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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FF85FFC45AB6FD314369CA7A" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus bidoupensis V. S. Dang &amp; Naiki 2019" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571570" ID-Zenodo-Dep="7571570" LSID="urn:lsid:plazi:treatment:43284262FF85FFC45AB6FD314369CA7A" httpUri="http://treatment.plazi.org/id/43284262FF85FFC45AB6FD314369CA7A" pageId="26">
<subSubSection box="[136,1142,660,687]" pageId="26" type="nomenclature">
<paragraph blockId="26.[136,1452,660,793]" box="[136,1142,660,687]" pageId="26">
<emphasis bold="true" box="[136,157,660,686]" pageId="26">4.</emphasis>
<taxonomicName ID-CoL="8Y54T" authority="V. S. Dang &amp; Naiki (2019: 191)" authorityName="V. S. Dang &amp; Naiki" authorityPageNumber="191" authorityYear="2019" box="[164,784,660,687]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="26" phylum="Tracheophyta" rank="species" species="bidoupensis">
<emphasis bold="true" box="[164,437,660,687]" italics="true" pageId="26">Lasianthus bidoupensis</emphasis>
V.S.
<bibRefCitation author="Dang, V. S. &amp; Naiki, A." box="[486,784,660,686]" pageId="26" pagination="33 - 40" refId="ref54157" refString="Dang, V. S. &amp; Naiki, A. (2019) Two new species of Lasianthus (Rubiaceae) from Hoang Lien National Park, northern Vietnam. Phytotaxa 423 (1): 33 - 40. https: // doi. org / 10.11646 / phytotaxa. 423.1.4" type="journal article" year="2019">Dang &amp; Naiki (2019: 191)</bibRefCitation>
</taxonomicName>
—
<figureCitation box="[815,894,660,686]" captionStart="FIGURE 18" captionStartId="27.[136,229,1755,1777]" captionTargetBox="[172,1415,194,1726]" captionTargetId="figure-18@27.[164,1421,193,1732]" captionTargetPageId="27" captionText="FIGURE 18. Lasianthus bidoupensis V.S.Dang &amp; Naiki (from Dang 250, VNM and the herbarium of Iriomote Station (Japan)): A. Branch with flower buds, B. Stipules, C. Abaxial leaf surfaces, D–F. Inflorescence showing lateral view bracts and flowers, G. Young fruit, H. Mature fruits, I. Cross section of fruit with 5 pyrenes." figureDoi="http://doi.org/10.5281/zenodo.7571602" httpUri="https://zenodo.org/record/7571602/files/figure.png" pageId="26">Fig. 18</figureCitation>
(plate),
<figureCitation box="[988,1068,660,687]" captionStart="FIGURE 15" captionStartId="22.[136,229,1778,1800]" captionTargetBox="[173,1409,201,1747]" captionTargetId="figure-24@22.[165,1418,193,1754]" captionTargetPageId="22" captionText="FIGURE 15. Distribution maps of Lasianthus annamicus, L. attenuatus, L. bachmaensis and L. bidoupensis." figureDoi="http://doi.org/10.5281/zenodo.7571596" httpUri="https://zenodo.org/record/7571596/files/figure.png" pageId="26">Fig. 15</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="26" type="materials_examined">
<paragraph blockId="26.[136,1452,660,793]" pageId="26">
<emphasis bold="true" box="[136,194,700,721]" pageId="26">Type:</emphasis>
—
<materialsCitation collectingDate="2018-02-10" collectionCode="VNM, HN" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang" country="Vietnam" elevation="1436" location="Dalat City" municipality="Bidoup Nui Ba National Park" pageId="26" specimenCode="VNM00042870, VNM00042871, VNM00042872" specimenCount="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry box="[219,332,700,721]" name="Vietnam" pageId="26">VIETNAM</collectingCountry>
.
<collectingRegion box="[344,548,700,722]" country="Vietnam" name="Lam Dong" pageId="26">Lam Dong Province</collectingRegion>
,
<collectingMunicipality box="[560,859,700,722]" pageId="26">Bidoup Nui Ba National Park</collectingMunicipality>
,
<quantity box="[872,936,700,722]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="26" unit="km" value="50.0">50 km</quantity>
northeast of
<location LSID="urn:lsid:plazi:treatment:43284262FF85FFC45AB6FD314369CA7A:CE5EA5AFFF85FFC45E11FD1944B0CE9A" box="[1071,1174,700,722]" country="Vietnam" municipality="Bidoup Nui Ba National Park" name="Dalat City" pageId="26" stateProvince="Lam Dong">Dalat City</location>
,
<quantity box="[1187,1263,700,722]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="26" unit="m" value="1436.0">
<elevation box="[1187,1263,700,722]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="26" unit="m" value="1436.0">1436 m</elevation>
</quantity>
elev.,
<date pageId="26" value="2018-02-10">
<collectingDate pageId="26" value="2018-02-10">10 February 2018</collectingDate>
</date>
,
<emphasis box="[252,610,736,758]" italics="true" pageId="26">
<collectorName box="[252,350,736,758]" pageId="26">Dang V.S.</collectorName>
&amp;
<collectorName box="[384,501,736,758]" pageId="26">Hoang N.S.</collectorName>
,
<collectorName box="[513,566,737,758]" pageId="26">Dang</collectorName>
250
</emphasis>
(
<typeStatus box="[625,711,736,758]" pageId="26">holotype</typeStatus>
<collectionCode box="[719,780,736,757]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[796,957,736,757]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM00042870</specimenCode>
]; isotypes
<collectionCode box="[1061,1102,736,757]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="26" type="Herbarium">HN</collectionCode>
!, VNMN!,
<collectionCode box="[1213,1274,736,757]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[1291,1448,736,757]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM00042871</specimenCode>
,
<specimenCode box="[189,349,772,793]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM00042872</specimenCode>
])
</materialsCitation>
.
</paragraph>
<paragraph blockId="26.[136,1452,840,1587]" pageId="26">
<emphasis bold="true" box="[136,394,840,866]" pageId="26">
<collectingRegion box="[136,262,840,866]" pageId="26">Specimens</collectingRegion>
examined:
</emphasis>
—
<materialsCitation accessionNumber="N10098" collectingDate="1839-05" collectingDateMax="2018-02-11" collectingDateMin="1839-05" collectionCode="VNM, FU" collectorName="Dang V. S. &amp; Hoang N. S. &amp; Dang &amp; Japan &amp; Toyama H. &amp; Tagane S. &amp; Nagamasu H. &amp; Naiki A. &amp; Tran H. &amp; Yang C. J." country="Vietnam" county="Bidoup Nui Ba National Park" elevation="1436" location="Iriomote Station" municipality="Dalat City" pageId="26" specimenCount="1" stateProvince="Lam Dong">
<collectingCountry box="[423,555,841,867]" name="Vietnam" pageId="26">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[566,667,840,866]" pageId="26">Central:</emphasis>
<collectingRegion box="[675,907,840,867]" country="Vietnam" name="Lam Dong" pageId="26">Lam Dong Province</collectingRegion>
,
<collectingCounty box="[919,1257,840,867]" pageId="26">Bidoup Nui Ba National Park</collectingCounty>
,
<quantity box="[1269,1341,840,867]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="26" unit="km" value="50.0">50 km</quantity>
northeast of
<collectingMunicipality box="[166,282,876,902]" pageId="26">Dalat City</collectingMunicipality>
,
<quantity box="[292,377,876,903]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="26" unit="m" value="1436.0">
<elevation box="[292,377,876,903]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="26" unit="m" value="1436.0">1436 m</elevation>
</quantity>
elev.,
<date box="[448,647,876,902]" pageId="26" value="2018-02-11">
<collectingDate box="[448,647,876,902]" pageId="26" value="2018-02-11">11 February 2018</collectingDate>
</date>
,
<emphasis box="[658,1069,876,903]" italics="true" pageId="26">
<collectorName box="[658,768,877,903]" pageId="26">Dang V.S.</collectorName>
&amp;
<collectorName box="[802,934,877,903]" pageId="26">Hoang N.S.</collectorName>
,
<collectorName box="[945,1007,877,902]" pageId="26">Dang</collectorName>
250a
</emphasis>
(
<collectionCode box="[1083,1153,876,902]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FF85FFC45AB6FD314369CA7A:CE5EA5AFFF85FFC45F75FCC940F1CFE2" country="Vietnam" county="Bidoup Nui Ba National Park" municipality="Dalat City" name="Iriomote Station" pageId="26" stateProvince="Lam Dong">Iriomote Station</location>
(
<collectorName box="[230,299,912,938]" pageId="26">Japan</collectorName>
)); in the same location,
<quantity box="[565,650,912,939]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="26" unit="m" value="1490.0">
<elevation box="[565,650,912,939]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="26" unit="m" value="1490.0">1490 m</elevation>
</quantity>
elev.,
<date box="[721,940,912,938]" pageId="26" value="2014-11-19">
<collectingDate box="[721,940,912,938]" pageId="26" value="2014-11-19">19 November 2014</collectingDate>
</date>
,
<emphasis italics="true" pageId="26">
<collectorName box="[950,1074,913,938]" pageId="26">Toyama H.</collectorName>
,
<collectorName box="[1085,1196,913,939]" pageId="26">Tagane S.</collectorName>
,
<collectorName box="[1208,1321,913,939]" pageId="26">Dang V.S.</collectorName>
,
<collectorName pageId="26">Nagamasu H.</collectorName>
,
<collectorName box="[178,271,948,975]" pageId="26">Naiki A.</collectorName>
,
<collectorName box="[284,374,949,974]" pageId="26">Tran H.</collectorName>
,
<collectorName box="[386,494,949,975]" pageId="26">Yang C.J.</collectorName>
<date box="[501,575,948,974]" pageId="26" value="1839-05">
<collectingDate box="[501,575,948,974]" pageId="26" value="1839-05">V1839</collectingDate>
</date>
&amp; V4364
</emphasis>
(
<collectionCode box="[699,739,948,974]" country="China" name="Fudan University, Department of Biology" pageId="26">FU</collectionCode>
,
<collectionCode box="[750,818,949,975]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FF85FFC45AB6FD314369CA7A:CE5EA5AFFF85FFC45E3FFC11449ECF86" box="[1025,1208,948,974]" country="Vietnam" county="Bidoup Nui Ba National Park" municipality="Dalat City" name="Iriomote Station" pageId="26" stateProvince="Lam Dong">Iriomote Station</location>
(
<collectorName box="[1224,1293,948,974]" pageId="26">Japan</collectorName>
)); in the same location,
<quantity box="[244,333,984,1011]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="26" unit="m" value="1438.0">
<elevation box="[244,333,984,1011]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="26" unit="m" value="1438.0">1438 m</elevation>
</quantity>
elev.,
<date box="[413,640,984,1010]" pageId="26" value="2017-11-26">
<collectingDate box="[413,640,984,1010]" pageId="26" value="2017-11-26">26 November 2017</collectingDate>
</date>
, in moist evergreen forest,
<emphasis box="[962,1314,984,1011]" italics="true" pageId="26">
<collectorName box="[962,1057,984,1011]" pageId="26">Naiki A.</collectorName>
&amp;
<collectorName box="[1100,1215,985,1011]" pageId="26">Dang V.S.</collectorName>
<accessionNumber box="[1225,1314,984,1010]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/N10098" pageId="26">N10098</accessionNumber>
</emphasis>
(
<collectionCode box="[1333,1403,984,1010]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="26" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FF85FFC45AB6FD314369CA7A:CE5EA5AFFF85FFC45B1EFC5941F1C85E" box="[288,471,1020,1046]" country="Vietnam" county="Bidoup Nui Ba National Park" municipality="Dalat City" name="Iriomote Station" pageId="26" stateProvince="Lam Dong">Iriomote Station</location>
(Japan))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="26" type="distribution">
<paragraph blockId="26.[136,1452,840,1587]" pageId="26">
<emphasis bold="true" box="[189,345,1056,1082]" pageId="26">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[513,608,1056,1082]" name="Vietnam" pageId="26">Vietnam</collectingCountry>
(currently known only from Bidoup Nui Ba National Park in
<collectingRegion country="Vietnam" name="Lam Dong" pageId="26">Lam Dong Province</collectingRegion>
).
</paragraph>
<paragraph blockId="26.[136,1452,840,1587]" pageId="26">
<emphasis bold="true" box="[189,423,1128,1154]" pageId="26">Habitat &amp; Ecology:</emphasis>
—Growing in evergreen forest at
<quantity box="[803,958,1128,1155]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="26" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</quantity>
elev. in Bidoup Nui Ba National Park,
<collectingRegion country="Vietnam" name="Lam Dong" pageId="26">Lam Dong Province</collectingRegion>
, southern
<collectingCountry box="[432,531,1164,1190]" name="Vietnam" pageId="26">Vietnam</collectingCountry>
. This species was collected in the understory of forest dominated by
<taxonomicName authorityName="Dumortier" authorityYear="1829" box="[1340,1447,1165,1191]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="26" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
,
<taxonomicName box="[136,243,1200,1226]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="26" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
,
<taxonomicName box="[254,374,1200,1226]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="26" phylum="Tracheophyta" rank="family">Rubiaceae</taxonomicName>
,
<taxonomicName box="[385,503,1201,1227]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="26" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
,
<taxonomicName box="[514,635,1201,1227]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="26" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
,
<taxonomicName box="[646,808,1200,1226]" class="Magnoliopsida" family="Symplocaceae" kingdom="Plantae" order="Ericales" pageId="26" phylum="Tracheophyta" rank="family">Symplocaceae</taxonomicName>
and
<taxonomicName box="[862,987,1200,1226]" class="Magnoliopsida" family="Clusiaceae" kingdom="Plantae" order="Malpighiales" pageId="26" phylum="Tracheophyta" rank="family">Clusiaceae</taxonomicName>
.
</paragraph>
<paragraph blockId="26.[136,1452,840,1587]" box="[189,1116,1236,1263]" pageId="26">
<emphasis bold="true" box="[189,323,1236,1262]" pageId="26">Phenology:</emphasis>
—Flowering and fruiting from November to February of the next year.
</paragraph>
<paragraph blockId="26.[136,1452,840,1587]" box="[189,638,1272,1301]" pageId="26">
<emphasis bold="true" box="[189,405,1272,1298]" pageId="26">Vernacular name:</emphasis>
—Xú hưƠng bidoup.
</paragraph>
<paragraph blockId="26.[136,1452,840,1587]" pageId="26">
<emphasis bold="true" box="[189,265,1308,1334]" pageId="26">Notes:</emphasis>
—
<taxonomicName box="[294,564,1308,1334]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="26" phylum="Tracheophyta" rank="species" species="bidoupensis">
<emphasis box="[294,564,1308,1334]" italics="true" pageId="26">Lasianthus bidoupensis</emphasis>
</taxonomicName>
was described in 2018 from our collections (
<emphasis box="[1118,1235,1308,1334]" italics="true" pageId="26">Dang 250</emphasis>
). It can easily be recognized by having elliptic-oblong to elliptic-lanceolate leaf blades, secondary veins 6–7 on each side of midrib, flowers fascicular, pinkish white to white, and fruits ovoid or ellipsoid, deep red when ripe.
</paragraph>
<paragraph blockId="26.[136,1452,840,1587]" pageId="26">
This species is closely related to
<taxonomicName authority="V. S. Dang, Tagane &amp; H. Tran (2015: 352)" authorityName="V. S. Dang, Tagane &amp; H. Tran" authorityPageNumber="352" authorityYear="2015" box="[558,1139,1416,1443]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="26" phylum="Tracheophyta" rank="species" species="yaharae">
<emphasis box="[558,679,1416,1442]" italics="true" pageId="26">L. yaharae</emphasis>
V.S.Dang, Tagane &amp; H.Tran (2015: 352)
</taxonomicName>
and
<taxonomicName authority="Wernham (1921: 138)" authorityName="Wernham" authorityPageNumber="138" authorityYear="1921" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="26" phylum="Tracheophyta" rank="species" species="dalatensis">
<emphasis box="[1194,1337,1416,1442]" italics="true" pageId="26">L. dalatensis</emphasis>
Wernham (1921: 138)
</taxonomicName>
based on leaf morphology, deeply dissected calyx lobes and cylindrical corolla, but differs from them in having larger leaves and bracts, longer petioles, stipules, calyx, and corolla tubes. Fruit color is also different when ripe. It is also similar to
<taxonomicName authority="Blume (1826: 998)" authorityName="Blume" authorityPageNumber="998" authorityYear="1826" box="[409,758,1524,1550]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="26" phylum="Tracheophyta" rank="species" species="inodorus">
<emphasis box="[409,538,1524,1550]" italics="true" pageId="26">L. inodorus</emphasis>
Blume (1826: 998)
</taxonomicName>
but differs from the latter in having larger bracts (
<quantity box="[1319,1413,1524,1551]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="26" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
vs.
<quantity box="[136,229,1560,1587]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="26" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
long) and longer calyx tubes (
<quantity box="[570,664,1560,1587]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="26" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
vs.
<quantity box="[709,774,1560,1587]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="26" unit="mm" value="2.0">2 mm</quantity>
long).
</paragraph>
</subSubSection>
</treatment>
</document>
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