treatments-xml/data/52/D0/30/52D0301278D5A22C9017D5EE848E90DF.xml

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<mods:title id="E7276EB9712B6AE356CDC208482EF300">Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<paragraph id="12F925DD9B32B1F9BBB040EE7827C5D4" pageId="13" pageNumber="130">
<taxonomicName id="32F55C6CF6ED6039FCE05490FEBC8D9E" ID-CoL="4R35Q" authority="Luze, 1904" authorityName="Luze" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) imitator" order="Coleoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="species" species="imitator" subGenus="Raphirus">Quedius (Raphirus) imitator Luze, 1904</taxonomicName>
Figs 3C, 15
</paragraph>
</subSubSection>
<subSubSection id="41706FA3FA105A9464146258BA9CDB54" pageId="14" pageNumber="131" type="reference_group">
<paragraph id="70F31F0FB0EF798E6F8C5B3035C5B344" pageId="14" pageNumber="131">
<taxonomicName id="972E4CECBCE6216A609E51072A6C1A96" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius tschinganensis" order="Coleoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="species" species="tschinganensis">
<pageBreakToken id="8FD6C08A4DCE7A6332A7B7DF42336D66" pageId="14" pageNumber="131" start="start">Quedius</pageBreakToken>
tschinganensis
</taxonomicName>
Coiffait, 1969, syn. n. (Fig. 14)
</paragraph>
<paragraph id="7103BCB67AB8F2D0BCD2B012C77A33B8" pageId="14" pageNumber="131">
<taxonomicName id="71231C051E3D00C0F88364FAFEA87E7F" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius imitator" order="Coleoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="species" species="imitator">Quedius imitator</taxonomicName>
Luze, 1904, 102 (original description); 
<bibRefCitation id="451377012052C6D5B1A9B7B13E93DBC7" author="Bernhauer, M" journalOrPublisher="zoologisch-botanischen Gesellschaft in Wien" pageId="25" pageNumber="142" pagination="580 - 596" title="13. Folge neuer Staphyliniden der palaearktischen Fauna, nebst Bemerkungen. Verhandlungen der k. k." url="http://www.zobodat.at/pdf/VZBG_55_0580-0596.pdf" volume="55" year="1905">Bernhauer 1905</bibRefCitation>
, 596 (notes); 
<bibRefCitation id="45F40D13DD67C87A9EAB10BBBC8FADF2" pageId="14" pageNumber="131">Bernhauer and Schubert 1916</bibRefCitation>
, 429 (list with synonyms); 
<bibRefCitation id="9EF7E168E940467BA3F3B92124849D14" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli 1924</bibRefCitation>
, 135 (characters, notes); 
<bibRefCitation id="5A16AE091EEAC62BBA017449A537432D" author="Coiffait, H" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="25" pageNumber="142" pagination="391 - 424" title="Quedius nouveaux ou mal connus." volume="103" year="1967">Coiffait 1967</bibRefCitation>
, 406 (characters); 
<bibRefCitation id="2064AF92EBF3B6946B1EE31441F8D269" pageId="14" pageNumber="131">Coiffait 1978</bibRefCitation>
, 237 (characters, distribution records); 
<bibRefCitation id="83B056F5945FF76AC5AE5AA05872CF40" author="Bohac, J" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="25" pageNumber="142" pagination="549 - 558" title="Novye i maloizvestnye zhuki-stafilinidy podsemeistva Staphylininae (Coleoptera, Staphylinidae)." volume="67" year="1988">
<normalizedToken id="BC5774AC865CA677B3AC38146DDD0D16" originalValue="Boháč">Bohac</normalizedToken>
1988
</bibRefCitation>
, 556 (distribution records); 
<bibRefCitation id="3DD02379A2844889D4B227C96AD1BF8D" pageId="14" pageNumber="131">Klimenko 1996</bibRefCitation>
, 121;
</paragraph>
<paragraph id="16903DFB43C8CAEE6F64F86F0F5B17AB" pageId="14" pageNumber="131">
<taxonomicName id="6145AE7E9346FC3B70FA6382265DEEA0" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius tschinganensis" order="Coleoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="species" species="tschinganensis">Quedius tschinganensis</taxonomicName>
Coiffait, 1969, 50 (original description); 
<bibRefCitation id="62476EAE1BFA0B9C46349AE737F6E9CF" author="Coiffait, H" journalOrPublisher="L'Entomologiste" pageId="25" pageNumber="142" pagination="142 - 148" title="Staphylinidae d'Ouzbekistan." volume="26" year="1970">Coiffait 1970</bibRefCitation>
, 143 (list); 
<bibRefCitation id="07883A68579DC14EBF18929A3F27750B" pageId="14" pageNumber="131">Coiffait 1978</bibRefCitation>
, 237 (characters); Kascheev 2001, 102 (distribution records);
</paragraph>
<paragraph id="AC256565F52918841A82D05ECED573EB" pageId="14" pageNumber="131">
<taxonomicName id="FF7B23824931D73B0496D33D948A8B27" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Quedius tschinganensis var. gracilicornis" order="Coleoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="variety" species="tschinganensis" variety="gracilicornis">Quedius tschinganensis var. gracilicornis</taxonomicName>
Coiffait, 1977, 139 (original description);
</paragraph>
<paragraph id="1FEB6C0226ECBE6E55D2AA84148A5D48" pageId="14" pageNumber="131">
<taxonomicName id="4CA1CA1F307DB1E6B93E9A7E3E7BBC7D" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Quedius tschinganensis var. debilicornis" order="Coleoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="variety" species="tschinganensis" variety="debilicornis">Quedius tschinganensis var. debilicornis</taxonomicName>
Coiffait, 1978, 237 (replacement name for 
<taxonomicName id="C4097BA15344EAEA5803C09C455AEBD3" family="Staphylinidae" genus="Amblyopinina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amblyopinina gracilicornis" order="Coleoptera" pageId="14" pageNumber="131" rank="species" species="gracilicornis">gracilicornis</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="B51D6FE055670636AAB036E009B9E112" pageId="14" pageNumber="131" type="type material">
<paragraph id="3647FCA12BFEE48AA2F2A421328FEB64" pageId="14" pageNumber="131">Type material examined.</paragraph>
<paragraph id="476DEB20190CB6CDB1AE391357325C64" pageId="14" pageNumber="131">
Quedius imitator: Tajikistan or Uzbekistan: Lectotype (here designated): 1 ♂, &quot;Seravschan Darch Glasunov 1892 [printed]/ Q. imitator Luze J. 
<normalizedToken id="8BA69C24F967ECB4939995F2FD4B8331" originalValue="Boháč">Bohac</normalizedToken>
det. 1983 [pre-printed]&quot;; paralectotypes: 1 ♂, &quot;[square orange piece of paper]/ Seravschan Putchin Pass. Glasunov, 1892 [printed]/ Quedius imitator Luze [handwritten]/ Q. imitator Luze J. 
<normalizedToken id="73BFA9B6AC5558A8AF373F10AE3BD657" originalValue="Boháč">Bohac</normalizedToken>
det. 1983 [pre-printed]&quot;; paralectotypes: 1 ♂, &quot;Seravschan Putchin Pass. Glasunov, 1892 [printed]/ Quedius imitator Luze [handwritten]/ Q. imitator Luze J. 
<normalizedToken id="2C01439BD927977E9E92A58775A9B157" originalValue="Boháč">Bohac</normalizedToken>
det. 1983 [pre-printed]/ Quedius sp.1. cf. suturalis Ksw. A. Solodovnikov 1997 [handwritten]&quot;; 3 ♂, &quot;Seravschan Putchin Pass. Glasunov 1892 [printed]/ Q. imitator Luze J. 
<normalizedToken id="A04E008929440B79C1A6CBA79A097660" originalValue="Boháč">Bohac</normalizedToken>
det. 1983 [pre-printed]/ Quedius sp.1. cf. suturalis Ksw. A. Solodovnikov 1997 [handwritten]&quot;; 2 ♀, &quot;Seravschan Putchin Pass. Glasunov 1892[printed]&quot;; 2 ♂, &quot;Seravschan Obburden Glasunov, 1892 [printed]/ Q. imitator Luze J. 
<normalizedToken id="1FD4EF6C5D50996C4ED434359514AD91" originalValue="Boháč">Bohac</normalizedToken>
det. 1983 [pre-printed]/ Quedius sp.1. cf. suturalis Ksw. A. Solodovnikov 1997 [handwritten]&quot;; 1 ♀, &quot;Iskander-Kul Iskander-Darja Glasunov 1892 [printed]/ Q. imitator Luze J. 
<normalizedToken id="A9D8680F085718DA89FD7CA2B838CCBD" originalValue="Boháč">Bohac</normalizedToken>
det. 1983 [pre-printed]&quot; (ZIN).
</paragraph>
<paragraph id="0587DBD9A4E59D6F8D565E0D38B07E2E" pageId="14" pageNumber="131">Quedius tschinganensis: Uzbekistan: Holotype: ♂, &quot;Ouzbekistan 8-68 Mts Tschingan 1500 m. H.C. [printed]/ Q. (Sauridus) tschinganus [sic!] Coiff. H. Coiffait det. 1968 [pre-printed]/ Holotype [printed]&quot;; 5 ♂, 1 ♀, &quot;Ouzbekistan 8-68 Mts Tschingan 1500 m. H.C. [printed]/ Paralectotype [printed]&quot; (Fig. 14D, H) (MNHN).</paragraph>
<paragraph id="9A018B00F007A07775C964B69AF6590B" pageId="14" pageNumber="131">Quedius tschinganensis gracilicornis: Tajikistan: ♂, &quot;Karatak Buchara [printed]/ Type [printed]/ Q. (Sauridus) tschinhganensis v. gracilicornis H. Coiffait det. [sic!] 1977 [pre-printed]&quot; (MNHN).</paragraph>
<caption id="5BB1C867771E33661B77BE71C0249BEB" pageId="14" pageNumber="131">
<paragraph id="506D7AAC1DECB49FD7A6E28FA34AECF5" pageId="14" pageNumber="131">
Figure 14. 
<taxonomicName id="804406A8020FD14BC6E15D3D9380AB10" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius tschinganensis" order="Coleoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="species" species="tschinganensis">Quedius tschinganensis</taxonomicName>
(new synonym of 
<taxonomicName id="CA0C52A83668620BE0077ACFDE6DBCD0" lsidName="Q. imitator" pageId="14" pageNumber="131" rank="species" species="imitator">Q. imitator</taxonomicName>
), types. 
<normalizedToken id="EAE64A47C21E13193B1979C8C2460661" originalValue="A–D">A-D</normalizedToken>
, holotype; 
<normalizedToken id="13116BBFAA435D700188A8D53560FF30" originalValue="E–H">E-H</normalizedToken>
, paratype. A, E, habitus. B, C, F, G, aedeagus: B, lateral view; C, dorsal view; F, median lobe, lateral view; G, paramare, underside. D, H, labels. Scale bars: A, E = 1 mm; B, C, F, G = 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EF10399D6F333E22633AA7046EC6D503" lastPageId="16" lastPageNumber="133" pageId="14" pageNumber="131" type="materials_examined">
<paragraph id="D0A636597F83BD600ECA3E3C42E5C42A" pageId="14" pageNumber="131">Additional material.</paragraph>
<paragraph id="567B088670AA954EC452630B7CDB530B" lastPageId="16" lastPageNumber="133" pageId="14" pageNumber="131">
Tajikistan: 3 ♂, Zeravshan Mt. Ridge, Chap-Dara River valley, 2500 m a.s.l., 26.VI.1983, S.K. Alekseev leg. (cRyv); 1 ♂, Pamir-Alai, Zeravshan Mt Ridge, Zavron valley, 2100-3000 m a.s.l., 12-13.VII.1990, M. 
<normalizedToken id="107F72320007DC03973F432921EFA528" originalValue="Schülke">Schuelke</normalizedToken>
&amp; D.W. Wrase leg (cSch); 1 ♂, Zeravshan Mt. Ridge, near Mazor, 14.VIII.1989, K.G. Michailov leg. (NHMD); Kazakhstan: 1 ♂, Makanchi District, Tarbagatay Mts, 6 km NE Kirovka (=Karatuma), Sholakterek River valley, ca. 1200 m a.s.l., 
<geoCoordinate id="802442D10DC2936A42424326CFBE674F" direction="north" orientation="latitude" precision="925" value="47.166668">47°10'N</geoCoordinate>
, 
<geoCoordinate id="B451CC0B8558C17497C0FFAA776E5DFC" direction="east" orientation="longitude" precision="925" value="82.1">82°06'E</geoCoordinate>
, highly disturbed 
<taxonomicName id="E38966CADDC8E2F7070015B13310E2F9" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
forest with 
<taxonomicName id="C011E2730677334E68DD30FCB66318B4" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="14" pageNumber="131" phylum="Chordata" rank="genus">Salix</taxonomicName>
, 
<taxonomicName id="19AB975C0F833DE19565D46514E66066" class="Magnoliopsida" family="Rosaceae" genus="Rosa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rosa" order="Rosales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Rosa</taxonomicName>
, 
<taxonomicName id="9DB32003ECBC518CE04545DEA21DB6AA" class="Magnoliopsida" family="Caprifoliaceae" genus="Lonicera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lonicera" order="Dipsacales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Lonicera</taxonomicName>
, 
<taxonomicName id="1E48575B141C216DC8915BF12C6561C3" class="Magnoliopsida" family="Rosaceae" genus="Crataegus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Crataegus" order="Rosales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Crataegus</taxonomicName>
, etc., 23-24.VI.2001, S.I. Golovatch leg. (cRyv); 1 ♂, Dzhungarskiy Alatau, S Koktuma, Alakol Lake, 05.VI.1962, L.V. Arnoldi leg. (ZIN); 2 ♂, Almaty Area, Dzhungarskiy Alatau Mts, 6 km NE Rudnichnyi, Koksu River canyon, 1300-1400 m a.s.l., 
<geoCoordinate id="A93113329EF80BF81B13A937278FBE61" direction="north" orientation="latitude" precision="925" value="44.683334">44°41'N</geoCoordinate>
, 
<geoCoordinate id="D035C8492B312BA303547796E25090B5" direction="east" orientation="longitude" precision="925" value="78.96667">78°58'E</geoCoordinate>
, 
<taxonomicName id="356B1CC6F16FF001EE26D30F25DCE259" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
sp., 
<taxonomicName id="70B56636A4CE0885AC9982F3EEDE76EC" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
, 
<taxonomicName id="4A38BE61CD27C3C62B7210BCE6D06BEC" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
etc. forest, 09-10.VI.2001, S.I. Golovatch leg. (cRyv); 2 ♂, Zalataysky Alatau, Krasnogorka [Sulutor], stream beach under tree, 75.13504E, 43.23457N, 28.VII.2010, V.A. Kastcheev leg. (ZIN); 2 ♂, 1 ♀, Lle-Alatau NP Talgar env., Ak-Bulak Resort horse and cow dung, 1750 m a.s.l., 43.26897N, 77.37145E, 08.V.2014, M. 
<normalizedToken id="529E36BFE42D88D05846799FE42BC503" originalValue="Kocián">Kocian</normalizedToken>
leg. (cKoc); 2 ♀, Lle-Alatau NP Talgar env., Ak-Bulak Resort, horse and cow dung, 1690 m a.s.l., 43.27039N, 77.37137E, 12-15.V.2014, M. 
<normalizedToken id="032E0E73123998FA090619DA7EC0C573" originalValue="Kocián">Kocian</normalizedToken>
leg. (cKoc); 1 ♂, Almaty Area, Zailiyskiy Alatau Mts, Medeo near Almaty, 1500-1600 m a.s.l., 
<geoCoordinate id="75095513C1418AE98E8D60F6FEE466AF" direction="north" orientation="latitude" precision="925" value="43.166668">43°10'N</geoCoordinate>
, 
<geoCoordinate id="9F140899FDB77CCE8790627B131FA388" direction="east" orientation="longitude" precision="925" value="77.066666">77°04'E</geoCoordinate>
, 
<taxonomicName id="0E6B0F00D55C901447D0325FC98B68B4" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
, 
<taxonomicName id="54D18F5F2621C41543D5CDCAF1201317" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
, etc. forest, 27.V.2001, S.I. Golovatch leg. (cRyv); 3 ♂, Almaty Area, Zailiyskiy Alatau Mts, ca. 20 km SE Turgen, Turgen River canyon, near Batun, 1750 m a.s.l., 
<geoCoordinate id="638B16EAD5B0E90107EF9D87682508AD" direction="north" orientation="latitude" precision="925" value="43.233334">43°14'N</geoCoordinate>
, 
<geoCoordinate id="597F5AF6E7B109CA74EB3D7055CB80BE" direction="east" orientation="longitude" precision="925" value="77.76667">77°46'E</geoCoordinate>
, 
<taxonomicName id="AF70DF4F0B86EB4FFA35E30ECF7DB861" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
, 
<taxonomicName id="643DF5F9B44CD2A99AB9973D08D5740B" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
sp., 
<taxonomicName id="E158E9FEE5EB0A3F3DD86C49D1FE3028" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="14" pageNumber="131" phylum="Chordata" rank="genus">Salix</taxonomicName>
, etc. forest, 25.V.2001, S.I. Golovatch leg. (cRyv); 1 ♂, Zailiysky Alatau, Chilik River, Sarybastau, 15.VI.1988, V.A. Kastcheev leg. (ZIN); 4 ♂, 1 ♀, Almaty Area, Uygurskiy Distr., Ketmen Mts, 5 km SE Kyrghyzsay (=Podgornoye), 1500-1900 m a.s.l., 
<geoCoordinate id="20D3BF6577E979A20835476951AEE662" direction="north" orientation="latitude" precision="925" value="43.283333">43°17'N</geoCoordinate>
, 
<geoCoordinate id="CA02B513108B872500E88A8779E0A7EB" direction="east" orientation="longitude" precision="925" value="79.51667">79°31'E</geoCoordinate>
, 
<taxonomicName id="6A62C911EEAA17B06156BB1B774FC759" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea" order="Pinales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
, 
<taxonomicName id="720511D34486B06E9169165C9C1613E4" class="Magnoliopsida" family="Betulaceae" genus="Betula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Betula" order="Fagales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Betula</taxonomicName>
sp., 
<taxonomicName id="F6296972C3489F3A891440C8804CB86B" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="14" pageNumber="131" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
, etc. forest, 01-02.VI.2001, S.I. Golovatch leg. (cRyv);1 ♂, Ketmen Mts, Malyi Kyrgisai, 28.VII.1987, V.A. Kastcheev leg., (ZIN); 3 ♂, Karatau Mts, 660 m a.s.l., 42°53' 
<geoCoordinate id="05107F95FF44F5E7B3016B50589ECADF" direction="north" orientation="latitude" precision="555" value="41.42">41.42N</geoCoordinate>
, 70°42' 
<geoCoordinate id="F5D9FD7CD85B4D9BF8D2428E36353A17" direction="east" orientation="longitude" precision="5555" value="56.6">56.6E</geoCoordinate>
, leaf litter along stream, 11.VII.2010, V.A. Kastcheev leg. (ZIN); 8 ♂, 1 ♀, Karatau Mts, Byzhi River, Rynagus stream, 757 m a.s.l., 43°57' 
<geoCoordinate id="DF4A49F5ACE9454A055CDA51132219EA" direction="north" orientation="latitude" precision="5555" value="8.7">08.7N</geoCoordinate>
, 68°12' 
<geoCoordinate id="860CA72C02CC7D667947FB04569618C2" direction="east" orientation="longitude" precision="5555" value="4.2">04.2E</geoCoordinate>
, 24-25.VII.2010, V.A. Kastcheev leg. (ZIN); 2 ♂, 1, Karatau Mts, Aktobe River, grove, 25.VII.2010, V.A. Kastcheev leg. (ZIN); 6 ♂, 1 ♀, Karatau Mts, Khantagi River, 570 m a.s.l., 43°33' 
<geoCoordinate id="838A9E1D15C08ABF5424F37E51D2264A" direction="north" orientation="latitude" precision="5555" value="32.4">32.4N</geoCoordinate>
, 68°40' 
<geoCoordinate id="41F06F69518A6272AEAF92171C8FABA0" direction="east" orientation="longitude" precision="5555" value="52.7">52.7E</geoCoordinate>
, 25.VI.2011, V.A. Kastcheev leg. (ZIN); same locality and collector, but 1 ♂, leaf litter under 
<taxonomicName id="7DE1411D811BF3D312A41133B9234A0D" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="14" pageNumber="131" phylum="Chordata" rank="genus">Salix</taxonomicName>
sp., 536 m a.s.l., 43°32' 
<geoCoordinate id="213055248512A6E1BE100EB94A5AC488" direction="north" orientation="latitude" precision="5555" value="46.5">46.5N</geoCoordinate>
, 68°39' 
<geoCoordinate id="450215AB4E8D559EA02065EEF8FCF99E" direction="east" orientation="longitude" precision="5555" value="50.6">50.6E</geoCoordinate>
, 21.VII.2010 (ZIN); ♂, 1 ♀, 27 km S Chulak-Kurgan, 04.VI.1983, B.V. Iskakov leg. (ZIN); 1 ♂, 1, Chimkent, Aksukent, Aksu River, 29.VI.1983, V.A. Kastcheev leg. (ZIN); 2 ♂, Aksu-Zhabagly Nature Reserve, Tokmak River, near border, under stones, 1600 m a.s.l., 30.V.1974, E.V. Ishkov leg. (ZIN); 1 ♂, 2 ♀, Aksu-Dzhabagly, Taldy-Bulak River, 15-25.VI.1983, B.V. Iskakov leg. (ZIN); 1 ♂, Aksu-Dzhabagly, Isbala River, 18.VII.1986, (ZIN); 1 ♂, Aksu-Djabagly, Djabagly River, tract Ulken-Kaindy, IV.1986, B.V. Iskakov leg. (ZIN); same locality and collector, but 1 ♂, Kshi-Kaindy River, 01.V.1986 (ZIN); Uzbekistan: 1 ♂, Kitab, 30.VII.1933, V.V. 
<pageBreakToken id="36C18A8EC0B6E51186E2474E8CEF08D8" pageId="15" pageNumber="132" start="start">Gussakovsky</pageBreakToken>
leg. (ZIN); Kyrgyzstan: 1 ♂, Terskey Alatoo Mts, Barskoon Valley, Chuli River, 15.VII.1983, S.K. Alekseev leg. (cRyv); 1 ♂, 1 ♀, Osh Area, Sary-Chelek Biosphere Reserve, near Arkit, Bakay-say Tract, 14.VII.1983, K.G. Mikhailov leg. (cRyv); 1 ♂, Chatkal Mt. Ridge, near Arkit, nut-fruit forest, 16.V.1961, V.A. Zaslavsky leg. (ZIN); 2 ♂, Tien Shan, Kichik-Alai Mt. Ridge, upper reaches of Kyrghyz-Ata River, Kara-Goy, 2400-2850 m a.s.l., 
<taxonomicName id="4433C520631D5F80B96894AD416CD741" class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus" order="Pinales" pageId="15" pageNumber="132" phylum="Tracheophyta" rank="genus">Juniperus</taxonomicName>
stand, 21-23.V.1993, S.I. Golovatch leg. (cRyv); 1 ♂, Osh Area, environs of Sary-Chelek Biosphere Reserve, confluent of Aflatun River, Batrakhan (=Baltyrkan) Tract, moss and leaf litter in birch forest &amp; under 
<taxonomicName id="B9EACB17F0CAABB6EF572FE058213791" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Picea schrenkiana" order="Pinales" pageId="15" pageNumber="132" phylum="Tracheophyta" rank="species" species="schrenkiana">Picea schrenkiana</taxonomicName>
at stream bank, 31.VII.1983, A.B. Ryvkin leg. (cRyv); 3 ♂, Tien Shan, Chatkal Mt. Ridge, Sary-Chelek Biosphere Reserve, 1550-2200 m a.s.l., forests, 29-31.V.1993, S.I.Golovatch leg. (cRyv); 2 ♂, 1 ♀, Osh Area, W Tien Shan, Ferganskiy Mt. Ridge, near Yarodar, 1100-1200 m a.s.l., 
<taxonomicName id="3F156AC69893B19CBC3E40CE229772E5" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juglans" order="Fagales" pageId="15" pageNumber="132" phylum="Tracheophyta" rank="genus">Juglans</taxonomicName>
forest, in leaf litter, 27-28.IX.1983, K.Yu. Eskov leg. (cRyv); 1 ♂, Osh Area, W Tien Shan, Ferganskiy Mt. Ridge, Yarodar, 1300 m a.s.l., rill bank, in leaf litter and under stones, 24-25.IX.1983, K.Yu. Eskov leg. (cRyv); 3 ♂, Tien Shan, Baubash-Ata Mt. Ridge, near Arslanbob, 1800-1900 m a.s.l., scrub, litter &amp; under stones. 19.V.1993. S.I. Golovatch leg. (cRyv); 10 ♂, 1 ♀, Tien Shan, Baubash-Ata 
<pageBreakToken id="A31F29FBCC2E407C39CF330B3B0C8E56" pageId="16" pageNumber="133" start="start">Mt</pageBreakToken>
. Ridge, near Yarodar, 1400-1700 m a.s.l., 
<taxonomicName id="3EAEFAD98BE414D9A019A66AC7C665D6" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juglans" order="Fagales" pageId="16" pageNumber="133" phylum="Tracheophyta" rank="genus">Juglans</taxonomicName>
forest, litter &amp; under bark. 16-17.V.1993. S.I. Golovatch leg. (cRyv); 4 ♂, 1 ♀, Ferganskiy Mt. Ridge, Kara-Alma, 1800 m a.s.l., 22-24.VI.1945, K.V. Arnoldi leg. (ZMMU); ♂ 1 ♂, 3 ♀, Gava, Fergana Valley, Jalal-Abad Region, near station of Forest Institute of Russian Academy of Sciences, 04.IX.1950, L.V. Arnoldi leg. (ZIN); 1 ♂, Fergana Valley, Kulun Lake, 3000 m a.s.l., 07.V.1993, I.I. Kabak leg. (ZIN); 2 ♂, 1 ♀, S Fergana Valley, Ak-Terek, 25.IX.1937, A.N. Kirichenko leg. (ZIN).
</paragraph>
</subSubSection>
<subSubSection id="11E10836EE6AC4499AC69C1E70ABC3AE" pageId="16" pageNumber="133" type="comments">
<paragraph id="A68C64D75CD0ADF24962977D86B59D40" pageId="16" pageNumber="133">Comments on the lectotype designation.</paragraph>
<paragraph id="5D8C2AAE7F9AD52CBAD960F57C3E6B87" pageId="16" pageNumber="133">
In the original description of 
<taxonomicName id="B2BCE85F9B5D1F406B0FCDF884A2772C" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
, 
<bibRefCitation id="5FD2D600173B9564807DE3330B507C87" pageId="16" pageNumber="133">Luze (1904)</bibRefCitation>
did not specify the number of syntypes but provided geographical data that indicated multiple syntypes collected in the localities &quot;Seravschan, Putchin-Pass, Darch, Obburden, Urmitan, Kumar; Jagnob, Varsaut; Iskander-Kul, Iskander-Darja&quot; (approximate coordinates as we interpret these localities are given in the Table. 2). Also, the syntype series must have included both sexes because male characters were specified separately in the description. In the ZIN collection we found 11 specimens from several localities along Zeravchan and Iskander Darya Rivers matching those in the original description (for details see 'Material 
<normalizedToken id="E5E3D5CB52A6B05A7892FEBCB27E6064" originalValue="examined’">examined'</normalizedToken>
above). Based on that and additional information from the specimen labels, there is no doubt that they are syntypes. Earlier they were identified by 
<bibRefCitation id="7047B65948987E8D16044C7646FFA7C1" author="Bohac, J" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="25" pageNumber="142" pagination="549 - 558" title="Novye i maloizvestnye zhuki-stafilinidy podsemeistva Staphylininae (Coleoptera, Staphylinidae)." volume="67" year="1988">
<normalizedToken id="C14DAF854BCC81A2EEC4A9A30B9CF1AA" originalValue="Boháč">Bohac</normalizedToken>
(1988)
</bibRefCitation>
as 
<taxonomicName id="A20CC395143B2FCF28E7E3D79E10BA06" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
without recognizing them as syntypes. In order to fix the identity of the species, we designate here one male syntype with more preciselocality &quot;Seravschan Darch Glasunov 1892&quot; (Darg, Sughd Distr.) as the lectotype. 
<bibRefCitation id="62559BD00838F246703B01F8F22A9E72" author="Bernhauer, M" journalOrPublisher="zoologisch-botanischen Gesellschaft in Wien" pageId="25" pageNumber="142" pagination="580 - 596" title="13. Folge neuer Staphyliniden der palaearktischen Fauna, nebst Bemerkungen. Verhandlungen der k. k." url="http://www.zobodat.at/pdf/VZBG_55_0580-0596.pdf" volume="55" year="1905">Bernhauer (1905)</bibRefCitation>
considered 
<taxonomicName id="D2290A5FD2DC44456610203B29DF07CA" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
as a &quot;rough form&quot; of 
<taxonomicName id="7018DEBEB46E1A69D43083FB16321360" lsidName="Q. oblitteratus" pageId="16" pageNumber="133" rank="species" species="oblitteratus">Q. oblitteratus</taxonomicName>
(now synonym of 
<taxonomicName id="C0F6DC0880EAA028E92D577D41A962E0" lsidName="Q. humeralis" pageId="16" pageNumber="133" rank="species" species="humeralis">Q. humeralis</taxonomicName>
Stephens, 1832). 
<bibRefCitation id="99556ADD74FE81CF94779F85ECF5624D" author="Gridelli, E" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="25" pageNumber="142" pagination="5 - 180" title="Studi sul genere Quedius Steph. (Coleoptera: Staphylinidae). Secondo contributo. Specie della regione paleartica." volume="3" year="1924">Gridelli (1924)</bibRefCitation>
not seeing types or any other material of 
<taxonomicName id="EC4D84B58ABF55C8F102F711A9151119" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
was not sure about the status of this species. Based on the non-type material, 
<bibRefCitation id="AE30F2D53B1E4EBF17DDF4E7823FD89B" author="Coiffait, H" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="25" pageNumber="142" pagination="391 - 424" title="Quedius nouveaux ou mal connus." volume="103" year="1967">Coiffait (1967</bibRefCitation>
, 
<bibRefCitation id="9F337CB42FE6EB2D6C2494A8115E98FE" pageId="16" pageNumber="133">1978</bibRefCitation>
) illustrated its aedeagus for the first time that here is shown to be the correct species interpretation. 
<bibRefCitation id="E15159994CA21E1D133BC425976D7001" author="Bohac, J" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="25" pageNumber="142" pagination="549 - 558" title="Novye i maloizvestnye zhuki-stafilinidy podsemeistva Staphylininae (Coleoptera, Staphylinidae)." volume="67" year="1988">
<normalizedToken id="06CB8E065FCDCB261678073EDB3AFF44" originalValue="Boháč">Bohac</normalizedToken>
(1988)
</bibRefCitation>
provided new records for the species from Uzbekistan and Turkmenistan that are reliable because he examined syntypes.
</paragraph>
</subSubSection>
<subSubSection id="38B5867BA996AAE91F1F61DCDB486339" pageId="16" pageNumber="133" type="comments">
<paragraph id="BB3AAF5791FC7D7B857BF7AED96413F9" pageId="16" pageNumber="133">Comments on the new synonym.</paragraph>
<paragraph id="6F3A5E57479A1D2AE42D5405163AFA52" pageId="16" pageNumber="133">
<bibRefCitation id="79FC7D3A2C4CD21FA5E7F9BDAF1C9F1A" author="Coiffait, H" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="25" pageNumber="142" pagination="44 - 54" title="Quedius nouveaux. 5 e note sur le genre Quedius." volume="105" year="1969">Coiffait (1969)</bibRefCitation>
described 
<taxonomicName id="0ECC420192662638D7FC9FD48A719DEC" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius tschinganensis" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="tschinganensis">Quedius tschinganensis</taxonomicName>
(Fig. 14) from Uzbekistan and separated it from 
<taxonomicName id="AFA6F12DD90CB2EBAA183EF6BADE73B7" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
by darker body coloration, antennal segment 3 longer than 2, presence of 
<normalizedToken id="4B52C0C1D8C4EBEE0D23AA28FBC43088" originalValue="‘lateral’">'lateral'</normalizedToken>
puncture on pronotum, denser punctation of the elytra and more elongated median lobe. Additionally 
<bibRefCitation id="57FB799320614ECCA72077C07664D258" author="Coiffait, H" journalOrPublisher="Nouvelle Revue d'Entomologie" pageId="25" pageNumber="142" pagination="133 - 143" title="Note sur quelques Quedius et Heterothops nouveaux ou mal connus (Coleoptera: Staphylinidae)." volume="7" year="1977">Coiffait (1977)</bibRefCitation>
described 
<taxonomicName id="8D64AB8FB9D2813A255F2C20A1AF9465" lsidName="Q. tschinganensis" pageId="16" pageNumber="133" rank="subspecies" species="tschinganensis" subspecies="gracilicornis">Q. tschinganensis gracilicornis</taxonomicName>
, a variety of 
<taxonomicName id="C7D2690D493CB515995DC9E1D13DDD47" lsidName="Q. tschinganensis" pageId="16" pageNumber="133" rank="species" species="tschinganensis">Q. tschinganensis</taxonomicName>
from Tajikistan based on some differences in coloration of the body and the proportions of antennae. Later (1977), he replaced the preoccupied name 
<taxonomicName id="82D69F7E1D838B1D111AE27D5A62D6C0" genus="Amblyopinina" lsidName="Amblyopinina gracilicornis" pageId="16" pageNumber="133" rank="species" species="gracilicornis">gracilicornis</taxonomicName>
by the new name 
<taxonomicName id="A2D31F47CA6B38D552E48680610009CA" genus="Amblyopinina" lsidName="Amblyopinina debilicornis" pageId="16" pageNumber="133" rank="species" species="debilicornis">debilicornis</taxonomicName>
. Both are unavailable names due to ICZN Article 15.2 as already noted in 
<bibRefCitation id="2C68976C76DD0B84A8D636B9A21728AE" author="Herman, LH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="25" pageNumber="142" pagination="3021 - 3840" title="Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second Millennium. VI. Staphylinine Group (Part 3). Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae: Incertae Sedis fossils, Protactinae." volume="265" year="2001">Herman (2001)</bibRefCitation>
.
</paragraph>
<paragraph id="164AFE60FA3B57C22FC9D9024C3D29D0" pageId="16" pageNumber="133">
Our examination of the material from Middle Asia, including types, showed continuous variability in the external morphology and aedeagus that connects the states of 
<taxonomicName id="3DF9A46B1E811F316A8EBDB715FA16B7" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
and 
<taxonomicName id="356A6CA1A99B0B8F1BFF11A9D2C67A10" lsidName="Q. tschinganensis" pageId="16" pageNumber="133" rank="species" species="tschinganensis">Q. tschinganensis</taxonomicName>
. The shape of the paramere varies from the state with narrow and sharp apex with lesser number of sensory peg setae arranged in regular rows away from the apex, to the state with obtuse apex and with more sensory peg setae arranged denser and closer to the apex (Figs 14C, G; 15). Shape of the median lobe is more stable and varies only slightly in length and degree of sharpness of its apex (Figs 14B, F; 15). Mapping of this variability across the species distribution does not show any geographical patterns. Externally all specimens including females, also show no traits that would correspond to variants different in the shape of the paramere. Thus we place 
<taxonomicName id="8785ECDDB96561ED91A49132EC779C3F" lsidName="Q. tschinganensis" pageId="16" pageNumber="133" rank="species" species="tschinganensis">Q. tschinganensis</taxonomicName>
Coiffait, 1969 in synonymy with 
<taxonomicName id="B25625A0D36659E630FD9BE414C4E281" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
Luze, 1904. Our study of the type specimen of 
<taxonomicName id="F3CF8B0AD23509D345AC0177245B8A72" lsidName="Q. tschinganensis" pageId="16" pageNumber="133" rank="subspecies" species="tschinganensis" subspecies="debilicornis">Q. tschinganensis debilicornis</taxonomicName>
also shows it to be conspecifc with 
<taxonomicName id="C48153E3C617D580E93E06EC0F409565" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="AAA4790F373C39A7093E999009581F67" pageId="16" pageNumber="133" type="distribution">
<paragraph id="476697885EBF83D2A4D397B7219C5DB2" pageId="16" pageNumber="133">Comments on taxonomy, distribution and bionomics.</paragraph>
<paragraph id="741D84C9CB657C471CAD922DF37321B3" pageId="16" pageNumber="133">
<taxonomicName id="3F33EB3E9D911A5BF61EF91F182ACEC0" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius imitator" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="imitator">Quedius imitator</taxonomicName>
can be diagnosed by the following character combination: body dark brown with darker head and abdomen; elytra with slightly yellowish anterior angles; antennae usually pale; scutellum without setiferous punctation; aedeagus with ventral tooth of median lobe located remotely from its apex, with median lobe and paramere very narrow, apex of paramere obtusely sharpened and sensory peg setae arranged in two regular rows convergent to each other. Among other 
<taxonomicName id="057CC8414D3F565EA6F38890C83E757B" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Raphirus" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="genus">Raphirus</taxonomicName>
that occur in Middle Asia, 
<taxonomicName id="85041267A0F8E0347B524E96EF8E67E8" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
is most similar to 
<taxonomicName id="916E78773F37115BE1687D24CE3CC0FB" lsidName="Q. cohaesus" pageId="16" pageNumber="133" rank="species" species="cohaesus">Q. cohaesus</taxonomicName>
from which it can be easily distinguished by the mentioned diagnostic characters of the aedeagus.
</paragraph>
<paragraph id="F14289C4A86906222B8DFB7489042E01" pageId="16" pageNumber="133">
Based on the examined material and literature (Table 1), 
<taxonomicName id="D35D543F52E59A872A658D1BD1F3FFA8" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
is widely distributed in all countries of Middle Asia (Fig. 15). According to the label data of the examined material, 
<taxonomicName id="BB7A351EDDA3D9B3D96BB6694BBB3508" lsidName="Q. imitator" pageId="16" pageNumber="133" rank="species" species="imitator">Q. imitator</taxonomicName>
inhabits ground based debris and leaf litter of mainly deciduous forests along rivers and streams at various elevations, up to 3000 m. Also it can be found in dung or under stones.
</paragraph>
<caption id="199C83FDB465B2FEC4B2D1A524B5D86C" pageId="16" pageNumber="133">
<paragraph id="480B1763E10B2C91267EBF4B815CA6D3" pageId="16" pageNumber="133">
Figure 15. 
<taxonomicName id="CA57C9D60757FED13099F36975CAD68E" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius imitator" order="Coleoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="imitator">Quedius imitator</taxonomicName>
, distribution, median lobe of the aedeagus laterally, and variability of the paramere (as an example specimens from one locality, indicated by black dot). Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>