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<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FF82FFC15AB6F8E445ACCD91" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus caeruleus Pitard 1924" docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571582" ID-Zenodo-Dep="7571582" LSID="urn:lsid:plazi:treatment:43284262FF82FFC15AB6F8E445ACCD91" httpUri="http://treatment.plazi.org/id/43284262FF82FFC15AB6F8E445ACCD91" lastPageId="31" pageId="29">
<subSubSection box="[136,979,1857,1884]" pageId="29" type="nomenclature">
<paragraph blockId="29.[136,1416,1857,1919]" box="[136,979,1857,1884]" pageId="29">
<emphasis bold="true" box="[136,157,1857,1883]" pageId="29">6.</emphasis>
<taxonomicName ID-CoL="3SCHV" authority="Pitard (1924: 390)" authorityName="Pitard" authorityPageNumber="390" authorityYear="1924" box="[164,625,1857,1884]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="29" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis bold="true" box="[164,412,1857,1884]" italics="true" pageId="29">Lasianthus caeruleus</emphasis>
<bibRefCitation author="Pitard, J." box="[419,625,1857,1883]" pageId="29" pagination="395" refId="ref55116" refString="Pitard, J. (1924) Rubiacees In: Lecomte, P. H., Humbert, H. &amp; Gagnepain, F. (Eds.) Flore generale de l'Indo-Chine 3: 395. Masson, Paris, pp. 20 - 442." type="journal article" year="1924">Pitard (1924: 390)</bibRefCitation>
</taxonomicName>
<figureCitation box="[655,732,1857,1883]" captionStart="FIGURE 21" captionStartId="30.[136,229,1929,1951]" captionTargetBox="[167,1418,358,1901]" captionTargetId="figure-86@30.[165,1418,356,1905]" captionTargetPageId="30" captionText="FIGURE 21. Lasianthus caeruleus Pit. (AG from Dang 411 and HJ from Dang 407, VNM and the herbarium of Iriomote Station (Japan)): A. Habit, B. Fruiting branch, C. Stipules, D. Abaxial leaf surfaces, EG. Inflorescence showing lateral view flowers and buds, HJ. Young fruits." figureDoi="http://doi.org/10.5281/zenodo.7571612" httpUri="https://zenodo.org/record/7571612/files/figure.png" pageId="29">Fig. 21</figureCitation>
(plate),
<figureCitation box="[826,905,1857,1883]" captionStart="FIGURE 20" captionStartId="29.[136,229,1789,1811]" captionTargetBox="[180,1403,201,1747]" captionTargetId="figure-18@29.[165,1418,193,1765]" captionTargetPageId="29" captionText="FIGURE 20. Distribution maps of Lasianthus biflorus, L. caeruleus, L. cambodianus and L. camlamensis." figureDoi="http://doi.org/10.5281/zenodo.7571608" httpUri="https://zenodo.org/record/7571608/files/figure.png" pageId="29">Fig. 20</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection lastPageId="30" pageId="29" type="materials_examined">
<paragraph blockId="29.[136,1416,1857,1919]" box="[136,1416,1897,1919]" pageId="29">
<emphasis bold="true" box="[136,194,1897,1918]" pageId="29">Type:</emphasis>
<materialsCitation box="[219,1412,1897,1919]" collectingDate="1913-12-06" collectorName="A. Chevalier" country="Vietnam" elevation="1450" location="Sapa" pageId="29" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<collectingCountry box="[219,332,1897,1918]" name="Vietnam" pageId="29">VIETNAM</collectingCountry>
.
<collectingRegion box="[341,512,1897,1919]" country="Vietnam" name="Lao Cai" pageId="29">Lao Cai Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:43284262FF82FFC15AB6F8E445ACCD91:CE5EA5AFFF82FFC35837F8CC421CCB36" box="[521,570,1897,1918]" country="Vietnam" name="Sapa" pageId="29" stateProvince="Lao Cai">Sapa</location>
,
<quantity box="[580,713,1897,1919]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="29" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
<elevation box="[580,713,1897,1919]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="29" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">14001500 m</elevation>
</quantity>
elev.,
<date box="[775,947,1897,1919]" pageId="29" value="1913-12-06">
<collectingDate box="[775,947,1897,1919]" pageId="29" value="1913-12-06">6 December 1913</collectingDate>
</date>
,
<emphasis box="[957,1145,1897,1918]" italics="true" pageId="29">
<collectorName box="[957,1079,1897,1918]" pageId="29">A. Chevalier</collectorName>
29398
</emphasis>
(
<typeStatus box="[1157,1243,1897,1919]" pageId="29">holotype</typeStatus>
P! [P04009929])
</materialsCitation>
.
</paragraph>
<paragraph blockId="29.[136,1452,1965,2064]" lastBlockId="30.[136,1451,158,328]" lastPageId="30" pageId="29">
<emphasis bold="true" box="[136,397,1965,1991]" pageId="29">
<collectingRegion box="[136,262,1965,1991]" pageId="29">Specimens</collectingRegion>
examined:
</emphasis>
<materialsCitation collectingDate="2019-06-26" collectingDateMax="2019-07-27" collectingDateMin="2019-06-26" collectionCode="VNM" collectorName="Dang V. S. &amp; Naiki A. &amp; Dang &amp; Hoang N. S. &amp; Japan" country="Vietnam" county="Thua Thien Hue Province" elevation="1193" lastPageId="30" location="Iriomote Station" municipality="Bach Ma National Park" pageId="29" specimenCount="1" stateProvince="Da Nang">
<collectingCountry box="[426,558,1965,1991]" name="Vietnam" pageId="29">VIETNAM</collectingCountry>
.
<emphasis bold="true" box="[572,673,1965,1991]" pageId="29">Central:</emphasis>
<collectingCounty box="[683,987,1965,1992]" pageId="29">Thua Thien Hue Province</collectingCounty>
,
<collectingMunicipality box="[1002,1279,1965,1992]" pageId="29">Bach Ma National Park</collectingMunicipality>
,
<quantity box="[1294,1382,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="29" unit="m" value="1193.0">
<elevation box="[1294,1382,1965,1992]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="29" unit="m" value="1193.0">1193 m</elevation>
</quantity>
elev.,
<date box="[136,287,2001,2027]" pageId="29" value="2019-07-26">
<collectingDate box="[136,287,2001,2027]" pageId="29" value="2019-07-26">26 July 2019</collectingDate>
</date>
,
<emphasis box="[299,821,2001,2028]" italics="true" pageId="29">
<collectorName box="[299,412,2002,2028]" pageId="29">Dang V.S.</collectorName>
&amp;
<collectorName box="[450,545,2001,2028]" pageId="29">Naiki A.</collectorName>
,
<collectorName box="[559,621,2002,2027]" pageId="29">Dang</collectorName>
407 &amp;
<collectorName box="[710,772,2002,2027]" pageId="29">Dang</collectorName>
411
</emphasis>
(
<collectionCode box="[837,907,2001,2027]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="29" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FF82FFC15AB6F8E445ACCD91:CE5EA5AFFF82FFC35E60F8744530CBA3" box="[1118,1302,2001,2027]" country="Vietnam" county="Thua Thien Hue Province" municipality="Bach Ma National Park" name="Iriomote Station" pageId="29" stateProvince="Da Nang">Iriomote Station</location>
(Japan));
<collectingRegion country="Vietnam" name="Da Nang" pageId="29">Da Nang</collectingRegion>
City,
<location LSID="urn:lsid:plazi:treatment:43284262FF82FFC15AB6F8E445ACCD91:CE5EA5AFFF82FFC35B35F853422BC458" box="[267,525,2038,2064]" country="Vietnam" county="Thua Thien Hue Province" municipality="Bach Ma National Park" name="Ba Na Nature Reserve" pageId="29" stateProvince="Da Nang">Ba Na Nature Reserve</location>
,
<quantity box="[538,624,2037,2064]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="29" unit="m" value="1480.0">
<elevation box="[538,624,2037,2064]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="29" unit="m" value="1480.0">1480 m</elevation>
</quantity>
elev.,
<date box="[700,855,2037,2064]" pageId="29" value="2019-06-26">
<collectingDate box="[700,855,2037,2064]" pageId="29" value="2019-06-26">26 June 2019</collectingDate>
</date>
,
<emphasis box="[868,1278,2037,2064]" italics="true" pageId="29">
<collectorName box="[868,980,2038,2064]" pageId="29">Dang V.S.</collectorName>
&amp;
<collectorName box="[1019,1153,2038,2064]" pageId="29">Hoang N.S.</collectorName>
, Dang 373
</emphasis>
(
<collectionCode box="[1295,1366,2037,2063]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="29" type="Herbarium">VNM</collectionCode>
); in the same location,
<quantity box="[304,389,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.489" pageId="30" unit="m" value="1489.0">
<elevation box="[304,389,158,185]" metricMagnitude="3" metricUnit="m" metricValue="1.489" pageId="30" unit="m" value="1489.0">1489 m</elevation>
</quantity>
elev.,
<date box="[461,608,158,184]" pageId="30" value="2019-07-27">
<collectingDate box="[461,608,158,184]" pageId="30" value="2019-07-27">27 July 2019</collectingDate>
</date>
,
<emphasis box="[619,981,158,185]" italics="true" pageId="30">
<collectorName box="[619,730,159,185]" pageId="30">Dang V.S.</collectorName>
&amp;
<collectorName box="[765,858,158,185]" pageId="30">Naiki A.</collectorName>
,
<collectorName box="[870,932,159,184]" pageId="30">Dang</collectorName>
420
</emphasis>
(
<collectionCode box="[995,1065,158,184]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="30" type="Herbarium">VNM</collectionCode>
, the herbarium of
<location LSID="urn:lsid:plazi:treatment:43284262FF82FFC15AB6F8E445ACCD91:CE5EA5AFFF81FFC05ECBFF3B458DCCF0" box="[1269,1451,158,184]" country="Vietnam" county="Thua Thien Hue Province" municipality="Bach Ma National Park" name="Iriomote Station" pageId="30" stateProvince="Da Nang">Iriomote Station</location>
(
<collectorName box="[144,213,194,220]" pageId="30">Japan</collectorName>
))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="31" pageId="30" type="distribution">
<paragraph blockId="30.[136,1451,158,328]" pageId="30">
<emphasis bold="true" box="[189,345,230,256]" pageId="30">Distribution:</emphasis>
—Endemic to
<collectingCountry box="[510,605,230,256]" name="Vietnam" pageId="30">Vietnam</collectingCountry>
(currently known as Sapa District in
<collectingRegion box="[1031,1231,230,257]" country="Vietnam" name="Lao Cai" pageId="30">Lao Cai Province</collectingRegion>
, Bach Ma National Park in Thua Thien Hue Province, Ba Na Nature Reserve in
<collectingRegion box="[812,913,267,293]" country="Vietnam" name="Da Nang" pageId="30">Da Nang</collectingRegion>
City, and Kon Chu Rang Nature Reserve in
<collectingRegion country="Vietnam" name="Gia Lai" pageId="30">Gia Lai Province</collectingRegion>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571612" ID-Zenodo-Dep="7571612" httpUri="https://zenodo.org/record/7571612/files/figure.png" pageId="30" startId="30.[136,229,1929,1951]" targetBox="[167,1418,358,1901]" targetPageId="30">
<paragraph blockId="30.[136,1452,1929,2023]" pageId="30">
<emphasis bold="true" box="[136,268,1929,1951]" pageId="30">FIGURE 21.</emphasis>
<taxonomicName authority="Pit." authorityName="Pitard" authorityYear="1924" box="[276,525,1929,1951]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="30" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis box="[276,483,1929,1950]" italics="true" pageId="30">Lasianthus caeruleus</emphasis>
Pit.
</taxonomicName>
(AG from
<emphasis box="[651,746,1929,1951]" italics="true" pageId="30">Dang 411</emphasis>
and HJ from
<emphasis box="[900,998,1929,1951]" italics="true" pageId="30">Dang 407</emphasis>
, VNM and the herbarium of Iriomote Station (Japan)):
<emphasis bold="true" box="[228,252,1965,1986]" pageId="30">A.</emphasis>
Habit,
<emphasis bold="true" box="[324,346,1965,1986]" pageId="30">B.</emphasis>
Fruiting branch,
<emphasis bold="true" box="[514,537,1965,1987]" pageId="30">C.</emphasis>
Stipules,
<emphasis bold="true" box="[633,657,1965,1986]" pageId="30">D.</emphasis>
Abaxial leaf surfaces,
<emphasis bold="true" box="[878,930,1965,1987]" pageId="30">EG.</emphasis>
Inflorescence showing lateral view flowers and buds,
<emphasis bold="true" box="[136,184,2001,2022]" pageId="30">HJ.</emphasis>
Young fruits.
</paragraph>
</caption>
<paragraph blockId="31.[136,1452,158,473]" pageId="31">
<emphasis bold="true" box="[189,432,158,184]" pageId="31">Habitat &amp; Ecology:</emphasis>
—Growing in primary and secondary montane forests or on the edge of the forest, on gravel soil at
<quantity box="[302,462,194,221]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="31" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">12001600 m</quantity>
elev. This species was found in the forest habitats where
<taxonomicName authority="Miq. (1867: 110)" authorityName="Miq." authorityPageNumber="110" authorityYear="1867" class="Magnoliopsida" family="Fabaceae" genus="Lasitnhus" kingdom="Plantae" order="Fabales" pageId="31" phylum="Tracheophyta" rank="species" species="japonicus">
<emphasis box="[1156,1386,194,220]" italics="true" pageId="31">Lasitnhus japonicus</emphasis>
Miq. (1867: 110)
</taxonomicName>
,
<taxonomicName box="[291,415,231,257]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="31" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[291,369,231,257]" italics="true" pageId="31">Croton</emphasis>
sp.
</taxonomicName>
,
<taxonomicName authority="Poir. (1804: 298)" authorityName="Poir." authorityPageNumber="298" authorityYear="1804" box="[433,903,230,256]" class="Magnoliopsida" family="Phyllanthaceae" genus="Phyllanthus" kingdom="Plantae" order="Malpighiales" pageId="31" phylum="Tracheophyta" rank="species" species="reticulatus">
<emphasis box="[433,693,230,256]" italics="true" pageId="31">Phyllanthus reticulatus</emphasis>
Poir. (1804: 298)
</taxonomicName>
,
<taxonomicName authority="(Bureau) Corner (1962: 226)" authorityName="Corner" authorityPageNumber="226" authorityYear="1962" baseAuthorityName="Bureau" box="[920,1447,230,256]" class="Magnoliopsida" family="Moraceae" genus="Streblus" kingdom="Plantae" order="Rosales" pageId="31" phylum="Tracheophyta" rank="species" species="indicus">
<emphasis box="[920,1099,230,256]" italics="true" pageId="31">Streblus indicus</emphasis>
(Bureau) Corner (1962: 226)
</taxonomicName>
,
<taxonomicName box="[136,438,266,292]" class="Magnoliopsida" family="Melastomataceae" genus="Melastoma" kingdom="Plantae" order="Myrtales" pageId="31" phylum="Tracheophyta" rank="species" species="malabathricum">
<emphasis box="[136,438,266,292]" italics="true" pageId="31">Melastoma malabathricum</emphasis>
</taxonomicName>
L. (1753: 390),
<taxonomicName box="[628,819,266,293]" class="Magnoliopsida" family="Annonaceae" genus="Anomianthus" kingdom="Plantae" order="Magnoliales" pageId="31" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[628,776,266,292]" italics="true" pageId="31">Anomianthus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName authority="L. T. Lu &amp; Boufford (2003: 255)" authorityName="L. T. Lu &amp; Boufford" authorityPageNumber="255" authorityYear="2003" box="[833,1451,266,293]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="31" phylum="Tracheophyta" rank="species" species="pluribracteatus">
<emphasis box="[833,1083,266,292]" italics="true" pageId="31">Rubus pluribracteatus</emphasis>
L.T.Lu &amp; Boufford (2003: 255)
</taxonomicName>
are dominant.
</paragraph>
<paragraph blockId="31.[136,1452,158,473]" box="[189,882,338,365]" pageId="31">
<emphasis bold="true" box="[189,323,338,364]" pageId="31">Phenology:</emphasis>
—Flowering and fruiting from June to December.
</paragraph>
<paragraph blockId="31.[136,1452,158,473]" box="[189,603,374,403]" pageId="31">
<emphasis bold="true" box="[189,405,374,400]" pageId="31">Vernacular name:</emphasis>
—Xú hưƠng lam.
</paragraph>
<paragraph blockId="31.[136,1452,158,473]" pageId="31">
<emphasis bold="true" box="[189,265,410,436]" pageId="31">Notes:</emphasis>
<taxonomicName authorityName="Pitard" authorityYear="1924" box="[294,534,410,436]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="31" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis box="[294,534,410,436]" italics="true" pageId="31">Lasianthus caeruleus</emphasis>
</taxonomicName>
was described in 1924 from a Vietnamese specimen (
<emphasis box="[1155,1377,410,437]" italics="true" pageId="31">A. Chevalier 29398</emphasis>
). This species was characterized in the field as having yellowish-green, velvety branches and then black, pubescent ones.
</paragraph>
</subSubSection>
</treatment>
</document>