treatments-xml/data/43/28/42/43284262FF11FF4E5AB6FDEB4100CE68.xml

<document ID-DOI="10.11646/phytotaxa.581.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7571550" approvalRequired="50" approvalRequired_for_taxonomicNames="15" approvalRequired_for_textStreams="30" approvalRequired_for_treatments="5" checkinTime="1674712547726" checkinUser="plazi" docAuthor="Dang, Van-Son &amp; Naiki, Akiyo" docDate="2023" docId="43284262FF11FF4E5AB6FDEB4100CE68" docLanguage="en" docName="phytotaxa.581.1.1.pdf" docOrigin="Phytotaxa 581 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.1:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="1" docTitle="Lasianthus yersinii V. S. Dang &amp; Naiki 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="1935" masterDocId="BF113A1AFF9FFFDE5A3EFFA54026CC48" masterDocTitle="A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam" masterLastPageNumber="1935" masterPageNumber="1921" pageNumber="1921" updateTime="1674713874756" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the genus Lasianthus Jack (Rubiaceae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dang, Van-Son</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8681-4141</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. &amp; dvsonitb @ gmail. com; https: // orcid. org / 0000 - 0001 - 8681 - 4141</mods:affiliation>
<mods:nameIdentifier type="email">dvsonitb@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naiki, Akiyo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3235-2823</mods:nameIdentifier>
<mods:affiliation>Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan. &amp; naiki @ lab. u-ryukyu. ac. jp; https: // orcid. org / 0000 - 0002 - 3235 - 2823 * Author for correspondence</mods:affiliation>
<mods:nameIdentifier type="email">naiki@lab.u-ryukyu.ac.jp</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>581</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1921</mods:start>
<mods:end>1935</mods:end>
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<mods:url>http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.581.1.1</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7571550</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7571904" ID-Zenodo-Dep="7571904" LSID="urn:lsid:plazi:treatment:43284262FF11FF4E5AB6FDEB4100CE68" httpUri="http://treatment.plazi.org/id/43284262FF11FF4E5AB6FDEB4100CE68" lastPageId="144" pageId="142">
<subSubSection box="[136,1093,590,617]" pageId="142" type="nomenclature">
<paragraph blockId="142.[136,1453,590,760]" box="[136,1093,590,617]" pageId="142">
<emphasis bold="true" box="[136,171,590,616]" pageId="142">75.</emphasis>
<taxonomicName authority="V.S.Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" authorityYear="2023" box="[178,614,590,617]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="142" phylum="Tracheophyta" rank="species" species="yersinii" status="sp. nov.">
<emphasis bold="true" box="[178,400,590,617]" italics="true" pageId="142">Lasianthus yersinii</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
, 
<emphasis box="[626,711,591,616]" italics="true" pageId="142">
<taxonomicNameLabel box="[626,711,591,616]" pageId="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
— 
<figureCitation box="[740,831,590,616]" captionStart="FIGURE 108" captionStartId="143.[136,229,1778,1800]" captionTargetBox="[168,1414,194,1750]" captionTargetId="figure-17@143.[165,1418,193,1754]" captionTargetPageId="143" captionText="FIGURE 108. Lasianthus yersinii V.S.Dang &amp; Naiki (from holotype, VNM): A. Branch with fruits, B. Close-up of expanding young leaves, C. Stipules, D. Abaxial leaf surfaces, E. Inflorescence showing lateral view flower buds, F–G. Mature fruits, H. Seeds (A &amp; F–G: photos by Shuichiro Tagane)." figureDoi="http://doi.org/10.5281/zenodo.7571884" httpUri="https://zenodo.org/record/7571884/files/figure.png" pageId="142">Fig. 108</figureCitation>
(plate), 
<figureCitation box="[925,1019,590,616]" captionStart="FIGURE 104" captionStartId="137.[136,229,1819,1841]" captionTargetBox="[177,1407,205,1770]" captionTargetId="figure-18@137.[165,1419,193,1796]" captionTargetPageId="137" captionText="FIGURE 104. Distribution maps of Lasianthus vietnamensis, L. viridiramulis, L. vuquangensis, L. yaharae and L. yersinii." figureDoi="http://doi.org/10.5281/zenodo.7571872" httpUri="https://zenodo.org/record/7571872/files/figure.png" pageId="142">Fig. 104</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection pageId="142" type="materials_examined">
<paragraph blockId="142.[136,1453,590,760]" pageId="142">
<emphasis bold="true" box="[136,194,630,651]" pageId="142">Type:</emphasis>
— 
<materialsCitation collectingDate="1964-05" collectingDateMax="2014-11-22" collectingDateMin="1964-05" collectionCode="VNM, FU" collectorName="Toyama H. &amp; Tagane S. &amp; Dang V. S. &amp; Nagamasu H. &amp; Naiki A. &amp; Tran H. &amp; Yang C. J. &amp; Cuong N. Q. &amp; Hieu H. N. P." country="Vietnam" elevation="617" latitude="12.111048" location="Iriomote Station" longLatPrecision="1" longitude="108.983116" municipality="Hon Ba Nature Reserve" pageId="142" specimenCode="VNM00042950, VNM00042951, VNM00042952" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<collectingCountry box="[219,332,630,651]" name="Vietnam" pageId="142">VIETNAM</collectingCountry>
. 
<collectingRegion box="[341,545,630,652]" country="Vietnam" name="Khanh Hoa" pageId="142">Khanh Hoa Province</collectingRegion>
, 
<collectingMunicipality box="[555,787,630,651]" pageId="142">Hon Ba Nature Reserve</collectingMunicipality>
, 
<quantity box="[796,858,630,652]" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="142" unit="m" value="617.0">
<elevation box="[796,858,630,652]" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="142" unit="m" value="617.0">617 m</elevation>
</quantity>
elev., 
<geoCoordinate box="[920,1074,630,652]" degrees="12" direction="north" minutes="06" orientation="latitude" pageId="142" precision="1" seconds="39.77" value="12.111048">12°06’39.77’’N</geoCoordinate>
, 
<geoCoordinate box="[1083,1246,630,652]" degrees="108" direction="east" minutes="58" orientation="longitude" pageId="142" precision="1" seconds="59.23" value="108.983116">108°58’59.23’’E</geoCoordinate>
, in evergreen forests, 
<date box="[189,379,666,688]" pageId="142" value="2014-11-22">
<collectingDate box="[189,379,666,688]" pageId="142" value="2014-11-22">22 November 2014</collectingDate>
</date>
, 
<emphasis italics="true" pageId="142">
<collectorName box="[390,499,666,687]" pageId="142">Toyama H.</collectorName>
, 
<collectorName box="[509,606,666,687]" pageId="142">Tagane S.</collectorName>
, 
<collectorName box="[618,717,666,687]" pageId="142">Dang V.S.</collectorName>
, 
<collectorName box="[727,863,666,687]" pageId="142">Nagamasu H.</collectorName>
, 
<collectorName box="[874,956,666,687]" pageId="142">Naiki A.</collectorName>
, 
<collectorName box="[967,1045,666,687]" pageId="142">Tran H.</collectorName>
, 
<collectorName box="[1056,1150,666,687]" pageId="142">Yang C.J.</collectorName>
with 
<collectorName box="[1205,1324,666,687]" pageId="142">Cuong N.Q.</collectorName>
, 
<collectorName box="[1335,1453,666,687]" pageId="142">Hieu H.N.P.</collectorName>
<date box="[189,252,702,723]" pageId="142" value="1964-05">
<collectingDate box="[189,252,702,723]" pageId="142" value="1964-05">V1964</collectingDate>
</date>
</emphasis>
(
<typeStatus box="[268,354,702,724]" pageId="142">holotype</typeStatus>
<collectionCode box="[364,425,702,723]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="142" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[443,604,702,724]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="142" type="Herbarium">VNM00042950</specimenCode>
]; isotypes 
<collectionCode box="[711,748,702,723]" country="China" name="Fudan University, Department of Biology" pageId="142">FU</collectionCode>
!, 
<collectionCode box="[765,826,702,723]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="142" type="Herbarium">VNM</collectionCode>
! [
<specimenCode box="[845,1002,702,724]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="142" type="Herbarium">VNM00042951</specimenCode>
, 
<specimenCode box="[1015,1172,702,724]" collectionCode="VNM" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="142" type="Herbarium">VNM00042952</specimenCode>
], the herbarium of 
<location LSID="urn:lsid:plazi:treatment:43284262FF11FF4E5AB6FDEB4100CE68:CE5EA5AFFF11FF505F67FD1B4127CEB0" country="Vietnam" latitude="12.111048" longLatPrecision="1" longitude="108.983116" municipality="Hon Ba Nature Reserve" name="Iriomote Station" pageId="142" stateProvince="Khanh Hoa">Iriomote Station</location>
(Japan))
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="142" type="diagnosis">
<paragraph blockId="142.[136,1452,806,976]" pageId="142">
<emphasis bold="true" box="[136,261,806,832]" pageId="142">Diagnosis:</emphasis>
—This new species is morphologically similar to 
<taxonomicName authority="(Blume ex DC.) Blume ex Miq. (1857: 320)" authorityName="Miq." authorityPageNumber="320" authorityYear="1857" baseAuthorityName="DC." class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="142" phylum="Tracheophyta" rank="species" species="latifolius">
<emphasis box="[839,1072,806,832]" italics="true" pageId="142">Lasianthus latifolius</emphasis>
(Blume ex DC.) Blume ex Miq. (1857: 320)
</taxonomicName>
based on leaf and inflorescences shapes, but differs from the latter in having more secondary veins (7–8 on each side of midrib vs. ca. 7 on each side of midrib), bracts 
<quantity box="[858,973,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="142" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long (vs. absent), larger calyx (
<quantity box="[1335,1452,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="142" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long vs. ca. 
<quantity box="[271,357,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="142" unit="mm" value="1.5">1.5 mm</quantity>
long for tubes and 
<quantity box="[574,688,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="142" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
long vs. ca. 
<quantity box="[829,915,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="142" unit="mm" value="2.5">2.5 mm</quantity>
long for lobes) and fruits with 5 (vs. 6) pyrenes (
<tableCitation box="[143,230,950,976]" captionStart="TABLE 3" captionStartId="142.[136,229,1022,1048]" captionText="TABLE 3. Comparison of Lasianthus yersinii with its morphologically similar species." pageId="142">Table 3</tableCitation>
).
</paragraph>
</subSubSection>
<caption box="[136,1115,1022,1048]" pageId="142" startId="142.[136,229,1022,1048]">
<paragraph blockId="142.[136,1115,1022,1048]" box="[136,1115,1022,1048]" pageId="142">
<emphasis bold="true" box="[136,257,1022,1048]" pageId="142">TABLE 3.</emphasis>
Comparison of 
<taxonomicName box="[438,652,1022,1048]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="142" phylum="Tracheophyta" rank="species" species="yersinii">
<emphasis box="[438,652,1022,1048]" italics="true" pageId="142">Lasianthus yersinii</emphasis>
</taxonomicName>
with its morphologically similar species.
</paragraph>
</caption>
<paragraph pageId="142">
<table box="[147,1426,1064,1552]" gridcols="3" gridrows="11" pageId="142">
<tr box="[147,1426,1064,1086]" gridrow="0" pageId="142">
<th box="[147,276,1064,1086]" gridcol="0" gridrow="0" pageId="142">
<emphasis bold="true" box="[147,262,1064,1086]" pageId="142">Characters</emphasis>
</th>
<th box="[435,886,1064,1086]" gridcol="1" gridrow="0" pageId="142">
<taxonomicName box="[435,535,1065,1086]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="142" phylum="Tracheophyta" rank="species" species="yersinii">
<emphasis bold="true" box="[435,535,1065,1086]" italics="true" pageId="142">L. yersinii</emphasis>
</taxonomicName>
</th>
<th box="[949,1426,1064,1086]" gridcol="2" gridrow="0" pageId="142">
<emphasis bold="true" box="[949,1064,1064,1086]" italics="true" pageId="142">L. latifolius</emphasis>
</th>
</tr>
<tr box="[147,1426,1104,1162]" gridrow="1" pageId="142">
<th box="[147,276,1104,1162]" gridcol="0" gridrow="1" pageId="142">Branches</th>
<td box="[435,886,1104,1162]" gridcol="1" gridrow="1" pageId="142">glabrous or subglabrous to sparsely strigose on young branches</td>
<td box="[949,1426,1104,1162]" gridcol="2" gridrow="1" pageId="142">glabrous except appressed-puberulous nodes</td>
</tr>
<tr box="[147,1426,1179,1237]" gridrow="2" pageId="142">
<th box="[147,276,1179,1237]" gridcol="0" gridrow="2" pageId="142">Leaf blades</th>
<td box="[435,886,1179,1237]" gridcol="1" gridrow="2" pageId="142">elliptic to elliptic-oblong, 15–20 × 4–6 cm; secondary veins 7–8 on each side of midrib</td>
<td box="[949,1426,1179,1237]" gridcol="2" gridrow="2" pageId="142">elliptic-oblong, 12–16 × 5–7 cm; secondary veins 6–7 on each side of midrib</td>
</tr>
<tr box="[147,1426,1255,1277]" gridrow="3" pageId="142">
<th box="[147,276,1255,1277]" gridcol="0" gridrow="3" pageId="142">Stipules</th>
<td box="[435,886,1255,1277]" gridcol="1" gridrow="3" pageId="142">2–3 mm long</td>
<td box="[949,1426,1255,1277]" gridcol="2" gridrow="3" pageId="142">ca. 3 mm long</td>
</tr>
<tr box="[147,1426,1294,1316]" gridrow="4" pageId="142">
<th box="[147,276,1294,1316]" gridcol="0" gridrow="4" pageId="142">Bracts</th>
<td box="[435,886,1294,1316]" gridcol="1" gridrow="4" pageId="142">1–1.5 mm long</td>
<td box="[949,1426,1294,1316]" gridcol="2" gridrow="4" pageId="142">absent</td>
</tr>
<tr box="[147,1426,1333,1355]" gridrow="5" pageId="142">
<th box="[147,276,1333,1355]" gridcol="0" gridrow="5" pageId="142">Calyx tubes</th>
<td box="[435,886,1333,1355]" gridcol="1" gridrow="5" pageId="142">1.5–2 mm long</td>
<td box="[949,1426,1333,1355]" gridcol="2" gridrow="5" pageId="142">ca. 1.5 mm long</td>
</tr>
<tr box="[147,1426,1372,1395]" gridrow="6" pageId="142">
<th box="[147,276,1372,1395]" gridcol="0" gridrow="6" pageId="142">Calyx lobes</th>
<td box="[435,886,1372,1395]" gridcol="1" gridrow="6" pageId="142">2.5–4 mm long</td>
<td box="[949,1426,1372,1395]" gridcol="2" gridrow="6" pageId="142">ca. 2.5 mm long</td>
</tr>
<tr box="[147,1426,1412,1434]" gridrow="7" pageId="142">
<th box="[147,276,1412,1434]" gridcol="0" gridrow="7" pageId="142">Corolla tubes</th>
<td box="[435,886,1412,1434]" gridcol="1" gridrow="7" pageId="142">unknown</td>
<td box="[949,1426,1412,1434]" gridcol="2" gridrow="7" pageId="142">6–8 mm long</td>
</tr>
<tr box="[147,1426,1451,1473]" gridrow="8" pageId="142">
<th box="[147,276,1451,1473]" gridcol="0" gridrow="8" pageId="142">Corolla lobes</th>
<td box="[435,886,1451,1473]" gridcol="1" gridrow="8" pageId="142">unknown</td>
<td box="[949,1426,1451,1473]" gridcol="2" gridrow="8" pageId="142">6–8 mm long</td>
</tr>
<tr box="[147,1426,1490,1513]" gridrow="9" pageId="142">
<th box="[147,276,1490,1513]" gridcol="0" gridrow="9" pageId="142">Fruits</th>
<td box="[435,886,1490,1513]" gridcol="1" gridrow="9" pageId="142">5–6 mm in diam.</td>
<td box="[949,1426,1490,1513]" gridcol="2" gridrow="9" pageId="142">ca. 5 mm in diam.</td>
</tr>
<tr box="[147,1426,1530,1552]" gridrow="10" pageId="142">
<th box="[147,276,1530,1552]" gridcol="0" gridrow="10" pageId="142">Pyrenes</th>
<td box="[435,886,1530,1552]" gridcol="1" gridrow="10" pageId="142">5</td>
<td box="[949,1426,1530,1552]" gridcol="2" gridrow="10" pageId="142">6</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="142" type="description">
<paragraph blockId="142.[136,1452,1635,2058]" pageId="142">
<emphasis bold="true" box="[189,337,1635,1661]" pageId="142">Description:</emphasis>
—Shrubs, 
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tall; branches and branchlets terete, 
<quantity box="[960,1056,1636,1662]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="142" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
diam., glabrous or subglabrous to sparsely strigose on young branches, internodes 
<quantity box="[674,757,1671,1698]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="142" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
long. Leaves opposite, elliptic to elliptic-oblong, 15–20 × 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="142" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
, coriaceous, glabrous on both surfaces except sparsely appressed-pubescent midrib and secondary vein abaxially, apex acuminate to cuspidate, aristate 
<quantity box="[580,699,1743,1770]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="142" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, base acute to cuneate, margin entire; midrib conspicuous adaxially, prominent abaxially; secondary veins 7–8 on each side of midrib, ascending at an angle of 50–60° from midrib, curved to the margin, flat to slightly adaxially, distinct abaxially; tertiary veins parallel; petioles 
<quantity box="[1329,1452,1815,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="142" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
long, sparsely appressed-pubescent. Stipules triangular, 
<quantity box="[778,874,1852,1878]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="142" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, appressed-pubescent. Inflorescences sessile, cymose, 1–7-flowered; bracts small, linear, 
<quantity box="[624,737,1887,1914]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="142" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long, appressed-pubescent. Flowers sessile; calyx campanulate, 
<quantity box="[136,230,1924,1950]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="142" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long, whitish pale to whitish purple, pubescent, calyx tube 
<quantity box="[907,1022,1923,1950]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="142" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long, calyx lobes 5, linear, 
<quantity box="[1337,1452,1923,1950]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="142" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
long; corolla not seen. Fruits drupaceous, globose, 
<quantity box="[715,809,1960,1986]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="142" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
long, 
<quantity box="[882,976,1959,1986]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="142" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
in diam., glabrous, crowned by persistent calyx lobes, orange when ripe; pyrenes 5. Seeds ovoid, 
<quantity box="[758,851,1996,2022]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="142" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, sulcate on the abaxial face, brown when dry.
</paragraph>
</subSubSection>
<subSubSection lastPageId="143" pageId="142" type="distribution">
<paragraph blockId="142.[136,1452,1635,2058]" box="[189,1367,2031,2058]" pageId="142">
<emphasis bold="true" box="[189,345,2031,2057]" pageId="142">Distribution:</emphasis>
— 
<collectingCountry box="[373,468,2032,2058]" name="Vietnam" pageId="142">Vietnam</collectingCountry>
(currently known only from the Hon Ba Nature Reserve, 
<collectingRegion box="[1115,1356,2032,2058]" country="Vietnam" name="Khanh Hoa" pageId="142">Khanh Hoa Province</collectingRegion>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7571884" ID-Zenodo-Dep="7571884" httpUri="https://zenodo.org/record/7571884/files/figure.png" pageId="143" startId="143.[136,229,1778,1800]" targetBox="[168,1414,194,1750]" targetPageId="143">
<paragraph blockId="143.[136,1452,1778,1873]" pageId="143">
<emphasis bold="true" box="[136,279,1778,1800]" pageId="143">FIGURE 108.</emphasis>
<taxonomicName authority="V. S. Dang &amp; Naiki" authorityName="V. S. Dang &amp; Naiki" box="[287,660,1779,1801]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="143" phylum="Tracheophyta" rank="species" species="yersinii">
<emphasis box="[287,473,1779,1800]" italics="true" pageId="143">Lasianthus yersinii</emphasis>
V.S.Dang &amp; Naiki
</taxonomicName>
(from holotype, VNM): 
<emphasis bold="true" box="[907,931,1778,1799]" pageId="143">A.</emphasis>
Branch with fruits, 
<emphasis bold="true" box="[1131,1153,1779,1800]" pageId="143">B.</emphasis>
Close-up of expanding young leaves, 
<emphasis bold="true" box="[208,231,1814,1836]" pageId="143">C.</emphasis>
Stipules, 
<emphasis bold="true" box="[325,349,1815,1836]" pageId="143">D.</emphasis>
Abaxial leaf surfaces, 
<emphasis bold="true" box="[566,589,1815,1836]" pageId="143">E.</emphasis>
Inflorescence showing lateral view flower buds, 
<emphasis bold="true" box="[1058,1109,1814,1836]" pageId="143">F–G.</emphasis>
Mature fruits, 
<emphasis bold="true" box="[1252,1278,1815,1836]" pageId="143">H.</emphasis>
Seeds (
<emphasis bold="true" box="[1352,1369,1814,1835]" pageId="143">A</emphasis>
&amp; 
<emphasis bold="true" box="[1399,1444,1814,1836]" pageId="143">F–G</emphasis>
: photos by Shuichiro Tagane).
</paragraph>
</caption>
<paragraph blockId="143.[136,1452,1919,2090]" pageId="143">
<emphasis bold="true" box="[189,420,1919,1945]" pageId="143">Habitat &amp; Ecology:</emphasis>
—The species was growing under the shade of the evergreen forest at about 
<quantity box="[1261,1387,1919,1946]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="143" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500–700 m</quantity>
elev., where 
<taxonomicName box="[213,399,1955,1981]" class="Gnetopsida" family="Gnetaceae" genus="Gnetum" kingdom="Plantae" order="Gnetales" pageId="143" phylum="Tracheophyta" rank="species" species="gnemon">
<emphasis box="[213,399,1955,1981]" italics="true" pageId="143">Gnetum gnemon</emphasis>
</taxonomicName>
L. (1767: 125), 
<taxonomicName authority="(Roxb.) Merr. (1952: 299)" authorityName="Merr." authorityPageNumber="299" authorityYear="1952" baseAuthorityName="Roxb." box="[587,1117,1955,1982]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="143" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis box="[587,809,1955,1981]" italics="true" pageId="143">Lasianthus hirsutus</emphasis>
(Roxb.) Merr. (1952: 299)
</taxonomicName>
, 
<taxonomicName authority="Poir. (1804: 298)" authorityName="Poir." authorityPageNumber="298" authorityYear="1804" class="Magnoliopsida" family="Phyllanthaceae" genus="Phyllanthus" kingdom="Plantae" order="Malpighiales" pageId="143" phylum="Tracheophyta" rank="species" species="reticulatus">
<emphasis box="[1130,1391,1955,1981]" italics="true" pageId="143">Phyllanthus reticulatus</emphasis>
Poir. (1804: 298)
</taxonomicName>
, 
<taxonomicName authority="(Bureau) Corner (1962: 226)" authorityName="Corner" authorityPageNumber="226" authorityYear="1962" baseAuthorityName="Bureau" box="[284,797,1991,2017]" class="Magnoliopsida" family="Moraceae" genus="Streblus" kingdom="Plantae" order="Rosales" pageId="143" phylum="Tracheophyta" rank="species" species="indicus">
<emphasis box="[284,463,1991,2017]" italics="true" pageId="143">Streblus indicus</emphasis>
(Bureau) Corner (1962: 226)
</taxonomicName>
and 
<taxonomicName box="[854,1087,1991,2017]" class="Magnoliopsida" family="Hamamelidaceae" genus="Eustigma" kingdom="Plantae" order="Saxifragales" pageId="143" phylum="Tracheophyta" rank="species" species="honbaense">
<emphasis box="[854,1087,1991,2017]" italics="true" pageId="143">Eustigma honbaense</emphasis>
</taxonomicName>
H. 
<collectingRegion box="[1119,1215,1992,2018]" country="Japan" name="Toyama" pageId="143">Toyama</collectingRegion>
, Tagane &amp; V.S. 
<bibRefCitation author="Dang, V. S. &amp; Tagane, S. &amp; Toyama, H. &amp; Naiki, A. &amp; Nagamasu, H. &amp; Yahara, T." pageId="143" pagination="263 - 266" refId="ref53867" refString="Dang, V. S., Tagane, S., Toyama, H., Naiki, A., Nagamasu, H. &amp; Yahara, T. (2016) Lasianthus honbaensis (Rubiaceae), a new species from southern Vietnam. Annales Botanici Fennici 53: 263 - 266. https: // doi. org / 10.5735 / 085.053.0414" type="journal article" year="2016">Dang (2016: 49)</bibRefCitation>
are dominant.
</paragraph>
<paragraph blockId="143.[136,1452,1919,2090]" box="[189,986,2063,2090]" pageId="143">
<emphasis bold="true" box="[189,323,2063,2089]" pageId="143">Phenology:</emphasis>
—Flowering and fruiting specimens were collected in July.
</paragraph>
</subSubSection>
<subSubSection pageId="144" type="etymology">
<paragraph blockId="144.[136,1452,158,544]" pageId="144">
<emphasis bold="true" box="[189,328,158,184]" pageId="144">Etymology:</emphasis>
—The specific epithet refers to Dr. Alexandre Emile Jean Yersin, who first discovered the top of Hòn Bà mountain.
</paragraph>
<paragraph blockId="144.[136,1452,158,544]" box="[189,629,230,259]" pageId="144">
<emphasis bold="true" box="[189,405,230,256]" pageId="144">Vernacular name:</emphasis>
—Xú hưƠng yersin.
</paragraph>
<paragraph blockId="144.[136,1452,158,544]" pageId="144">
<emphasis bold="true" box="[189,636,266,292]" pageId="144">Preliminary conservation assessment:</emphasis>
— 
<taxonomicName box="[664,876,266,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="144" phylum="Tracheophyta" rank="species" species="yersinii">
<emphasis box="[664,876,266,292]" italics="true" pageId="144">Lasianthus yersinii</emphasis>
</taxonomicName>
was collected from two populations with fewer than 200 individuals in the evergreen forest of Hon Ba Nature Reserve, where is well protected. The Extent of Occurrence (EOO) is less than 
<quantity box="[345,429,338,364]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="144" unit="km" value="200.0">200 km</quantity>
<superScript attach="left" box="[429,437,338,352]" fontSize="6" pageId="144">2</superScript>
and the known Area of Occupancy (AOO) is less than 
<quantity box="[1045,1115,338,364]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="144" unit="km" value="40.0">40 km</quantity>
<superScript attach="left" box="[1115,1123,338,352]" fontSize="6" pageId="144">2</superScript>
. However, the natural habitat of this species is close to the road and easily affected by anthropogenic activities. Therefore, we assigned a preliminary status of this species as Vulnerable (VU) based on the IUCN (2019) guidelines.
</paragraph>
<paragraph blockId="144.[136,1452,158,544]" pageId="144">
<emphasis bold="true" box="[189,265,446,472]" pageId="144">Notes:</emphasis>
— 
<taxonomicName box="[294,510,446,472]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="144" phylum="Tracheophyta" rank="species" species="yersinii">
<emphasis box="[294,510,446,472]" italics="true" pageId="144">Lasianthus yersinii</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName authority="Dunn (1912: 367)" authorityName="Dunn" authorityPageNumber="367" authorityYear="1912" box="[724,1078,446,473]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="144" phylum="Tracheophyta" rank="species" species="calycinus">
<emphasis box="[724,862,446,472]" italics="true" pageId="144">L. calycinus</emphasis>
Dunn (1912: 367)
</taxonomicName>
which is endemic to 
<collectingCountry box="[1332,1402,446,472]" name="China" pageId="144">China</collectingCountry>
, but differs from the latter in having longer petioles (
<quantity box="[673,795,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="144" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
vs. 
<quantity box="[840,947,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="144" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long), bracts 
<quantity box="[1099,1213,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="144" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
long (vs. absent) and smaller calyx.
</paragraph>
</subSubSection>
</treatment>
</document>