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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.123.1448" ID-GBIF-Dataset="f1dbd205-6f2f-4feb-867e-d52041258d12" ID-PMC="PMC3175121" ID-Pensoft-Pub="1313-2970-123-1" ID-PubMed="21998529" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-123-1" ModsDocOrigin="ZooKeys 123" ModsDocTitle="Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia" checkinTime="1451249962699" checkinUser="pensoft" docAuthor="Rix, Michael G. &amp; Harvey, Mark S." docDate="2011" docId="2188336B0C2E920BEDE4458F04264628" docLanguage="en" docName="ZooKeys 123: 1-100" docOrigin="ZooKeys 123" docSource="http://dx.doi.org/10.3897/zookeys.123.1448" docTitle="Austrarchaea alani Kroombit Tops Assassin Spider Rix &amp; Harvey, sp. n." docType="treatment" docVersion="5" lastPageNumber="26" masterDocId="A428FFEAFD3A7353FFACFFBCFFFA6D47" masterDocTitle="Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia" masterLastPageNumber="100" masterPageNumber="1" pageNumber="24" updateTime="1668152002706" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Australian Assassins, Part I: A review of the Assassin Spiders (Araneae, Archaeidae) of mid-eastern Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>123</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>100</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.123.1448</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.123.1448</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-123-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030504" LSID="urn:lsid:zoobank.org:act:C227AAB1-B201-40F7-A94D-4390A30C8518" httpUri="http://treatment.plazi.org/id/2188336B0C2E920BEDE4458F04264628" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection pageId="23" pageNumber="24" type="nomenclature">
<paragraph pageId="23" pageNumber="24">
<taxonomicName LSID="urn:lsid:zoobank.org:act:C227AAB1-B201-40F7-A94D-4390A30C8518" authority="Rix &amp; Harvey" class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea alani" name="Kroombit Tops Assassin Spider" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="vernacular name" species="alani" vernacular="vernacular">Austrarchaea alani Kroombit Tops Assassin Spider Rix &amp; Harvey</taxonomicName>
<taxonomicNameLabel pageId="23" pageNumber="24">sp. n.</taxonomicNameLabel>
Figs 4A-E7A8A1836
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="type material">
<paragraph pageId="23" pageNumber="24">Type material.</paragraph>
<paragraph pageId="23" pageNumber="24">
Holotype male: Kroombit Tops National Park, creek crossing off Tablelands Road, Queensland, Australia,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-24.377777">24°22'40&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="150.99611">150°59'46&quot;E</geoCoordinate>
, sifting elevated leaf litter, subtropical rainforest with emergent eucalypts, 799 m, 26.X.2010, M. &amp; A. Rix (QMB S90195).
</paragraph>
<paragraph lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Paratypes: Allotype female, same data as holotype (QMB S90194); 1 female, same data as holotype (QMB S90196); 2 females and 3 juveniles, same data as holotype
<pageBreakToken pageId="24" pageNumber="25" start="start">(</pageBreakToken>
WAM T112550DNA: KT-63-F/KT-64-J/KT-65-J); 1 male and 2 juveniles, Kroombit Tops National Park, Rainforest Walk off Tablelands Road, near Munholme Creek, Queensland, Australia,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-24.413055">24°24'47&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="151.03944">151°02'22&quot;E</geoCoordinate>
, sifting elevated leaf litter, subtropical rainforest, 753 m, 26.X.2010, M. &amp; A. Rix (WAM T112551DNA: KT-66-M/KT-67-J).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="additional material (not examined)">
<paragraph pageId="24" pageNumber="25">Additional material (not examined).</paragraph>
<paragraph pageId="24" pageNumber="25">AUSTRALIA: Queensland: Kroombit Tops National Park: Lower Dry Creek, pitfall trap, rainforest, 13-18.XII.1983, G. Monteith, V. Davies, J. Gallon, G. Thompson, 1♂ (QMB S30812); Three Moon Creek, rainforest, 9-19.XII.1983, V. Davies, J. Gallon, 2♀ (QMB S30816); Beauty Spot 98, rainforest, 9-19.XII.1983, V. Davies, J. Gallon, 1♀ (QMB S30803).</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="etymology">
<paragraph pageId="24" pageNumber="25">Etymology.</paragraph>
<paragraph pageId="24" pageNumber="25">The specific epithet is a patronym in honour of Alan Rix, for his great assistance in helping to collect this species, and for a lifetime of generosity and support to the senior author.</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="25" type="diagnosis">
<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea alani" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="alani">Austrarchaea alani</taxonomicName>
can be distinguished from all other
<taxonomicName family="Archaeidae" lsidName="" pageId="24" pageNumber="25" rank="family">Archaeidae</taxonomicName>
from mid-eastern Australia except
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea aleenae" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="aleenae">Austrarchaea aleenae</taxonomicName>
by the very large, porrect tegular sclerite 3 (TS 3) (Figs 18D-F); and from
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea aleenae" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="aleenae">Austrarchaea aleenae</taxonomicName>
by the short comb of accessory setae on the male chelicerae (Fig. 18C).
</paragraph>
<paragraph pageId="24" pageNumber="25">This species can also be distinguished from other genotyped taxa from mid-eastern Australia (see Fig. 3B) by the following three unique nucleotide substitutions for COI and COII (n = 5): T(684), A(1218), C(1347).</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph pageId="24" pageNumber="25">Description.</paragraph>
<paragraph pageId="24" pageNumber="25">
Holotype male: Total length 2.69; leg I femur 2.83; F1/CL ratio 2.68. Cephalothorax reddish-brown; legs tan-brown with darker annulations; abdomen mottled grey-brown and beige, with darker reddish-brown dorsal scute and sclerites (Fig. 18B). Carapace very tall (CH/CL ratio 2.28); 1.06 long, 2.41 high, 0.97 wide;
<normalizedToken originalValue="neck">'neck'</normalizedToken>
0.49 wide; bearing two pairs of rudimentary horns; highest point of pars cephalica (HPC) near posterior margin of
<normalizedToken originalValue="head">'head'</normalizedToken>
(ratio of HPC to post-ocular length 0.85), carapace gently sloping anterior to HPC;
<normalizedToken originalValue="head">'head'</normalizedToken>
moderately elevated postero-dorsally (post-ocular ratio 0.35) (Fig. 8A). Chelicerae with short comb of accessory setae on anterior face of paturon (Fig. 18C). Abdomen 1.38 long, 0.92 wide; with three pairs of dorsal hump-like tubercles (HT 1-6); dorsal scute fused anteriorly to epigastric sclerites, extending posteriorly to first pair of hump-like tubercles; HT 3-6 each covered by separate dorsal sclerites. Unexpanded pedipalp (Figs 18D-F) with broad, obliquely-angled foliate conductor; tegular sclerite 1 (TS 1) relatively short, almost triangular, obscured by conductor in retrolateral view; TS 2 thin, spiniform, longer than TS 1; TS 2a sinuous, largely obscured by TS 2; TS 3 very large, porrect, with broadly-pointed rectangular apex projecting well beyond retro-distal rim of tegulum.
</paragraph>
<paragraph lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Allotype female: Total length 3.41; leg I femur 3.06; F1/CL ratio 2.66. Cephalothorax dark reddish-brown; legs tan-brown with darker annulations; abdomen mottled grey-brown and beige (Fig. 18A). Carapace very tall (CH/CL ratio 2.30); 1.15 long, 2.65 high, 1.04 wide;
<normalizedToken originalValue="neck">'neck'</normalizedToken>
0.56 wide; bearing two pairs of rudimentary horns; highest point of pars cephalica (HPC) near posterior margin of
<normalizedToken originalValue="head">'head'</normalizedToken>
(ratio of HPC to post-ocular length 0.81), carapace sloping gently anterior to HPC;
<normalizedToken originalValue="head">'head'</normalizedToken>
strongly elevated postero-dorsally (post-ocular ratio 0.38) (Fig. 7A). Chelicerae without accessory setae on anterior face of paturon. Abdomen 1.97 long, 1.38 wide; with three pairs of dorsal
<pageBreakToken pageId="25" pageNumber="26" start="start">hump-like</pageBreakToken>
tubercles (HT 1-6). Internal genitalia with dense cluster of ≤ 15 variably shaped spermathecae on either side of gonopore, clusters meeting near midline of genital plate (Fig. 18G); innermost (anterior) spermathecae longest, sausage-shaped, curved antero-laterally; outermost (posterior) spermathecae bulbous; other spermathecae variably pyriform, straight, directed antero-laterally.
</paragraph>
<paragraph pageId="25" pageNumber="26">Variation: Males (n=2): total length 2.26-2.69; carapace length 1.03-1.06; carapace height 2.33-2.41; CH/CL ratio 2.28 (invariable). Females (n=4): total length 3.18-3.44; carapace length 1.15-1.18; carapace height 2.58-2.82; CH/CL ratio 2.23-2.39.</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="distribution">
<paragraph pageId="25" pageNumber="26">Distribution and habitat.</paragraph>
<paragraph pageId="25" pageNumber="26">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea alani" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alani">Austrarchaea alani</taxonomicName>
is known only from rainforest habitats in the Kroombit Tops National Park of south-eastern Queensland (Fig. 36).
</paragraph>
<paragraph pageId="25" pageNumber="26">
Conservation status. This species appears to be a short-range endemic taxon (
<bibRefCitation author="Harvey, MS" journalOrPublisher="Invertebrate Systematics" pageId="39" pageNumber="40" pagination="555 - 570" title="Short-range endemism among the Australian fauna: some examples from non-marine environments." url="doi:10.1071/IS02009" volume="16" year="2002 b">Harvey 2002b</bibRefCitation>
), which although potentially restricted in distribution, is abundant within the Kroombit Tops National Park (M. Rix, pers. obs.). It is not considered to be of conservation concern.
</paragraph>
</subSubSection>
</treatment>
</document>