246 lines
25 KiB
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246 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.295.4905" ID-GBIF-Dataset="59cbf2a9-41ca-4f5a-a350-ef034991798b" ID-PMC="PMC3677376" ID-Pensoft-Pub="1313-2970-295-1" ID-PubMed="23794874" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-295-1" ModsDocOrigin="ZooKeys 295" ModsDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" checkinTime="1451247354359" checkinUser="pensoft" docAuthor="Branstetter, Michael G." docDate="2013" docId="43B26ED53939BE3C3A00C1397438A34E" docLanguage="en" docName="ZooKeys 295: 1-277" docOrigin="ZooKeys 295" docSource="http://dx.doi.org/10.3897/zookeys.295.4905" docTitle="Stenamma tico Branstetter, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="219" masterDocId="FFB3D3328201C177FF99FFC5FFA8A72C" masterDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="215" updateTime="1668155749461" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branstetter, Michael G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>295</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>277</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.295.4905</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.295.4905</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-295-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152043643" LSID="urn:lsid:zoobank.org:act:65486F53-70F0-40A3-90EC-F4C24090422F" httpUri="http://treatment.plazi.org/id/43B26ED53939BE3C3A00C1397438A34E" lastPageId="218" lastPageNumber="219" pageId="214" pageNumber="215">
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<subSubSection pageId="214" pageNumber="215" type="nomenclature">
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<paragraph pageId="214" pageNumber="215">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:65486F53-70F0-40A3-90EC-F4C24090422F" class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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<taxonomicNameLabel pageId="214" pageNumber="215">sp. n.</taxonomicNameLabel>
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Worker: Figure 162; Queen: Figure 163; Map: Figure 164
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</paragraph>
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</subSubSection>
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<subSubSection pageId="214" pageNumber="215" type="type material">
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<paragraph pageId="214" pageNumber="215">Type material.</paragraph>
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<paragraph pageId="214" pageNumber="215">
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Holotype worker. COSTA RICA, Heredia Prov.: 10km NE Vara Blanca,
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.233">10.233°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-84.067">84.067°W</geoCoordinate>
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, 1500m, 9 Mar 2005 (Project ALAS, collection 15/WF/02) [INBio,specimen CASENT0622416]. Paratypes:same data as holotype [1w, CAS, CASENT0622417], [1w, FMNH, CASENT0600104], [1w, ICN, CASENT0623501], [1w, INBio, CASENT0623502], [1w, JTLC, CASENT0623503], [1w, LACM, CASENT0623504], [1w, MGBPC, CASENT0623505], [1w, MCZ, CASENT0623506], [1w, MZSP, CASENT0623507], [1w, UCD, CASENT0622414], [1w, USNM, CASENT0622415].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="214" pageNumber="215" type="worker diagnosis">
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<paragraph pageId="214" pageNumber="215">Worker diagnosis.</paragraph>
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<paragraph pageId="214" pageNumber="215">
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Integument shining, largely black to red-black; medium-sized species (see HL, ML, PrW below); head mostly smooth and shiny; mesosoma reticulately costate to coarsely rugoreticulate; propodeal spines reduced to small upward projecting points, or sharp right angles at interface between dorsal and declivitous faces of propodeum (PSL 0.14-0.18, PSI 1.4-1.9); eyes large and somewhat bulging (EL 0.15-0.18, REL 23-24), with 7-10 ommatidia at greatest diameter; anterior margin of clypeus with shallow median emargination; basal margin of mandible straight, without notch or substantial depression; pilosity on gastral dorsum long, flexuous, and relatively sparse. Similar species:
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma lobinodus" order="Hymenoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="lobinodus">Stenamma lobinodus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="214" pageNumber="215" type="geographic range">
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<paragraph pageId="214" pageNumber="215">Geographic range.</paragraph>
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<paragraph pageId="214" pageNumber="215">Nicaragua to Panama.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="216" lastPageNumber="217" pageId="214" pageNumber="215" type="worker description">
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<paragraph pageId="214" pageNumber="215">Worker description.</paragraph>
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<paragraph lastPageId="216" lastPageNumber="217" pageId="214" pageNumber="215">
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(11 measured) HL 0.71-0.85 (0.84), HW 0.63-0.76 (0.76), FLD 0.22-0.27 (0.26), PCW 0.06-0.07 (0.07), SL 0.60-0.77 (0.77), EL 0.15-0.18 (0.18), ACL 0.54-0.68 (0.68), ML 0.91-1.11 (1.11), PrW 0.48-0.58 (0.57), PSL
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<pageBreakToken pageId="215" pageNumber="216" start="start">0.14</pageBreakToken>
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-0.18 (0.14), SDL 0.08-0.10 (0.09), PL 0.37-0.46 (0.44), PH 0.21-0.26 (0.26), PW 0.16-0.22 (0.20), PPL 0.19-0.25 (0.23), PPH 0.19-0.25 (0.24), PPW 0.20-0.25 (0.25), MFL 0.72-0.91 (0.91), MTL 0.56-0.71 (0.71), CI 87-93 (91), SI 95-103
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<pageBreakToken pageId="216" pageNumber="217" start="start">(</pageBreakToken>
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101), REL 23-24 (24), FLI 33-36 (34), PSI 1.4-1.9 (1.4), MFI 82-88 (84), ACI1 63-66 (64) ACI2 84-91 (88).
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</paragraph>
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<paragraph pageId="216" pageNumber="217">
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Medium-sized species; general body color black to red-black, with brown patches on waist and gaster; mandibles, clypeus and appendages dark brown to yellow-brown; setae golden; mandible with 6-7 teeth, consisting of 3 distinct apical teeth, a basal tooth, and 2-3 inner teeth, which are often worn and indistinct; basal margin of mandible straight, without any notch or significant depression; dorsal surface of mandible mostly smooth and shiny, with scattered piligerous punctae and variable number of basal striae; median lobe of clypeus with a pair of vestigial longitudinal carinulae and/or several ill-defined rugulae, apex of lobe with a short transverse carinula, remainder of clypeus smooth and shiny; posterior extension of clypeus between frontal lobes rather broad (PCW 0.06-0.07), with subparallel sides; frontal lobes relatively well developed, but not markedly expanded dorsolaterally (as in
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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), nor completely obscuring the underlying torular lobes in full-face view; head roughly oval-shaped, slightly, but distinctly longer than broad (CI 87-93), posterior margin gently convex, never depressed medially; eyes relatively large and somewhat bulging (EL 0.15-0.18, REL 23-24), oval-shaped, with 7-10 ommatidia at greatest diameter; face mostly smooth and shining, with variable amount of weak carinulae/rugulae and punctae confined to lower
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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of head; most specimens with some carinulae around antennal sockets and on gena, others with more extensive carinulae fanning outward from frontal carinae, sometimes with reticulae between antennal carinae and eyes; scape relatively long (SI 95-103), reaching and slightly surpassing posterior margin of head when laid back; dorsal surface of scape striate; flagellum with distinct 4-segmented antennal club; mesosoma shiny, almost entirely reticulately costate to coarsely rugoreticulae; propodeal declivity smooth, sometimes with a few transverse carinulae; promesonotum in profile low-domed and roughly symmetrically, anterior slope a little steeper and longer than posterior slope; propodeal spines reduced to small upward projecting points, or only forming sharp right angles where dorsal and declivitous surfaces of propodeum meet (PSL 0.14-0.18, PSI 1.4-1.9); petiole moderately long (PL/HW 0.57-0.63) and robust, node roughly half length of petiole and asymmetrical, with anterior face long and gradually sloping, and posterior face nearly vertical, but slightly angled so that apex of petiole points posteriad; dorsum of petiolar node viewed from posterior side weakly rounded; postpetiole in profile similar in size to petiolar node (PPH/PH 0.91-0.97), roughly circular, although with anterior face slightly longer and more sloping than posterior face; petiole and postpetiole shiny, nodes mostly smooth, with some rugulae and faint punctae on sides; venter of petiole and postpetiole weakly punctate; gaster mostly smooth and shiny, with scattered piligerous punctae; most of body with single layer of long standing setae; setae on legs and scapes varying from predominately suberect to mostly decumbent; coxae and femoral venters always with a row of longer suberect setae.
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</paragraph>
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<caption pageId="216" pageNumber="217">
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<paragraph pageId="216" pageNumber="217">
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Figure 162.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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holotype worker (CASENT0622416) A Profile B Face C Dorsum D Anterior clypeal margin in anterodorsal view E Gaster.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="217" lastPageNumber="218" pageId="216" pageNumber="217" type="queen description">
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<paragraph pageId="216" pageNumber="217">Queen description.</paragraph>
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<paragraph lastPageId="217" lastPageNumber="218" pageId="216" pageNumber="217">
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(5 measured) HL 0.71-0.85 (0.79), HW 0.63-0.76 (0.73), FLD 0.22-0.27 (0.24), PCW 0.07-0.09 (0.07), SL 0.64-0.78 (0.69), EL 0.20-0.24 (0.21), ACL 0.53-0.66 (0.60), ML 1.05-1.29 (1.12), PrW 0.57-0.70 (0.60), PSL 0.18-0.21 (0.19), SDL 0.09-0.11 (0.10), PL 0.43-0.53 (0.48), PH 0.24-0.29 (0.25),
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<pageBreakToken pageId="217" pageNumber="218" start="start">PW</pageBreakToken>
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0.20-0.25 (0.22), PPL 0.20-0.26 (0.23), PPH 0.23-0.29 (0.25), PPW 0.25-0.31 (0.26), MFL 0.77-0.94 (0.81), MTL 0.61-0.73 (0.64), CI 91-93 (93), SI 93-95 (94), REL 28-30 (28), FLI 34-37 (24), PSI 2.0-2.2 (2.0), MFI 87-90 (90), ACI1 63-66 (64), ACI2 82-90 (87).
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</paragraph>
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<paragraph pageId="217" pageNumber="218">Same as worker except for standard queen modifications and the following: face usually more sculptured, with light fan of rugulae/carinulae extending from frontal lobes to ocelli, sculpture around antennal sockets more distinct; costae on mesoscutum with a decidedly longitudinal orientation, but often wavy, and usually with some reticulation anteriorly; costae on side of promesonotum longitudinal in orientation; mesopleuron mostly smooth and shiny; propodeal spines always present, short, projecting dorsoposteriad (PSL 0.18-0.21, PSI 2.0-2.2); wing venation as in Figure 163D.</paragraph>
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<caption pageId="217" pageNumber="218">
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<paragraph pageId="217" pageNumber="218">
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Figure 163.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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A Queen (CASENT0600122), profile B Same, head C Same, dorsum D Queen (INB0003732299).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="217" pageNumber="218" type="male">
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<paragraph pageId="217" pageNumber="218">Male.</paragraph>
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<paragraph pageId="217" pageNumber="218">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="217" pageNumber="218" type="biology">
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<paragraph pageId="217" pageNumber="218">Biology.</paragraph>
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<paragraph pageId="217" pageNumber="218">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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is restricted to wet forest habitats from sea level to about 1500 m, being most abundant at mid elevations. Nests have never been collected of this species, with nearly all specimens coming from extracts of sifted leaf litter taken from the forest floor. Once at a site in Nicaragua I found a trail of workers migrating across a clay bank in montane wet forest. Although a nest site could not be found, the number of workers involved suggests that they were relocating the nest. The presence of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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on clay banks indicates that it may have biological characteristics very similar to the closely related
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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, which is known to nest in clay bank environments (see biology of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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above).
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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has large, bulging eyes and it may be that it is more active at night than
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="217" pageNumber="218" type="comments">
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<paragraph pageId="217" pageNumber="218">Comments.</paragraph>
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<paragraph pageId="217" pageNumber="218">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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together form the diversum species group (diagnosis given under
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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above).
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</paragraph>
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<paragraph pageId="217" pageNumber="218">
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Within its range
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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is unlikely to be confused with any other species. Only in northern Nicaragua does it occur in sympatry with its sister species
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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. As mentioned above (see comments under
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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),
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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can be separated from
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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by its smaller propodeal spines and narrower frontal lobes.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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also can be separated from
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma diversum" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="diversum">Stenamma diversum</taxonomicName>
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, as well as from the similar looking
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma lobinodus" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="lobinodus">Stenamma lobinodus</taxonomicName>
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, by its relatively long scape (SI> 95 vs. <93) and metafemur (MFI <89 vs.> 94).
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</paragraph>
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<paragraph pageId="217" pageNumber="218">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
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shows some elevational variation with specimens from higher localities becoming larger, darker and more sculptured (especially on the face). This variation appears to be continuous, and thus is unlikely due to the existence of cryptic species.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="218" lastPageNumber="219" pageId="217" pageNumber="218" type="material examined">
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<paragraph pageId="217" pageNumber="218">Material examined.</paragraph>
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<paragraph lastPageId="218" lastPageNumber="219" pageId="217" pageNumber="218">
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COSTA RICA:Cartago: Navarro Farm, [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="9.884">9.884°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-83.883">83.883°W</geoCoordinate>
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], [ca. 1100m] (W. M. Mann); Guanacaste: Est. Pitilla, 9km S Sta. Cecilia, P.N. Guanacaste, [ca.
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.989">10.989°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-85.426">85.426°W</geoCoordinate>
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], 700m (C. Moraga); P.N.
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<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
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Tenorio,
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<normalizedToken originalValue="Volcán">Volcan</normalizedToken>
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Tenorio,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.6764">10.6764°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-84.9775">84.9775°W</geoCoordinate>
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, 1650m (J. Azofeita); Heredia: 9km NE Vara Blanca,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.23">10.23°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-84.1">84.10°W</geoCoordinate>
|
|
, 1450-1550m (ALAS); 11km ESE La Virgen,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.35">10.35°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-84.05">84.05°W</geoCoordinate>
|
|
, 250-350m (ALAS); 11km SE La Virgen,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.33">10.33°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-84.07">84.07°W</geoCoordinate>
|
|
, 550m (ALAS); 16km SSE La Virgen,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.267">10.267°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-84.083">84.083°W</geoCoordinate>
|
|
, 1050-1150m (ALAS); 12km N Vol. Barba,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.25">10.250°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-84.083">84.083°W</geoCoordinate>
|
|
, 1420m (J. Longino); Est. Biol. La Selva,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.433">10.433°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-84.017">84.017°W</geoCoordinate>
|
|
, 50-150m (J. Longino); P.N. Braulio Carrillo,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.4">10.40°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-84.03">84.03°W</geoCoordinate>
|
|
, 150-180m (TEAM); P.N. Braulio Carrillo,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.4">10.40°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-84.07">84.07°W</geoCoordinate>
|
|
, 300m (TEAM); Puntarenas
|
|
<pageBreakToken pageId="218" pageNumber="219" start="start">:</pageBreakToken>
|
|
Monteverde,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="555" value="10.3">10.30°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="555" value="-84.8">84.80°W</geoCoordinate>
|
|
, 1500m (J. Longino); NICARAGUA: Jinotega: P.N. Cerro Saslaya,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="13.77172">13.77172°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-85.01277">85.01277°W</geoCoordinate>
|
|
, 1110m (LLAMA); Matagalpa: R.N. Cerro
|
|
<normalizedToken originalValue="Musún">Musun</normalizedToken>
|
|
,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="12.96071">12.96071°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-85.2325">85.23250°W</geoCoordinate>
|
|
, 750m (LLAMA); PANAMA: Bocas del Toro:
|
|
<normalizedToken originalValue="Fortuna-Chiriquí">Fortuna-Chiriqui</normalizedToken>
|
|
Grande Road,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="8.783">8.783°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-82.183">82.183°W</geoCoordinate>
|
|
, 500m (D. M. Olson).
|
|
</paragraph>
|
|
<caption pageId="218" pageNumber="219">
|
|
<paragraph pageId="218" pageNumber="219">
|
|
Figure 164. Distribution map of
|
|
<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma tico" order="Hymenoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="tico">Stenamma tico</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |