141 lines
19 KiB
XML
141 lines
19 KiB
XML
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<mods:title id="7CDA943D7184837458E2921A4D658B2F">Agra Fabricius (Coleoptera: Carabidae: Harpalinae: Lebiini: Agrina), Arboreal Beetles of Neotropical Forests: The Rare, Non-Neotropical Texas Species at the Generic Northern Limit, with Notes on Their Way of Life</mods:title>
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<treatment id="2F1787E8FFD88274FEBEFE8E8F4DF3D5" ID-DOI="http://doi.org/10.5281/zenodo.5157167" ID-GBIF-Taxon="183910539" ID-Zenodo-Dep="5157167" LSID="urn:lsid:plazi:treatment:2F1787E8FFD88274FEBEFE8E8F4DF3D5" httpUri="http://treatment.plazi.org/id/2F1787E8FFD88274FEBEFE8E8F4DF3D5" lastPageNumber="641" pageId="2" pageNumber="641">
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<paragraph id="A70136FEFFD88274FEBEFE8E8C0CF65E" blockId="2.[273,559,326,374]" box="[273,559,326,348]" pageId="2" pageNumber="641">
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<heading id="FC498192FFD88274FEBEFE8E8C0CF65E" box="[273,559,326,348]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="641" reason="2">
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<emphasis id="95CAEAECFFD88274FEBEFE8E8C0CF65E" bold="true" box="[273,559,326,348]" pageId="2" pageNumber="641">
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Genus
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<taxonomicName id="60BE4D7DFFD88274FEF6FE8E8C0CF65E" ID-CoL="8KV2T" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[345,559,326,348]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="641" phylum="Arthropoda" rank="genus">
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<emphasis id="95CAEAECFFD88274FEF6FE8E8FA9F659" bold="true" box="[345,394,326,348]" italics="true" pageId="2" pageNumber="641">Agra</emphasis>
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<bibRefCitation id="C32F4B0FFFD88274FE3DFE8E8C0CF65E" author="Fabricius, J. C." box="[402,559,326,347]" pageId="2" pageNumber="641" refId="ref6193" refString="Fabricius, J. C. 1801. Systema eleutheratorum: secundum ordines, genera, species adiecitis symomymis, locis, observationibus, descriptionibus. Tome 1, xxiv + 506 pp; Tome 2, 687 pp. Bibliopolii Academici Novi, Kiliae." type="book" year="1801">Fabricius, 1801</bibRefCitation>
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</emphasis>
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<paragraph id="A70136FEFFD88274FE83FEA98C30F673" blockId="2.[273,559,326,374]" box="[300,531,353,374]" pageId="2" pageNumber="641">Elegant Canopy Beetles</paragraph>
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</subSubSection>
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<subSubSection id="EFA46575FFD88274FF02FE438FC2F55F" pageId="2" pageNumber="641" type="reference_group">
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<paragraph id="A70136FEFFD88274FF02FE438F4DF6F5" blockId="2.[173,660,395,602]" pageId="2" pageNumber="641">
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<treatmentCitation id="261F10EFFFD88274FF02FE438F9FF6A4" author="Fabricius, J. C." box="[173,444,395,417]" page="224" pageId="2" pageNumber="641" year="1801">
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<taxonomicName id="60BE4D7DFFD88274FF02FE438F9FF6A4" ID-CoL="8KV2T" authority="Fabricius 1801: 224" authorityName="Fabricius" authorityPageNumber="224" authorityYear="1801" box="[173,444,395,417]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="641" phylum="Arthropoda" rank="genus">
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<emphasis id="95CAEAECFFD88274FF02FE438EFEF6A4" box="[173,221,395,417]" italics="true" pageId="2" pageNumber="641">Agra</emphasis>
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<bibRefCitation id="C32F4B0FFFD88274FF45FE448F9FF6A4" author="Fabricius, J. C." box="[234,444,396,417]" pageId="2" pageNumber="641" refId="ref6193" refString="Fabricius, J. C. 1801. Systema eleutheratorum: secundum ordines, genera, species adiecitis symomymis, locis, observationibus, descriptionibus. Tome 1, xxiv + 506 pp; Tome 2, 687 pp. Bibliopolii Academici Novi, Kiliae." type="book" year="1801">Fabricius 1801: 224</bibRefCitation>
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.
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<typeStatus id="7805885CFFD88274FE62FE448FDFF6A4" box="[461,508,396,417]" pageId="2" pageNumber="641">Type</typeStatus>
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species:
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<treatmentCitation id="261F10EFFFD88274FDCCFE438FF4F6BE" author="Fabricius, J. C." page=")" pageId="2" pageNumber="641" year="1801">
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<taxonomicName id="60BE4D7DFFD88274FDCCFE438FF4F6BE" ID-CoL="B2FM" authority="Fabricius (1801: 224)" authorityName="Fabricius" authorityPageNumber="224" authorityYear="1801" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="641" phylum="Arthropoda" rank="species" species="aenea">
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<emphasis id="95CAEAECFFD88274FDCCFE438F23F6BE" italics="true" pageId="2" pageNumber="641">Agra aenea</emphasis>
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<bibRefCitation id="C32F4B0FFFD88274FEA9FE6E8FF4F6BE" author="Fabricius, J. C." box="[262,471,422,443]" pageId="2" pageNumber="641" refId="ref6193" refString="Fabricius, J. C. 1801. Systema eleutheratorum: secundum ordines, genera, species adiecitis symomymis, locis, observationibus, descriptionibus. Tome 1, xxiv + 506 pp; Tome 2, 687 pp. Bibliopolii Academici Novi, Kiliae." type="book" year="1801">Fabricius (1801: 224)</bibRefCitation>
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, named first among three species described by Fabricius. Designated by
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<bibRefCitation id="C32F4B0FFFD88274FF48FE138F49F6F5" author="Erwin, T. L." box="[231,362,475,496]" pageId="2" pageNumber="641" pagination="39 - 71" refId="ref5623" refString="Erwin, T. L. 1982. Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic Entomology 7: 39 - 71." type="journal article" year="1982">Erwin (1982)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="A70136FEFFD88274FF02FE3D8FC2F55F" blockId="2.[173,660,395,602]" pageId="2" pageNumber="641">
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<treatmentCitation id="261F10EFFFD88274FF02FE3D8F9FF50E" author="Chaudoir, M. de" box="[173,444,501,523]" page="109" pageId="2" pageNumber="641" year="1861">
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<taxonomicName id="60BE4D7DFFD88274FF02FE3D8F9FF50E" ID-CoL="8LQSB" authority="Chaudoir 1861: 109" authorityName="Chaudoir" authorityPageNumber="109" authorityYear="1861" box="[173,444,501,523]" class="Insecta" family="Carabidae" genus="Agridia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="641" phylum="Arthropoda" rank="genus">
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<emphasis id="95CAEAECFFD88274FF02FE3D8ED6F50E" box="[173,245,501,523]" italics="true" pageId="2" pageNumber="641">Agridia</emphasis>
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<bibRefCitation id="C32F4B0FFFD88274FF54FE3E8F9FF50E" author="Chaudoir, M. de" box="[251,444,502,523]" pageId="2" pageNumber="641" pagination="109 - 138" refId="ref5377" refString="Chaudoir, M. de. 1861. Revision du Genre Agra. Annales de la Societe Entomologique de France 1: 109 - 138." type="journal article" year="1861">Chaudoir 1861: 109</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="7805885CFFD88274FE6AFE3E8FD7F50E" box="[453,500,502,523]" pageId="2" pageNumber="641">Type</typeStatus>
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species:
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<treatmentCitation id="261F10EFFFD88274FDE5FE3D8C31F520" author="Chaudoir, M. de" page=")" pageId="2" pageNumber="641" year="1861">
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<taxonomicName id="60BE4D7DFFD88274FDE5FE3D8C31F520" ID-CoL="B3JJ" authority="Chaudoir (1861: 109)" authorityName="Chaudoir" authorityPageNumber="109" authorityYear="1861" class="Insecta" family="Carabidae" genus="Agridia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="641" phylum="Arthropoda" rank="species" species="platyscelis">
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<emphasis id="95CAEAECFFD88274FDE5FE3D8F09F523" italics="true" pageId="2" pageNumber="641">Agridia platyscelis</emphasis>
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<bibRefCitation id="C32F4B0FFFD88274FE98FDD88C31F520" author="Chaudoir, M. de" box="[311,530,528,549]" pageId="2" pageNumber="641" pagination="109 - 138" refId="ref5377" refString="Chaudoir, M. de. 1861. Revision du Genre Agra. Annales de la Societe Entomologique de France 1: 109 - 138." type="journal article" year="1861">Chaudoir (1861: 109)</bibRefCitation>
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</taxonomicName>
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, named first between two species described by Chaudoir. Designated by
|
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<bibRefCitation id="C32F4B0FFFD88274FEF6FD8D8FFFF55F" author="Erwin, T. L." box="[345,476,581,602]" pageId="2" pageNumber="641" pagination="39 - 71" refId="ref5623" refString="Erwin, T. L. 1982. Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic Entomology 7: 39 - 71." type="journal article" year="1982">Erwin (1982)</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EFA46575FFD88274FF6AFDBB8F4DF3D5" pageId="2" pageNumber="641" type="diagnosis">
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<paragraph id="A70136FEFFD88274FF6AFDBB8C72F385" blockId="2.[173,660,627,1232]" pageId="2" pageNumber="641">
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<emphasis id="95CAEAECFFD88274FF6AFDBB8F0FF58D" bold="true" box="[197,300,627,648]" pageId="2" pageNumber="641">Diagnosis.</emphasis>
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During the evolution toward a canopy domain and away from a likely understory sistergroup (
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<bibRefCitation id="C32F4B0FFFD88274FF5BFD608F49F5B8" author="Erwin, T. L." box="[244,362,680,701]" pageId="2" pageNumber="641" pagination="539 - 592" refId="ref5515" refString="Erwin, T. L. 1979. Thoughts on the evolutionary history of ground beetles: Hypotheses generated from comparative faunal analyses of lowland forest sites in temperate and tropical regions [pp. 539 - 592]. In: Carabid Beetles: Their Evolution, Natural History, and Classification. Proceedings of the First International Symposium of Carabidology (T. L. Erwin, G. E. Ball, D. R. Whitehead, and A. L. Halpern, editors). Dr. W. Junk b. v. Publishers, The Hague, The Netherlands." type="book chapter" year="1979">Erwin 1979</bibRefCitation>
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,
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<bibRefCitation id="C32F4B0FFFD88274FED8FD608F8AF5B8" author="Erwin, T. L." box="[375,425,680,701]" pageId="2" pageNumber="641" pagination="437 - 472" refId="ref5654" refString="Erwin, T. L. 1985. The taxon pulse: A general pattern of lineage radiation and extinction among carabid beetles [pp. 437 - 472]. In: Taxonomy, Phylogeny, and Zoogeography of Beetles and Ants: A Volume Dedicated to the Memory of Philip Jackson Darlington Jr. 1904 - 1983 (G. E. Ball, editor). Dr. W. Junk b. v. Publishers, The Hague, The Netherlands." type="book chapter" year="1985">1985</bibRefCitation>
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,
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<bibRefCitation id="C32F4B0FFFD88274FE19FD608FCAF5B8" author="Erwin, T. L." box="[438,489,680,701]" pageId="2" pageNumber="641" pagination="491 - 510" refId="ref5782" refString="Erwin, T. L. 1998. Evolution at the equator: arboreal and alticolous beetles and their taxon pulses with descriptions of a new Agra subclade and its species (Coleoptera: Carabidae: Lebiini) [pp. 491 - 510]. In: Phylogeny and Classification of Caraboidea (Coleoptera: Adephaga). Proceedings of a Symposium (28 August 1996, Florence, Italy), XX International Congress of Entomology (G. E. Ball, A. Casale, and A. Vigna Taglianti editors.). Museo Regionale Scienze Naturali- Torino, Atti. Torino, Italy. Taglianti editors.). Museo Regionale Scienze Naturali- Torino, Atti. Torino, Italy." type="proceedings paper" year="1998">1998</bibRefCitation>
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),
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<taxonomicName id="60BE4D7DFFD88274FE53FD6F8C0FF5B8" authorityName="Fabricius" authorityYear="1801" box="[508,556,679,701]" class="Insecta" family="Carabidae" genus="Agra" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="641" phylum="Arthropoda" rank="genus">
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<emphasis id="95CAEAECFFD88274FE53FD6F8C0FF5B8" box="[508,556,679,701]" italics="true" pageId="2" pageNumber="641">Agra</emphasis>
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</taxonomicName>
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adults acquired numerous generic-level autapotypic features: head elongate (cranium extended behind eyes), with prognathous mandibles, securiform ultimate labial palpomeres (
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<bibRefCitation id="C32F4B0FFFD88274FE83FCDA8F84F422" author="Erwin, T. L." box="[300,423,786,807]" pageId="2" pageNumber="641" pagination="39 - 71" refId="ref5623" refString="Erwin, T. L. 1982. Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic Entomology 7: 39 - 71." type="journal article" year="1982">Erwin 1982</bibRefCitation>
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||
), and constricted neck; prothorax elongate and tubular, pleural sutures effaced; tarsomeres dilated with setiferous pads beneath (
|
||
<bibRefCitation id="C32F4B0FFFD88274FF42FCAA8F42F472" author="Erwin, T. L." box="[237,353,866,887]" pageId="2" pageNumber="641" pagination="39 - 71" refId="ref5623" refString="Erwin, T. L. 1982. Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic Entomology 7: 39 - 71." type="journal article" year="1982">Erwin 1982</bibRefCitation>
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||
), claws explanate and pectinate; elytron with latero-basal sinus and latero-apical callus, apex medially and laterally toothed or somewhat produced, apical margin truncate, sinuate, or medially lobed; male venter variously adorned with setal or pubescent patches; female reproductive system adapted to egg-laying telescopically deep in substrate (
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||
<bibRefCitation id="C32F4B0FFFD88274FE7FFBD48C6BF334" author="Erwin, T. L." box="[464,584,1052,1073]" pageId="2" pageNumber="641" pagination="1 - 68" refId="ref5951" refString="Erwin, T. L. 2002. The beetle family Carabidae of Costa Rica: twenty-nine new species of Costa Rican Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina). Zootaxa 119: 1 - 68." type="journal article" year="2002">Erwin 2002</bibRefCitation>
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||
), stylomere 2 stout (usually and with apical ensiform setae. Defense system very large (
|
||
<bibRefCitation id="C32F4B0FFFD88274FE7FFB998C6FF363" author="Erwin, T. L." box="[464,588,1105,1126]" pageId="2" pageNumber="641" pagination="39 - 71" refId="ref5623" refString="Erwin, T. L. 1982. Agra, arboreal beetles of Neotropical forests: erythropus group systematics (Carabidae). Systematic Entomology 7: 39 - 71." type="journal article" year="1982">Erwin 1982</bibRefCitation>
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||
). Size: ABL = 6.0–29.0 mm; TW = 1.5–6.0 mm.
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||
</paragraph>
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||
<paragraph id="A70136FEFFD88274FF6AFB4E8F4DF3D5" blockId="2.[173,660,627,1232]" pageId="2" pageNumber="641">
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||
<emphasis id="95CAEAECFFD88274FF6AFB4E8ED9F39E" bold="true" box="[197,250,1158,1179]" pageId="2" pageNumber="641">Note.</emphasis>
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||
For descriptions of species-groups and subgroups previously recognized, refer to
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<bibRefCitation id="C32F4B0FFFD88274FDF6FB698EC4F3D5" author="Erwin, T. L." pageId="2" pageNumber="641" pagination="17 - 21" refId="ref5740" refString="Erwin, T. L. 1996. Arboreal beetles of Neotropical Forests: Agra Fabricius, the cayennensis complex (Coleoptera: Carabidae: Lebiini: Calleidina). Annales Zoologici Fennici 33 (1): 17 - 21." type="journal article" year="1996">Erwin (1996</bibRefCitation>
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,
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,
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<bibRefCitation id="C32F4B0FFFD88274FE80FB738F40F3D5" author="Erwin, T. L." box="[303,355,1211,1232]" pageId="2" pageNumber="641" pagination="1 - 68" refId="ref5951" refString="Erwin, T. L. 2002. The beetle family Carabidae of Costa Rica: twenty-nine new species of Costa Rican Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina). Zootaxa 119: 1 - 68." type="journal article" year="2002">2002</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |