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<document id="B9232AB906B12984C4EEAD90CFECD74B" ID-CLB-Dataset="7702" ID-DOI="10.11646/zootaxa.4941.3.6" ID-GBIF-Dataset="2520ef1f-eadd-49cd-8957-3ddb1199b29d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4595423" ID-ZooBank="9B925C69-87D3-47FB-925A-1879948C65FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615421227401" checkinUser="plazi" docAuthor="Pellinen, Markku J. &amp; Wahlberg, Niklas" docDate="2021" docId="2474F637FF80FFF2FF6CFAD4BB90F8F6" docLanguage="en" docName="zootaxa.4941.3.6.pdf" docOrigin="Zootaxa 4941 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Beanoides Pellinen &amp; Wahlberg 2021, gen. nov" docType="treatment" docVersion="7" lastPageNumber="421" masterDocId="D84D8E4FFF86FFF4FFFBFFEEBA53FFD2" masterDocTitle="Changes in the genus Beana Walker (Beaninae Zahiri &amp; Holloway) with two new species and removing Beana nitida Tams to a new genus Beanoides (gen. nov.)" masterLastPageNumber="424" masterPageNumber="415" pageNumber="421" updateTime="1698932975053" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6EEE09227E460D80F62B2DD0918CDDE2">Changes in the genus Beana Walker (Beaninae Zahiri &amp; Holloway) with two new species and removing Beana nitida Tams to a new genus Beanoides (gen. nov.)</mods:title>
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<mods:name id="334B11A93C9B452B3B90E1FE63DE5C6B" type="personal">
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<mods:namePart id="7F342F9B918812B9DF45D0D3B06ACD4D">Pellinen, Markku J.</mods:namePart>
<mods:affiliation id="F5F7995C4B37B8AEB241DFDC19A5BBBF">175, M 1 Muban Phichai, Mueang Lampang, Lampang, 52000 THAILAND.</mods:affiliation>
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<mods:namePart id="5F78E9FB0813986C29F2E9682927A958">Wahlberg, Niklas</mods:namePart>
<mods:affiliation id="BD0B1125587E9F4A2FE2601C8DE3F491">Department of Biology, Sölvegatan 37, Lund University, SE- 223 62 Lund, SWEDEN.</mods:affiliation>
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<mods:title id="E2B6F69198D0EB6E35823E1AF53AB350">Zootaxa</mods:title>
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<mods:part id="113112D34B7888B70C715BAAEA8D0FF5">
<mods:date id="46CA54159926A7DF7FBE40E7F851F951">2021</mods:date>
<mods:detail id="D12C2FF6D7DB95F5DFFA9F5464142819" type="pubDate">
<mods:number id="55CDD3292599DDFA88C13A2B902FECC8">2021-03-09</mods:number>
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<mods:detail id="52894E8E3FC1130269A6CE3D3E5D7AF4" type="volume">
<mods:number id="B5893F7D95CFDD4063E27143A7F224DE">4941</mods:number>
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<mods:number id="A825BBE1BDDF45070B584268C23529B9">3</mods:number>
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<mods:classification id="6F0534C66E53FE02848441B8B6CF16D3">journal article</mods:classification>
<mods:identifier id="438AE83B1DA6759EADDAC32BB3B776FF" type="CLB-Dataset">7702</mods:identifier>
<mods:identifier id="9EBEA35B05D20E9F22F1F64852C954EF" type="DOI">10.11646/zootaxa.4941.3.6</mods:identifier>
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<mods:identifier id="286A4D8080A6BAF750F0426CAC860D23" type="ISSN">1175-5326</mods:identifier>
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<treatment id="2474F637FF80FFF2FF6CFAD4BB90F8F6" ID-DOI="http://doi.org/10.5281/zenodo.4595481" ID-GBIF-Taxon="180586519" ID-Zenodo-Dep="4595481" LSID="urn:lsid:plazi:treatment:2474F637FF80FFF2FF6CFAD4BB90F8F6" httpUri="http://treatment.plazi.org/id/2474F637FF80FFF2FF6CFAD4BB90F8F6" lastPageNumber="421" pageId="6" pageNumber="421">
<subSubSection id="E4C714AAFF80FFF2FF6CFAD4BBA2FA87" box="[151,497,1338,1365]" pageId="6" pageNumber="421" type="nomenclature">
<paragraph id="AC624721FF80FFF2FF6CFAD4BBA2FA87" blockId="6.[151,497,1338,1402]" box="[151,497,1338,1365]" pageId="6" pageNumber="421">
<heading id="F72AF04DFF80FFF2FF6CFAD4BBA2FA87" bold="true" box="[151,497,1338,1365]" fontSize="11" level="1" pageId="6" pageNumber="421" reason="1">
<emphasis id="9EA99B33FF80FFF2FF6CFAD4BBA2FA87" bold="true" box="[151,497,1338,1365]" pageId="6" pageNumber="421">
<taxonomicName id="6BDD3CA2FF80FFF2FF6CFAD4BBD2FA86" authority="Pellinen" authorityName="Pellinen &amp; Wahlberg" authorityYear="2021" box="[151,385,1338,1365]" class="Insecta" family="Nolidae" genus="Beanoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9EA99B33FF80FFF2FF6CFAD4BB47FA87" bold="true" box="[151,276,1338,1365]" italics="true" pageId="6" pageNumber="421">Beanoides</emphasis>
Pellinen
</taxonomicName>
<taxonomicNameLabel id="859A2648FF80FFF2FE73FAD5BBA2FA87" box="[392,497,1339,1365]" pageId="6" pageNumber="421" rank="genus">gen. nov</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E4C714AAFF80FFF2FF6CFAB1B8DCF965" pageId="6" pageNumber="421" type="description">
<paragraph id="AC624721FF80FFF2FF6CFAB1BBCBFAA8" blockId="6.[151,497,1338,1402]" box="[151,408,1374,1402]" pageId="6" pageNumber="421">
(
<figureCitation id="34E65BA4FF80FFF2FF5BFAB1BABEFAAB" box="[160,237,1375,1401]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1796]" captionTargetBox="[158,1427,355,1737]" captionTargetId="figure-91@5.[149,1438,345,1749]" captionTargetPageId="5" captionText="FIGURE 4. a. Beanoides nitida male Thailand, Lampang, Chae Hom 12.ix.2015 b. male Thailand, Lampang, Muban Phichai 28.vii.2012 c. female Thailand, Lampang, Muban Phichai 10.iii.2014 d. Beana terminigera male Thailand, Chiangmai, Pa Pae 12.vii.2012 e. female Thailand, Lampang, Muban Phichai 23.vi.2017 f. female Thailand, Lampang, Muban Phichai 27.vii.2008" figureDoi="http://doi.org/10.5281/zenodo.4595447" httpUri="https://zenodo.org/record/4595447/files/figure.png" pageId="6" pageNumber="421">Fig.s 4</figureCitation>
a–c and 5 a–d)
</paragraph>
<paragraph id="AC624721FF80FFF2FF6CFA4AB878FA69" blockId="6.[151,555,1443,1468]" box="[151,555,1443,1468]" pageId="6" pageNumber="421">
<typeStatus id="7366F983FF80FFF2FF6CFA4ABA9AFA6E" box="[151,201,1444,1468]" pageId="6" pageNumber="421">Type</typeStatus>
species:
<taxonomicName id="6BDD3CA2FF80FFF2FEDDFA4AB87BFA69" authority="Tams, 1924" authorityName="Tams" authorityYear="1924" box="[294,552,1443,1467]" class="Insecta" family="Nolidae" genus="Beana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="9EA99B33FF80FFF2FEDDFA4ABBFBFA69" box="[294,424,1443,1467]" italics="true" pageId="6" pageNumber="421">Beana nitida</emphasis>
<bibRefCitation id="C84C3AD0FF80FFF2FE55FA4AB87BFA69" author="Tams, W. H. T." box="[430,552,1443,1467]" pageId="6" pageNumber="421" pagination="229 - 289" refId="ref3733" refString="Tams, W. H. T. (1924) List of the moth, collected in Siam by E. J. Godfrey B. SC., F. E. S. Journal of theNatural History Society of Siam, 6, 229 - 289." type="journal article" year="1924">Tams, 1924</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="AC624721FF80FFF2FF6CFA07B8DCF965" blockId="6.[151,1437,1513,1828]" pageId="6" pageNumber="421">
Currently
<taxonomicName id="6BDD3CA2FF80FFF2FEF1FA07BB2CF9D1" authorityName="Pellinen &amp; Wahlberg" authorityYear="2021" box="[266,383,1513,1539]" class="Insecta" family="Nolidae" genus="Beanoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="9EA99B33FF80FFF2FEF1FA07BB2CF9D1" box="[266,383,1513,1539]" italics="true" pageId="6" pageNumber="421">Beanoides</emphasis>
</taxonomicName>
is a monotypic genus, including
<taxonomicName id="6BDD3CA2FF80FFF2FD09FA04B901F9D1" box="[754,850,1513,1539]" class="Insecta" family="Nolidae" genus="Beana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="9EA99B33FF80FFF2FD09FA04B901F9D1" box="[754,850,1513,1539]" italics="true" pageId="6" pageNumber="421">B. nitida</emphasis>
</taxonomicName>
only. It belongs to
<taxonomicName id="6BDD3CA2FF80FFF2FBD0FA07BE80F9D1" authorityName="Stainton" authorityYear="1859" box="[1067,1235,1513,1539]" class="Insecta" family="Nolidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="subFamily" subFamily="Chloephorinae">Chloephorinae</taxonomicName>
, best placed close to genus
<taxonomicName id="6BDD3CA2FF80FFF2FF04F9E3BBA5F9F5" authority="Walker 1864" authorityName="Walker" authorityYear="1864" box="[255,502,1549,1575]" class="Insecta" family="Nolidae" genus="Ariolica" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="9EA99B33FF80FFF2FF04F9E3BB08F9F5" box="[255,347,1549,1575]" italics="true" pageId="6" pageNumber="421">Ariolica</emphasis>
Walker 1864
</taxonomicName>
. Externally
<taxonomicName id="6BDD3CA2FF80FFF2FD7BF9E3B8A6F9F5" authorityName="Pellinen &amp; Wahlberg" authorityYear="2021" box="[640,757,1549,1575]" class="Insecta" family="Nolidae" genus="Beanoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="9EA99B33FF80FFF2FD7BF9E3B8A6F9F5" box="[640,757,1549,1575]" italics="true" pageId="6" pageNumber="421">Beanoides</emphasis>
</taxonomicName>
is different from
<taxonomicName id="6BDD3CA2FF80FFF2FC45F9E3BE78F9F5" box="[958,1067,1549,1575]" class="Insecta" family="Nolidae" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="tribe" tribe="Ariolicini">Ariolicini</taxonomicName>
species, but male genitalia have similarities; apical form of valva, exterior basal hair-pencil (not strong in
<taxonomicName id="6BDD3CA2FF80FFF2FC3BF9DFBE66F999" authorityName="Pellinen &amp; Wahlberg" authorityYear="2021" box="[960,1077,1585,1611]" class="Insecta" family="Nolidae" genus="Beanoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="9EA99B33FF80FFF2FC3BF9DFBE66F999" box="[960,1077,1585,1611]" italics="true" pageId="6" pageNumber="421">Beanoides</emphasis>
</taxonomicName>
) in valves, aedeagus vesica with several small spines also tymbal organ is not present in the basal sternal sclerite, though there are other features that connect
<taxonomicName id="6BDD3CA2FF80FFF2FF0DF997BB38F941" authorityName="Pellinen &amp; Wahlberg" authorityYear="2021" box="[246,363,1657,1683]" class="Insecta" family="Nolidae" genus="Beanoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="9EA99B33FF80FFF2FF0DF997BB38F941" box="[246,363,1657,1683]" italics="true" pageId="6" pageNumber="421">Beanoides</emphasis>
</taxonomicName>
to
<taxonomicName id="6BDD3CA2FF80FFF2FE74F997BBB8F941" authorityName="Walker" authorityYear="1864" box="[399,491,1657,1683]" class="Insecta" family="Nolidae" genus="Ariolica" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="9EA99B33FF80FFF2FE74F997BBB8F941" box="[399,491,1657,1683]" italics="true" pageId="6" pageNumber="421">Ariolica</emphasis>
</taxonomicName>
like the narrow basal sternal sclerite with somewhat longer and narrower apodemes on either side of a generally concave margin.
</paragraph>
</subSubSection>
<subSubSection id="E4C714AAFF80FFF2FF3CF92FBB90F8F6" pageId="6" pageNumber="421" type="diagnosis">
<paragraph id="AC624721FF80FFF2FF3CF92FBB90F8F6" blockId="6.[151,1437,1513,1828]" pageId="6" pageNumber="421">
<emphasis id="9EA99B33FF80FFF2FF3CF92FB855F909" bold="true" box="[199,518,1729,1755]" pageId="6" pageNumber="421">Diagnosis and Description.</emphasis>
The new genus resembles
<taxonomicName id="6BDD3CA2FF80FFF2FCCBF92CBE51F909" box="[816,1026,1730,1755]" class="Insecta" family="Nolidae" genus="Beana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="species" species="terminigera">
<emphasis id="9EA99B33FF80FFF2FCCBF92CBE51F909" box="[816,1026,1730,1755]" italics="true" pageId="6" pageNumber="421">Beana terminigera</emphasis>
</taxonomicName>
(
<figureCitation id="34E65BA4FF80FFF2FBEAF92FBE0EF909" box="[1041,1117,1729,1755]" captionStart="FIGURE 4" captionStartId="5.[151,250,1772,1796]" captionTargetBox="[158,1427,355,1737]" captionTargetId="figure-91@5.[149,1438,345,1749]" captionTargetPageId="5" captionText="FIGURE 4. a. Beanoides nitida male Thailand, Lampang, Chae Hom 12.ix.2015 b. male Thailand, Lampang, Muban Phichai 28.vii.2012 c. female Thailand, Lampang, Muban Phichai 10.iii.2014 d. Beana terminigera male Thailand, Chiangmai, Pa Pae 12.vii.2012 e. female Thailand, Lampang, Muban Phichai 23.vi.2017 f. female Thailand, Lampang, Muban Phichai 27.vii.2008" figureDoi="http://doi.org/10.5281/zenodo.4595447" httpUri="https://zenodo.org/record/4595447/files/figure.png" pageId="6" pageNumber="421">Fig.s 4</figureCitation>
d–f and 5 e–h) externally by size (wingspan ~
<quantity id="6B25EAC4FF80FFF2FEADF90BBB86F8D2" box="[342,469,1765,1792]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="6" pageNumber="421" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">34–35 mm</quantity>
), although it is more variable in the forewing pattern. There is no clear sexual dimorphism in
<taxonomicName id="6BDD3CA2FF80FFF2FF04F8E7BBEFF8F1" box="[255,444,1801,1827]" class="Insecta" family="Nolidae" genus="Beanoides" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="421" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="9EA99B33FF80FFF2FF04F8E7BBEFF8F1" box="[255,444,1801,1827]" italics="true" pageId="6" pageNumber="421">Beanoides nitida</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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