151,264,1134,1157]" captionTargetBox="[301,1310,253,1072]" captionTargetId="figure@5.[238,1348,193,1113]" captionTargetPageId="5" captionText="FIGURES 3 – 4. Male genitalia of Pachytrechodes brevis sp. n.: a—median lobe of aedeagus; b—left paramere, c—right paramere; d—genital and last abdominal segments." httpUri="https://zenodo.org/record/283603/files/figure.png" pageId="4" pageNumber="69">Figs. 3–4</figureCitation>
). Median lobe of aedeagus rather short and straight, bent only at basal portion; basal bulb bilobed. Dorsal wall not sclerotized. Endophallus armature consisting of one large heavily sclerotized plate gradually narrowing and pointed apically and a few less distinct plates in median part. Right and left parameres differ significantly in shape, left slightly triangular, right with distinctly attenuated apical half, more typical for most trechines, each paramere bearing four apical setae.
</paragraph>
<caption id="BDA6660BE70C1015BEB70278FD0FFB5D" httpUri="https://zenodo.org/record/283603/files/figure.png" pageId="5" pageNumber="70" targetBox="[301,1310,253,1072]" targetPageId="5">
<paragraph id="E9663683E70C1015BEB70278FD0FFB5D" blockId="5.[151,1435,1134,1187]" pageId="5" pageNumber="70">
<emphasis id="DBADEA91E70C1015BEB70278FEAEFB7D" bold="true" box="[151,318,1134,1157]" pageId="5" pageNumber="70">FIGURES 3–4.</emphasis>
Male genitalia of 
<taxonomicName id="2ED94D00E70C1015BC280278FD60FB7A" box="[520,752,1134,1156]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70C1015BC280278FD60FB7A" box="[520,752,1134,1156]" italics="true" pageId="5" pageNumber="70">Pachytrechodes brevis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C09E57EAE70C1015BCDB0279FCA8FB7A" box="[763,824,1135,1156]" pageId="5" pageNumber="70" rank="species">
<emphasis id="DBADEA91E70C1015BCDB0279FCA8FB7A" bold="true" box="[763,824,1135,1156]" pageId="5" pageNumber="70">sp. n.</emphasis>
</taxonomicNameLabel>
: a—median lobe of aedeagus; b—left paramere, c—right paramere; d—genital and last abdominal segments.
</paragraph>
</caption>
<paragraph id="E9663683E70C1015BEE702D9FEEBFAD1" blockId="5.[151,1436,1231,2011]" pageId="5" pageNumber="70">Sexual dimorphism. Male and female specimens are nearly identical in morphometric characters: males differ in having slightly larger eyes, longer antennae, smaller body size, and narrower head. Quantitative differences are indicated in Table 1.</paragraph>
<paragraph id="E9663683E70C1015BEE7032DFDFFF9F9" blockId="5.[151,1436,1231,2011]" pageId="5" pageNumber="70">
<emphasis id="DBADEA91E70C1015BEE7032DFEDAFAAA" bold="true" box="[199,330,1339,1364]" pageId="5" pageNumber="70">Bionomics.</emphasis>
The specimens were collected by searching in moist leaf litter on the forest floor of a montane tropical forest at an altitude of ca. 
<quantity id="2E219B66E70C1015BC630349FD0EFA89" box="[579,670,1375,1399]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="70" unit="m" value="1500.0">1500 m</quantity>
above sea level, Udzungwa Mountains National Park (formerly Mwanihana Forest Reserve). The vegetation of the area was dominated by tall trees (
<emphasis id="DBADEA91E70C1015BA500393FB59FA62" box="[1136,1225,1413,1436]" italics="true" pageId="5" pageNumber="70">Dryptes</emphasis>
sp., 
<taxonomicName id="2ED94D00E70C1015BB220393FA05FA62" box="[1282,1429,1412,1436]" class="Magnoliopsida" family="Rutaceae" genus="Vepris" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="70" phylum="Tracheophyta" rank="species" species="stolzii">
<emphasis id="DBADEA91E70C1015BB220393FA05FA62" box="[1282,1429,1412,1436]" italics="true" pageId="5" pageNumber="70">Vepris stolzii</emphasis>
</taxonomicName>
, 
<taxonomicName id="2ED94D00E70C1015BEB703BEFEE6FA41" box="[151,374,1448,1471]" class="Magnoliopsida" family="Myrtaceae" genus="Syzigium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="70" phylum="Magnoliophyta" rank="species" species="guineense">
<emphasis id="DBADEA91E70C1015BEB703BEFEE6FA41" box="[151,374,1448,1471]" italics="true" pageId="5" pageNumber="70">Syzigium guineense</emphasis>
</taxonomicName>
, and 
<taxonomicName id="2ED94D00E70C1015BF9803BEFDE4FA41" box="[440,628,1447,1471]" class="Magnoliopsida" family="Chrysobalanaceae" genus="Parinari" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="70" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="DBADEA91E70C1015BF9803BEFDE4FA41" box="[440,628,1447,1471]" italics="true" pageId="5" pageNumber="70">Parinari excelsa</emphasis>
</taxonomicName>
). Tree ferns (
<taxonomicName id="2ED94D00E70C1015BD3403B1FC4EFA41" box="[788,990,1447,1471]" class="Liliopsida" family="Arecaceae" genus="Phoenix" kingdom="Plantae" order="Arecales" pageId="5" pageNumber="70" phylum="Tracheophyta" rank="species" species="reclinata">
<emphasis id="DBADEA91E70C1015BD3403B1FC4EFA41" box="[788,990,1447,1471]" italics="true" pageId="5" pageNumber="70">Phoenix reclinata</emphasis>
</taxonomicName>
) were also common. The beetles were active from dusk to midnight under trees where there was fruit windfall. During the day they hid in the forest litter and could be collected by sifting the litter.
</paragraph>
<paragraph id="E9663683E70C1016BEE70005FEFCFEE2" blockId="5.[151,1436,1231,2011]" lastBlockId="6.[151,1436,151,464]" lastPageId="6" lastPageNumber="71" pageId="5" pageNumber="70">
<emphasis id="DBADEA91E70C1015BEE70005FE21F9D2" bold="true" box="[199,433,1555,1580]" pageId="5" pageNumber="70">Comparative notes.</emphasis>
The new species agrees well with the three other known congeners both in external and genitalic characters (Jeannel 1960; Uéno 1987). All four species of 
<taxonomicName id="2ED94D00E70C1015BDB80021FBDAF9B1" box="[920,1098,1591,1615]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70C1015BDB80021FBDAF9B1" box="[920,1098,1591,1615]" italics="true" pageId="5" pageNumber="70">Pachytrechodes</emphasis>
</taxonomicName>
share the following character states: small and weakly produced eyes which are not significantly longer than convex tempora; small and robust body shape with relatively short legs and brief ovate elytra; microsculpture of dorsum more or less reduced, and inner elytral striae well impressed. The structure of the male genitalia is also very similar including both the shape of the aedeagus and the presence of the well-sclerotized copulatory pieces. In its small size (it is the smallest species of the genus) and a narrower pronotum, 
<taxonomicName id="2ED94D00E70C1015BCEC00FAFC5CF8FA" box="[716,972,1772,1796]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70C1015BCEC00FAFC5CF8FA" box="[716,972,1772,1796]" italics="true" pageId="5" pageNumber="70">Pachytrechodes brevis</emphasis>
</taxonomicName>
<emphasis id="DBADEA91E70C1015BDF600FAFB89F8FA" bold="true" box="[982,1049,1772,1796]" pageId="5" pageNumber="70">
<taxonomicNameLabel id="C09E57EAE70C1015BDF600FAFB89F8FA" box="[982,1049,1772,1796]" pageId="5" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is more similar to 
<taxonomicName id="2ED94D00E70C1015BADC00FBFEA1F8D9" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="DBADEA91E70C1015BADC00FBFA0CF8FA" box="[1276,1436,1772,1796]" italics="true" pageId="5" pageNumber="70">P. basilewskyi</emphasis>
Jeannel, 1960
</taxonomicName>
and 
<taxonomicName id="2ED94D00E70C1015BF490106FDCCF8D9" authority="Ueno, 1987" authorityName="Ueno" authorityYear="1987" box="[361,604,1807,1831]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="DBADEA91E70C1015BF490106FE41F8D9" box="[361,465,1807,1831]" italics="true" pageId="5" pageNumber="70">P. leleupi</emphasis>
Uéno, 1987
</taxonomicName>
than to 
<emphasis id="DBADEA91E70C1015BC990106FCAEF8D9" box="[697,830,1808,1831]" italics="true" pageId="5" pageNumber="70">P. u n c i n a t u s</emphasis>
Uéno, 1987; however, 
<emphasis id="DBADEA91E70C1015BA6A0106FB37F8D9" box="[1098,1191,1807,1831]" italics="true" pageId="5" pageNumber="70">P. b re vi s</emphasis>
shares with the latter species the shape of the aedeagal median lobe with apex distinctly attenuated in lateral view and the presence of a large acuminate copulatory piece within the endophallus. Additionally, 
<taxonomicName id="2ED94D00E70C1015BDF90141FB49F891" box="[985,1241,1879,1903]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="brevis">
<emphasis id="DBADEA91E70C1015BDF90141FB49F891" box="[985,1241,1879,1903]" italics="true" pageId="5" pageNumber="70">Pachytrechodes brevis</emphasis>
</taxonomicName>
differs from the other species of the genus in having clearly protruding eyes (versus very flat in 
<taxonomicName id="2ED94D00E70C1015BA09016BFB24F86A" box="[1065,1204,1917,1940]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="uncinatus">
<emphasis id="DBADEA91E70C1015BA09016BFB24F86A" box="[1065,1204,1917,1940]" italics="true" pageId="5" pageNumber="70">P. uncinatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="2ED94D00E70C1015BACD016BFA1BF86A" box="[1261,1419,1916,1940]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="DBADEA91E70C1015BACD016BFA1BF86A" box="[1261,1419,1916,1940]" italics="true" pageId="5" pageNumber="70">P. basilewskyi</emphasis>
</taxonomicName>
), combined with the apex of the aedeagus attenuated in lateral view (versus very blunt, not produced in 
<taxonomicName id="2ED94D00E70C1015BB0301B6FA1BF849" box="[1315,1419,1951,1975]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="DBADEA91E70C1015BB0301B6FA1BF849" box="[1315,1419,1951,1975]" italics="true" pageId="5" pageNumber="70">P. leleupi</emphasis>
</taxonomicName>
), and some other characters of minor importance. The shape of the aedeagus in dorsal view is intermediate between that of 
<emphasis id="DBADEA91E70F1016BEC7068FFEFDFF4E" box="[231,365,153,176]" italics="true" pageId="6" pageNumber="71">P. u nc i na tu s</emphasis>
and 
<emphasis id="DBADEA91E70F1016BF88068FFD87FF4E" box="[424,535,152,176]" italics="true" pageId="6" pageNumber="71">
<taxonomicName id="2ED94D00E70F1016BF88068FFD84FF4E" box="[424,532,152,176]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="leleupi">P. leleupi</taxonomicName>
.
</emphasis>
The large sclerotized plate in the endophallus of the new species does not seem to be homologous with that of 
<taxonomicName id="2ED94D00E70F1016BFD106ABFDEAFF2A" box="[497,634,189,212]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="uncinatus">
<emphasis id="DBADEA91E70F1016BFD106ABFDEAFF2A" box="[497,634,189,212]" italics="true" pageId="6" pageNumber="71">P. uncinatus</emphasis>
</taxonomicName>
, as this plate is much longer in the new species and is located closer to the basal part of the aedeagus (this difference is even more noticeable when considering the position of the other copulatory pieces).
</paragraph>
</subSubSection>
<subSubSection id="A1C36508E70F1016BEE70731FD58FE31" pageId="6" pageNumber="71" type="distribution">
<paragraph id="E9663683E70F1016BEE70731FD58FE31" blockId="6.[151,1436,151,464]" pageId="6" pageNumber="71">
<emphasis id="DBADEA91E70F1016BEE70731FEC9FEBE" bold="true" box="[199,345,295,320]" pageId="6" pageNumber="71">Distribution</emphasis>
. 
<taxonomicName id="2ED94D00E70F1016BF4B073EFDFAFEBE" box="[363,618,296,320]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70F1016BF4B073EFDFAFEBE" box="[363,618,296,320]" italics="true" pageId="6" pageNumber="71">Pachytrechodes brevis</emphasis>
</taxonomicName>
<emphasis id="DBADEA91E70F1016BC55073EFD29FEBE" bold="true" box="[629,697,296,320]" pageId="6" pageNumber="71">
<taxonomicNameLabel id="C09E57EAE70F1016BC55073EFD29FEBE" box="[629,697,296,320]" pageId="6" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is the first member of the genus found outside of the Uluguru Mountains, where all the previously described species were found (Jeannel 1960, Uéno 1987). The 
<typeStatus id="36628821E70F1016BAD2075BFAB2FE9B" box="[1266,1314,333,357]" pageId="6" pageNumber="71">type</typeStatus>
locality of the new species is situated in an eastern part of the Udzungwa Mountains, approximately 
<quantity id="2E219B66E70F1016BAA80779FB4CFE79" box="[1160,1244,367,391]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="71" unit="km" value="110.0">110 km</quantity>
southwest of the 
<typeStatus id="36628821E70F1016BEB70783FF57FE53" box="[151,199,405,429]" pageId="6" pageNumber="71">type</typeStatus>
localities of 
<taxonomicName id="2ED94D00E70F1016BF440783FE61FE52" box="[356,497,405,428]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="uncinatus">
<emphasis id="DBADEA91E70F1016BF440783FE61FE52" box="[356,497,405,428]" italics="true" pageId="6" pageNumber="71">P. uncinatus</emphasis>
</taxonomicName>
and 
<emphasis id="DBADEA91E70F1016BC100783FD08FE52" box="[560,664,404,428]" italics="true" pageId="6" pageNumber="71">P. l e l e u p i</emphasis>
(
<figureCitation id="71E22A06E70F1016BC8E0782FD67FE52" box="[686,759,404,428]" captionStart="FIGURE 5" captionStartId="6.[151,250,1342,1365]" captionText="FIGURE 5. Distribution of Pachytrechodes species." pageId="6" pageNumber="71">Fig. 5</figureCitation>
). The Uluguru and Udzungwa mountains are two distinct neighboring blocks of the Eastern Arc Mountains.
</paragraph>
</subSubSection>
<caption id="BDA6660BE70F1016BEB70328FD3CFAAB" box="[151,684,1342,1365]" pageId="6" pageNumber="71">
<paragraph id="E9663683E70F1016BEB70328FD3CFAAB" blockId="6.[151,684,1342,1365]" box="[151,684,1342,1365]" pageId="6" pageNumber="71">
<emphasis id="DBADEA91E70F1016BEB70328FE9CFAAA" bold="true" box="[151,268,1342,1365]" pageId="6" pageNumber="71">FIGURE 5</emphasis>
. Distribution of 
<taxonomicName id="2ED94D00E70F1016BF950329FDC6FAAB" box="[437,598,1343,1365]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70F1016BF950329FDC6FAAB" box="[437,598,1343,1365]" italics="true" pageId="6" pageNumber="71">Pachytrechodes</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<subSubSection id="A1C36508E70F1017BEB703B0FC26F910" lastPageId="7" lastPageNumber="72" pageId="6" pageNumber="71" type="key">
<paragraph id="E9663683E70F1016BEB703B0FCF0FA40" blockId="6.[151,864,1445,1470]" box="[151,864,1445,1470]" pageId="6" pageNumber="71">
<emphasis id="DBADEA91E70F1016BEB703B0FD8AFA40" bold="true" box="[151,538,1445,1470]" pageId="6" pageNumber="71">
Key to species of 
<taxonomicName id="2ED94D00E70F1016BF4403B3FD8AFA40" ID-CoL="8KY6Z" box="[356,538,1445,1470]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70F1016BF4403B3FD8AFA40" bold="true" box="[356,538,1445,1470]" italics="true" pageId="6" pageNumber="71">Pachytrechodes</emphasis>
</taxonomicName>
</emphasis>
(modified from Uéno, 1987)
</paragraph>
<keyStep id="522D2E26E70F1016BEB703FBFA0CF94C" pageId="6" pageNumber="71">
<paragraph id="E9663683E70F1016BEB703FBFA0CF9C3" blockId="6.[151,1436,1517,2037]" pageId="6" pageNumber="71">
<keyLead id="522895B6E70F1016BEB703FBFA0CF9C3" pageId="6" pageNumber="71">
<key id="9D4861B2E70F1016BEB703FBFA0CF9C3" pageId="6" pageNumber="71">
1 Smaller species, body length less than 
<quantity id="2E219B66E70F1016BC7903FBFD22F9FC" box="[601,690,1517,1538]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="71" unit="mm" value="2.5">2.50 mm</quantity>
; elytra smaller, ratio maximum elytral width / maximum pronotal width normally lower than 1.48; aedeagus smaller, not gutter-shaped; endophallus armature without large uncinate copulatory piece near apical orifice......................................................................................... 2
</key>
</keyLead>
</paragraph>
<paragraph id="E9663683E70F1016BEB70053FA0CF94C" blockId="6.[151,1436,1517,2037]" pageId="6" pageNumber="71">
<keyLead id="522895B6E70F1016BEB70053FA0CF94C" pageId="6" pageNumber="71">
- Larger species, body length 
<quantity id="2E219B66E70F1016BFD40053FDDAF9A4" box="[500,586,1605,1626]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="71" unit="mm" value="2.7">2.70 mm</quantity>
or more; elytra larger, ratio maximum elytral width / maximum pronotal width usually higher than 1.50; aedeagus larger and heavily sclerotized, gutter-shaped, nearly enveloping endophallus which bears a large uncinate copulatory piece near apical orifice. Southern part of the Uluguru Mts: Lukwangule Plateau..................................................................................................... 
<taxonomicName id="2ED94D00E70F1016BA8C008BFA0CF94C" ID-CoL="4BVTB" authority="Ueno, 1987" authorityName="Ueno" authorityYear="1987" box="[1196,1436,1693,1714]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="uncinatus">
<emphasis id="DBADEA91E70F1016BA8C008BFAB6F94C" bold="true" box="[1196,1318,1693,1714]" italics="true" pageId="6" pageNumber="71">P. uncinatus</emphasis>
Uéno, 1987
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="522D2E26E70F1016BEB700ADFA0CF881" pageId="6" pageNumber="71">
<paragraph id="E9663683E70F1016BEB700ADFA0CF8F4" blockId="6.[151,1436,1517,2037]" pageId="6" pageNumber="71">
<keyLead id="522895B6E70F1016BEB700ADFA0CF8F4" pageId="6" pageNumber="71">
2 Body length 
<quantity id="2E219B66E70F1016BF4400ADFE66F92E" box="[356,502,1723,1744]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.45" metricValueMin="2.15" pageId="6" pageNumber="71" unit="mm" value="2.3" valueMax="2.45" valueMin="2.15">2.15–2.45 mm</quantity>
. Pronotum less transverse (on average ratio width/length 1.40). Elytra less wide, with more oblique humeri. Median lobe of aedeagus rather poorly sclerotized except for the ventral part, the apex not attenuated in profile, blunt or pointed. Endophallus armature without large acuminate copulatory piece in proximal part of aedeagus........ 3
</keyLead>
</paragraph>
<paragraph id="E9663683E70F1016BEB70105FA0CF881" blockId="6.[151,1436,1517,2037]" pageId="6" pageNumber="71">
<keyLead id="522895B6E70F1016BEB70105FA0CF881" pageId="6" pageNumber="71">
- The smallest species of the genus, body length 2.0– 
<quantity id="2E219B66E70F1016BCF30105FCB0F8D6" box="[723,800,1811,1832]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="71" unit="mm" value="2.2">2.2 mm</quantity>
. Pronotum more transverse (on average, ratio width/length 1.45), its shape approximately as in 
<emphasis id="DBADEA91E70F1016BC250126FD12F8BB" box="[517,642,1840,1861]" italics="true" pageId="6" pageNumber="71">
<taxonomicName id="2ED94D00E70F1016BC250126FDEEF8BB" ID-CoL="4BVTB" box="[517,638,1840,1861]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="uncinatus">P. uncinatus</taxonomicName>
.
</emphasis>
Elytra wider, with more distinct humeri. Median lobe of aedeagus more strongly sclerotized, its apex distinctly attenuated in profile. Endophallus armature with a large acuminate copulatory piece in proximal part of aedeagus (
<figureCitation id="71E22A06E70F1016BFAB017DFE78F87E" box="[395,488,1899,1920]" captionStart="FIGURES 3 – 4" captionStartId="5.[151,264,1134,1157]" captionTargetBox="[301,1310,253,1072]" captionTargetId="figure@5.[238,1348,193,1113]" captionTargetPageId="5" captionText="FIGURES 3 – 4. Male genitalia of Pachytrechodes brevis sp. n.: a—median lobe of aedeagus; b—left paramere, c—right paramere; d—genital and last abdominal segments." httpUri="https://zenodo.org/record/283603/files/figure.png" pageId="6" pageNumber="71">Figs. 3–4</figureCitation>
). Eastern part of the Udzungwa Mts..................................... 
<emphasis id="DBADEA91E70F1016BB2D017DFA0CF881" bold="true" box="[1293,1436,1898,1920]" pageId="6" pageNumber="71">
<taxonomicName id="2ED94D00E70F1016BB2D017DFAF0F87E" ID-CoL="4BVT9" box="[1293,1376,1899,1920]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70F1016BB2D017DFAF0F87E" bold="true" box="[1293,1376,1899,1920]" italics="true" pageId="6" pageNumber="71">P. brevis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C09E57EAE70F1016BB46017CFA0CF881" box="[1382,1436,1898,1919]" pageId="6" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="E9663683E70F1016BEB7019EFA0CF826" blockId="6.[151,1436,1517,2037]" pageId="6" pageNumber="71">
<keyLead id="522895B6E70F1016BEB7019EFA0CF826" pageId="6" pageNumber="71">
<keyStep id="522D2E26E70F1016BEB7019EFA0CF826" pageId="6" pageNumber="71">
3 Eyes flat; elytral striae shallower, stria 1 abbreviated near base; aedeagal apical lobe narrow and slightly reflexed in profile; endophallus armed with a single claw-like copulatory piece near apical orifice; Central part of the Uluguru Mts: Kidunda Mt............................................................................... 
<taxonomicName id="2ED94D00E70F1016BAAB01D5FA0CF826" ID-CoL="4BVT8" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[1163,1436,1987,2008]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="71" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="DBADEA91E70F1016BAAB01D5FA82F826" bold="true" box="[1163,1298,1987,2008]" italics="true" pageId="6" pageNumber="71">P. basilewskyi</emphasis>
Jeannel, 1960
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="E9663683E70F1017BEB701F6FE85FB2D" blockId="6.[151,1436,1517,2037]" lastBlockId="7.[151,277,1210,1235]" lastPageId="7" lastPageNumber="72" pageId="6" pageNumber="71">
<keyLead id="522895B6E70F1016BEB701F6FA0CF80B" box="[151,1436,2016,2037]" pageId="6" pageNumber="71">- Eye slightly convex; elytral striae deeper, stria 1 usually traceable to base; median lobe of aedeagus very short and more or less</keyLead>
blunt in profile; endophallus armed with at least two copulatory pieces lying one after the other at the median part. Southern part of the Uluguru Mts: ca. Chenzema Village............................................. 
<taxonomicName id="2ED94D00E70E1017BAEC06A3FA0CFF34" ID-CoL="75J9D" authority="Ueno, 1987" authorityName="Ueno" authorityYear="1987" box="[1228,1436,181,202]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis id="DBADEA91E70E1017BAEC06A3FAB6FF34" bold="true" box="[1228,1318,181,202]" italics="true" pageId="7" pageNumber="72">P. leleupi</emphasis>
Uéno, 1987
</taxonomicName>
<emphasis id="DBADEA91E70E1017BEB702ACFE85FB2D" bold="true" box="[151,277,1210,1235]" pageId="7" pageNumber="72">Discussion</emphasis>
</paragraph>
<caption id="BDA6660BE70E1017BEB70245FD6CFB97" box="[151,764,1107,1130]" httpUri="https://zenodo.org/record/283604/files/figure.png" pageId="7" pageNumber="72" targetBox="[238,1348,254,1086]" targetPageId="7">
<paragraph id="E9663683E70E1017BEB70245FD6CFB97" blockId="7.[151,764,1107,1130]" box="[151,764,1107,1130]" pageId="7" pageNumber="72">
<emphasis id="DBADEA91E70E1017BEB70245FE82FB97" bold="true" box="[151,274,1107,1130]" pageId="7" pageNumber="72">FIGURE 6.</emphasis>
Typical biotope of 
<taxonomicName id="2ED94D00E70E1017BFF80242FD2CFB94" ID-CoL="4BVT9" box="[472,700,1108,1130]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
<emphasis id="DBADEA91E70E1017BFF80242FD2CFB94" box="[472,700,1108,1130]" italics="true" pageId="7" pageNumber="72">Pachytrechodes brevis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C09E57EAE70E1017BCE30242FD6CFB97" box="[707,764,1108,1129]" pageId="7" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E9663683E70E1017BEB70314FC26F910" blockId="7.[151,1437,1282,1774]" pageId="7" pageNumber="72">
Members of the subtribe Trechodina are mostly ripicolous. So far, only a very few humicolous trechodines are known. Apart from 
<taxonomicName id="2ED94D00E70E1017BFA10330FDA3FAC0" ID-CoL="8KY6Z" box="[385,563,1318,1342]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70E1017BFA10330FDA3FAC0" box="[385,563,1318,1342]" italics="true" pageId="7" pageNumber="72">Pachytrechodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="2ED94D00E70E1017BC640330FC67FAC0" ID-CoL="6M96K" authority="Ueno, 1981" authorityName="Ueno" authorityYear="1981" box="[580,1015,1318,1342]" class="Insecta" family="Carabidae" genus="Himalotrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="DBADEA91E70E1017BC640330FCF6FAC0" box="[580,870,1318,1342]" italics="true" pageId="7" pageNumber="72">Himalotrechodes insignis</emphasis>
Uéno, 1981
</taxonomicName>
and 
<taxonomicName id="2ED94D00E70E1017BA150330FF42FA9C" ID-CoL="55ZR3" authority="Jeannel, 1938" authorityName="Jeannel" authorityYear="1938" class="Insecta" family="Carabidae" genus="Thalassophilus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="DBADEA91E70E1017BA150330FAA6FAC1" box="[1077,1334,1318,1343]" italics="true" pageId="7" pageNumber="72">Thalassophilus caecus</emphasis>
Jeannel, 1938
</taxonomicName>
, which were all discussed by Uéno (1987), there are three more species of the humicolous genus 
<taxonomicName id="2ED94D00E70E1017BEB70378FE67FA78" ID-CoL="8KYT5" authority="Geginat, 2006" authorityName="Geginat" authorityYear="2006" box="[151,503,1390,1414]" class="Insecta" family="Carabidae" genus="Scaurotrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70E1017BEB70378FEC1FA78" box="[151,337,1390,1414]" italics="true" pageId="7" pageNumber="72">Scaurotrechodes</emphasis>
Geginat, 2006
</taxonomicName>
described from the South African Cape region (Geginat 2006, 2011). Surprisingly, the only known member of the humicolous genus 
<taxonomicName id="2ED94D00E70E1017BCCD0385FBD2FA54" ID-CoL="8KX8D" authority="Ueno, 1981" authorityName="Ueno" authorityYear="1981" box="[749,1090,1426,1451]" class="Insecta" family="Carabidae" genus="Himalotrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70E1017BCCD0385FC3DFA55" box="[749,941,1427,1451]" italics="true" pageId="7" pageNumber="72">Himalotrechodes</emphasis>
Uéno, 1981
</taxonomicName>
, from the eastern Himalayas, demonstrates much more affinities with African 
<taxonomicName id="2ED94D00E70E1017BCEF03A0FB8EFA30" ID-CoL="8KY6Z" authority="Ueno 1981" authorityName="Ueno" authorityYear="1981" box="[719,1054,1462,1486]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70E1017BCEF03A0FC11FA30" box="[719,897,1462,1486]" italics="true" pageId="7" pageNumber="72">Pachytrechodes</emphasis>
(Uéno 1981)
</taxonomicName>
than the latter with members of 
<emphasis id="DBADEA91E70E1017BEB703CDFEC9FA0D" box="[151,345,1499,1523]" italics="true" pageId="7" pageNumber="72">
<taxonomicName id="2ED94D00E70E1017BEB703CDFEC5FA0D" ID-CoL="8KYT5" box="[151,341,1499,1523]" class="Insecta" family="Carabidae" genus="Scaurotrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">Scaurotrechodes</taxonomicName>
.
</emphasis>
This similarity is related to the reduction of hind wings, small eyes, swollen tempora, less strongly impressed frontal furrows etc., characters which may have evolved independently in various taxa when originally ripicolous lineages adapt to the humicolous microhabitats of the forest floor. On the other hand, the four species of 
<taxonomicName id="2ED94D00E70E1017BF2C0050FE2EF9A0" ID-CoL="8KY6Z" box="[268,446,1606,1630]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70E1017BF2C0050FE2EF9A0" box="[268,446,1606,1630]" italics="true" pageId="7" pageNumber="72">Pachytrechodes</emphasis>
</taxonomicName>
, in addition to the above set of adaptive characters, share a similar shape of the aedeagus and the structure of the copulatory pieces, what suggests a monophyletic relationship between them, a hypothesis also supported by their geographic distribution restricted to the neighboring areas of the Uluguru and Udzungwa Mountains (
<figureCitation id="71E22A06E70E1017BF0200A4FEFAF934" box="[290,362,1714,1738]" captionStart="FIGURE 5" captionStartId="6.[151,250,1342,1365]" captionText="FIGURE 5. Distribution of Pachytrechodes species." pageId="7" pageNumber="72">Fig. 5</figureCitation>
). Since the latter is the largest block of the Eastern Arc Mountains, it is expected that new species of 
<taxonomicName id="2ED94D00E70E1017BF2D00C0FE2FF910" ID-CoL="8KY6Z" box="[269,447,1750,1774]" class="Insecta" family="Carabidae" genus="Pachytrechodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="DBADEA91E70E1017BF2D00C0FE2FF910" box="[269,447,1750,1774]" italics="true" pageId="7" pageNumber="72">Pachytrechodes</emphasis>
</taxonomicName>
will be therein discovered in the next future.
</paragraph>
</subSubSection>
</treatment>
</document>