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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FFF6FFB75AB6FA77446BC464" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus nhatrangensis V. S. Dang &amp; Naiki 2021" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571834" ID-Zenodo-Dep="7571834" LSID="urn:lsid:plazi:treatment:43284262FFF6FFB75AB6FA77446BC464" httpUri="http://treatment.plazi.org/id/43284262FFF6FFB75AB6FA77446BC464" pageId="105">
<subSubSection box="[136,1170,1490,1517]" pageId="105" type="nomenclature">
<paragraph blockId="105.[136,1452,1490,1624]" box="[136,1170,1490,1517]" pageId="105">
<emphasis bold="true" box="[136,171,1490,1516]" pageId="105">52.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki (2021: 16)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="16" authorityYear="2021" box="[178,813,1490,1517]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="105" phylum="Tracheophyta" rank="species" species="nhatrangensis">
<emphasis bold="true" box="[178,479,1490,1517]" italics="true" pageId="105">Lasianthus nhatrangensis</emphasis>
V.S.Dang &amp; Naiki (2021: 16)
</taxonomicName>
<figureCitation box="[844,923,1490,1516]" captionStart="FIGURE 80" captionStartId="107.[136,229,1789,1811]" captionTargetBox="[167,1418,194,1763]" captionTargetId="figure-18@107.[165,1418,193,1765]" captionTargetPageId="107" captionText="FIGURE 80. Lasianthus nhatrangensis V.S.Dang &amp; Naiki (from V1563, VNM): A. Branch with young fruits, B. Close-up of expanding young leaves, C. Stipules, D. Abaxial leaf surfaces, E. Inflorescence showing side view flower buds, FG. Fruits, J. Seeds (G: photos by Shuichiro Tagane)." figureDoi="http://doi.org/10.5281/zenodo.7571798" httpUri="https://zenodo.org/record/7571798/files/figure.png" pageId="105">Fig. 80</figureCitation>
(plate),
<figureCitation box="[1017,1096,1490,1516]" captionStart="FIGURE 78" captionStartId="104.[136,229,1799,1821]" captionTargetBox="[176,1405,209,1763]" captionTargetId="figure-25@104.[165,1418,193,1775]" captionTargetPageId="104" captionText="FIGURE 78. Distribution maps of Lasianthus nghiasonii, L. nhatrangensis, L. ngoclinhensis, L. nuichuaensis and L. oblongilobus." figureDoi="http://doi.org/10.5281/zenodo.7571792" httpUri="https://zenodo.org/record/7571792/files/figure.png" pageId="105">Fig. 78</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="105" type="materials_examined">
<paragraph blockId="105.[136,1452,1490,1624]" pageId="105">
<emphasis bold="true" box="[136,194,1530,1551]" pageId="105">Type:</emphasis>
<materialsCitation collectingDate="2014-07-13" collectionCode="VNM, FU" collectorName="Tagane S. &amp; Kanemitsu H. &amp; Dang V. S. &amp; Tran H. &amp; Hanh N. &amp; Loi X. N. &amp; Thach N. D. &amp; Dinh N. &amp; Hieu P. N. H. V" country="Vietnam" elevation="400" location="Iriomote Station" municipality="Hon Ba Nature Reserve" pageId="105" specimenCode="VNM00042923, VNM00042924, VNM00042925" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<collectingCountry box="[219,332,1530,1551]" name="Vietnam" pageId="105">VIETNAM</collectingCountry>
.
<collectingRegion box="[343,552,1530,1552]" country="Vietnam" name="Khanh Hoa" pageId="105">Khanh Hoa Province</collectingRegion>
,
<collectingMunicipality box="[564,803,1530,1551]" pageId="105">Hon Ba Nature Reserve</collectingMunicipality>
,
<quantity box="[814,878,1530,1551]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="105" unit="m" value="400.0">
<elevation box="[814,878,1530,1551]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="105" unit="m" value="400.0">400 m</elevation>
</quantity>
elev.,
<date box="[945,1076,1530,1552]" pageId="105" value="2014-07-13">
<collectingDate box="[945,1076,1530,1552]" pageId="105" value="2014-07-13">13 July 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="105">
<collectorName box="[1088,1187,1530,1552]" pageId="105">Tagane S.</collectorName>
,
<collectorName box="[1199,1336,1530,1552]" pageId="105">Kanemitsu H.</collectorName>
,
<collectorName box="[1348,1448,1530,1552]" pageId="105">Dang V.S.</collectorName>
,
<collectorName box="[189,265,1567,1588]" pageId="105">Tran H.</collectorName>
with
<collectorName box="[317,402,1566,1588]" pageId="105">Hanh N.</collectorName>
,
<collectorName box="[411,494,1566,1588]" pageId="105">Loi X.N.</collectorName>
,
<collectorName box="[503,617,1566,1588]" pageId="105">Thach N.D.</collectorName>
,
<collectorName box="[626,705,1566,1588]" pageId="105">Dinh N.</collectorName>
,
<collectorName box="[714,851,1566,1588]" pageId="105">
Hieu P.N.H.
<date box="[836,851,1566,1587]" pageId="105" value="1563-05">V</date>
</collectorName>
1563
</emphasis>
(
<typeStatus box="[911,997,1566,1588]" pageId="105">holotype</typeStatus>
<collectionCode box="[1002,1063,1566,1587]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1077,1237,1566,1587]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM00042923</specimenCode>
]; isotypes
<collectionCode box="[1337,1374,1566,1587]" country="China" name="Fudan University, Department of Biology" pageId="105">FU</collectionCode>
!,
<collectionCode box="[1387,1448,1566,1587]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[195,352,1602,1623]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM00042924</specimenCode>
,
<specimenCode box="[361,518,1602,1624]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM00042925</specimenCode>
], the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFF6FFB75AB6FA77446BC464:CE5EA5AFFFF6FFB75885F9E7437ECA10" box="[699,856,1602,1624]" country="Vietnam" municipality="Hon Ba Nature Reserve" name="Iriomote Station" pageId="105" stateProvince="Khanh Hoa">Iriomote Station</location>
(Japan))
</materialsCitation>
.
</paragraph>
<paragraph blockId="105.[136,1452,1670,2093]" pageId="105">
<emphasis bold="true" box="[136,403,1670,1696]" pageId="105">
<collectingRegion box="[136,262,1670,1696]" pageId="105">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2014-07-13" collectionCode="FU, VNM" collectorName="Tagane S. &amp; Kanemitsu H. &amp; Dang V. S. &amp; Tran H. &amp; Hanh N. &amp; Loi X. N. &amp; Thach N. D. &amp; Dinh N. &amp; Hieu P. N. H. V &amp; Japan" country="Vietnam" elevation="400" latitude="12.108617" location="Iriomote Station" longLatPrecision="1" longitude="108.98722" municipality="Hon Ba Nature Reserve" pageId="105" specimenCount="1" stateProvince="Khanh Hoa">
<collectingCountry box="[432,564,1670,1696]" name="Vietnam" pageId="105">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[583,684,1670,1696]" pageId="105">Central:</emphasis>
<collectingRegion box="[701,957,1670,1697]" country="Vietnam" name="Khanh Hoa" pageId="105">Khanh Hoa Province</collectingRegion>
,
<collectingMunicipality box="[978,1275,1671,1697]" pageId="105">Hon Ba Nature Reserve</collectingMunicipality>
,
<quantity box="[1296,1376,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="105" unit="m" value="400.0">
<elevation box="[1296,1376,1670,1697]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="105" unit="m" value="400.0">400 m</elevation>
</quantity>
elev.,
<geoCoordinate box="[136,315,1706,1732]" degrees="12" direction="north" minutes="06" orientation="latitude" pageId="105" precision="1" seconds="31.02" value="12.108617">12°0631.02N</geoCoordinate>
,
<geoCoordinate box="[328,518,1706,1733]" degrees="108" direction="east" minutes="59" orientation="longitude" pageId="105" precision="1" seconds="14.01" value="108.98722">108°5914.01E</geoCoordinate>
, in evergreen forests,
<date box="[771,922,1706,1732]" pageId="105" value="2014-07-13">
<collectingDate box="[771,922,1706,1732]" pageId="105" value="2014-07-13">13 July 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="105">
<collectorName box="[935,1048,1707,1733]" pageId="105">Tagane S.</collectorName>
,
<collectorName box="[1062,1221,1707,1732]" pageId="105">Kanemitsu H.</collectorName>
,
<collectorName box="[1234,1349,1707,1733]" pageId="105">Dang V.S.</collectorName>
,
<collectorName box="[1362,1452,1707,1732]" pageId="105">Tran H.</collectorName>
with
<collectorName box="[195,296,1742,1768]" pageId="105">Hanh N.</collectorName>
,
<collectorName box="[310,411,1743,1768]" pageId="105">Loi X.N.</collectorName>
,
<collectorName box="[425,561,1742,1768]" pageId="105">Thach N.D.</collectorName>
,
<collectorName box="[576,670,1742,1768]" pageId="105">Dinh N.</collectorName>
,
<collectorName box="[685,852,1742,1768]" pageId="105">
Hieu P.N.H.
<date box="[834,852,1742,1768]" pageId="105" value="1563-05">V</date>
</collectorName>
1563
</emphasis>
(
<collectionCode box="[925,965,1742,1768]" country="China" name="Fudan University, Department of Biology" pageId="105">FU</collectionCode>
,
<collectionCode box="[979,1047,1743,1769]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="105" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FFF6FFB75AB6FA77446BC464:CE5EA5AFFFF6FFB75ECFF96B458DCAA0" box="[1265,1451,1742,1768]" country="Vietnam" latitude="12.108617" longLatPrecision="1" longitude="108.98722" municipality="Hon Ba Nature Reserve" name="Iriomote Station" pageId="105" stateProvince="Khanh Hoa">Iriomote Station</location>
(
<collectorName box="[144,213,1778,1804]" pageId="105">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="105" type="distribution">
<paragraph blockId="105.[136,1452,1670,2093]" box="[189,1394,1814,1841]" pageId="105">
<emphasis bold="true" box="[189,345,1814,1840]" pageId="105">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[508,603,1814,1840]" name="Vietnam" pageId="105">Vietnam</collectingCountry>
(known only from the Hon Ba Nature Reserve,
<collectingRegion box="[1142,1382,1814,1841]" country="Vietnam" name="Khanh Hoa" pageId="105">Khanh Hoa Province</collectingRegion>
).
</paragraph>
<paragraph blockId="105.[136,1452,1670,2093]" box="[189,1222,1850,1877]" pageId="105">
<emphasis bold="true" box="[189,423,1850,1876]" pageId="105">Habitat &amp; Ecology:</emphasis>
—Low montane forest under the shade of the canopy at ca.
<quantity box="[1091,1162,1850,1877]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="105" unit="m" value="400.0">400 m</quantity>
elev.
</paragraph>
<paragraph blockId="105.[136,1452,1670,2093]" box="[189,834,1886,1913]" pageId="105">
<emphasis bold="true" box="[189,323,1886,1912]" pageId="105">Phenology:</emphasis>
—Flowering and fruiting from March to July.
</paragraph>
<paragraph blockId="105.[136,1452,1670,2093]" box="[189,666,1922,1951]" pageId="105">
<emphasis bold="true" box="[189,405,1922,1948]" pageId="105">Vernacular name:</emphasis>
—Xú hưƠng nha trang.
</paragraph>
<paragraph blockId="105.[136,1452,1670,2093]" pageId="105">
<emphasis bold="true" box="[189,265,1958,1984]" pageId="105">Notes:</emphasis>
<taxonomicName box="[294,584,1958,1984]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="105" phylum="Tracheophyta" rank="species" species="nhatrangensis">
<emphasis box="[294,584,1958,1984]" italics="true" pageId="105">Lasianthus nhatrangensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName authority="Naiki (2015: 156)" authorityName="Naiki" authorityPageNumber="156" authorityYear="2015" box="[738,1092,1958,1985]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="105" phylum="Tracheophyta" rank="species" species="bokoensis">
<emphasis box="[738,879,1958,1984]" italics="true" pageId="105">L. bokoensis</emphasis>
<bibRefCitation author="Naiki, A. &amp; Tagane, S. &amp; Chhang, P. &amp; Toyama, H. &amp; Zhu, H. &amp; Dang, V. S. &amp; Yahara, T." box="[888,1092,1958,1985]" pageId="105" pagination="153 - 179" refId="ref54656" refString="Naiki, A., Tagane, S., Chhang, P., Toyama, H., Zhu, H., Dang, V. S. &amp; Yahara, T. (2015) Flora of Bokor National Park, Cambodia II: four new species and nine new records of Lasianthus (Rubiaceae) from Cambodia. Acta Phytotaxonomica et Geobotanica 66 (3): 153 - 179. https: // doi. org / 10.18942 / apg. KJ 00010115702" type="journal article" year="2015">Naiki (2015: 156)</bibRefCitation>
</taxonomicName>
which was recently discovered from Bokor National Park,
<collectingCountry box="[447,565,1994,2020]" name="Cambodia" pageId="105">Cambodia</collectingCountry>
, in the shade of the leaves and fruits, number of pyrenes, but differs in having shorter petioles (
<quantity box="[323,417,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="105" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long vs.
<quantity box="[519,612,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="105" unit="mm" value="7.5" valueMax="9.0" valueMin="6.0">69 mm</quantity>
long), smaller stipules (
<quantity box="[881,997,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="105" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long vs.
<quantity box="[1099,1192,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="105" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long) and calyx (
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="105" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
long vs.
<quantity box="[282,396,2066,2093]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="105" unit="mm" value="1.85" valueMax="2.0" valueMin="1.7">1.72 mm</quantity>
long for tubes and
<quantity box="[613,706,2066,2093]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="105" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long vs.
<quantity box="[808,922,2066,2093]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="105" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">34.5 mm</quantity>
long for lobes).
</paragraph>
</subSubSection>
</treatment>
</document>