179 lines
18 KiB
XML
179 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="1FEAD213B864538489F3547034B0111B" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Lithocharodes dubia Irmler 2024, sp. nov." docType="treatment" docUuid="7A616F9F-4F34-40FE-820D-00CC23B6D61A" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
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<mods:mods id="E7AE7E9348C4BD0DC2946999DD037340" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="69CCF2597B75104466E927CA3DF24A83">
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<mods:title id="5926536D60B7C4786AE0341E7751A7AA">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="D27FAB17496695EC379237FF60F7879F" type="personal">
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<mods:role id="6E1B9BAD5226F58DD9DCF72E3C863294">
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<mods:roleTerm id="9185789E326D6A986AB650E8B14D39C7">Author</mods:roleTerm>
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<mods:namePart id="DBA527AC6983EEE3C28864C5623215B0">Irmler, Ulrich</mods:namePart>
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<mods:affiliation id="D5947C2A5922F9C4B4F270CCC55A1E4B">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
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<mods:nameIdentifier id="4A9B307F6F8BB2CB8EAB65935B9F7045" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
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<mods:typeOfResource id="AA563D441D46F2D053763C7164061334">text</mods:typeOfResource>
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<mods:titleInfo id="99C71467FD2F671A9AB36EAB39A1294E">
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<mods:title id="5546022FF16C1487C8401190F174D428">Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="A258C519E38EE6ADE2207A95DB758A04">
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<mods:date id="7AF405A678F58C4E7AEC001ED4767683">2024</mods:date>
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<mods:detail id="3AD60C74C2A8688515F575810AF5CE0F" type="pubDate">
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<mods:number id="3C09597985279657E13BF1BE2B13A713">2024-01-31</mods:number>
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<mods:detail id="5D1229269DE50B6A77EF9D9F6995A1FF" type="volume">
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<mods:number id="FDCE243216DC5F99B9DEA68062884783">74</mods:number>
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</mods:detail>
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<mods:detail id="D054A1E562CCFC57581612500E41D58F" type="issue">
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<mods:number id="737138192F1C1C229386788AE5B7ECF8">1</mods:number>
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<mods:extent id="2C89CF5632600F7ECE8C933BAD8132D7" unit="page">
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<mods:start id="3EF914B3610FB237FE97325B44F68206">13</mods:start>
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<mods:end id="4F3DEE68FBE1D8D8BF5D5B024534BA48">34</mods:end>
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<mods:url id="3B54C02289ED5151B14C6AC27CFC2083">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
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</mods:location>
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<mods:classification id="B4B1AAFD1B28C1DFD3904723A127C888">journal article</mods:classification>
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<mods:identifier id="9C88FBEA6312557F40B3DB322C44AB03" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
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<treatment id="1FEAD213B864538489F3547034B0111B" LSID="urn:lsid:zoobank.org:act:7A616F9F-4F34-40FE-820D-00CC23B6D61A" httpUri="http://treatment.plazi.org/id/1FEAD213B864538489F3547034B0111B" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection id="181D14BCBC54C051B133C3E459EA1B1B" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="9D3706E1C861EC4C9D413B986617B473" pageId="0" pageNumber="13">
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<taxonomicName id="50167325BF8D7DA5D9F0EFD7FE4DBCC9" LSID="https://zoobank.org/7A616F9F-4F34-40FE-820D-00CC23B6D61A" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes dubia" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="dubia" status="sp. nov.">Lithocharodes dubia</taxonomicName>
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<taxonomicNameLabel id="B7BA619DA73FE90821DAC2B08236330F" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="77655C673D26430E1540CAB35D314914" pageId="0" pageNumber="13" type="description">
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<paragraph id="268FBAEAD7552458686665B340D4DC29" pageId="0" pageNumber="13">
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<figureCitation id="70394C15C7E857B0A9F00B901E08E2CA" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. 1. Lithocharodes andersoni; 2. L. dubia; 3. L. esmeraldae; 4. L. falini. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures1-4" httpUri="https://binary.pensoft.net/fig/975715" pageId="0" pageNumber="13">Figs 2a-d</figureCitation>
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<figureCitation id="94E4832B66C22DBCB8BAC4D4BC1C23BA" captionStart="Figure 19" captionStartId="F6" captionText="Figure 19. A. Fore-body of Lithocharodes andersoni; B. L. dubia; C. L. esmeraldae; D. L. falini; E. L. hansoni; F. L. hibbsi. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure19" httpUri="https://binary.pensoft.net/fig/975720" pageId="0" pageNumber="13">, 19B</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="B4B6F9E1589B08A15CA23006A59BB1F3" pageId="0" pageNumber="13" type="materials_examined">
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<paragraph id="89A806ED71097ADFE7F5489F3CB7D298" pageId="0" pageNumber="13">Material examined.</paragraph>
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<paragraph id="00E64226299E34877AAE7E419A251DDD" pageId="0" pageNumber="13">
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<materialsCitation id="CB10E845A18C8694029F1D9A55CCB280" collectingDate="2001-06-24" collectorName="R. Anderson" country="COSTA RICA" elevation="500" latitude="8.693334" location="Osa Peninsula" longLatPrecision="21" longitude="-83.5925" specimenCount="1" stateProvince="Puntarenas" typeStatus="Holotype">
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<emphasis id="8007931E907C6E7C9E881AEA9507CBE2" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="29A6876FD2E034FC270E5D4ADC10B438" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="42CC58A1B62E5AC22A9FE2862269E891" name="Costa Rica">COSTA RICA</collectingCountry>
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male;
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<collectingRegion id="847E61105197D43D7A4B6B69B20C4B09" country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
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;
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<location id="6C485C5B97210723D32E05F5CF7F9130" LSID="urn:lsid:plazi:treatment:1FEAD213B864538489F3547034B0111B:6C485C5B97210723D32E05F5CF7F9130" country="COSTA RICA" latitude="8.693334" longLatPrecision="21" longitude="-83.5925" name="Osa Peninsula" stateProvince="Puntarenas">Osa Peninsula</location>
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;
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<locationDeviation id="BCC70DFE1B6A4D11F94CCB99D02B57D0" location="Rancho Quemado">
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<quantity id="C5267AAE87BA03E62F5361298255921B" metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
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W Rancho Quemado
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</locationDeviation>
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;
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<geoCoordinate id="D320BB1B26C4780D2784B8BCA41785C7" degrees="8" direction="north" minutes="41" orientation="latitude" precision="15" seconds="36" value="8.693334">8°41'36"N</geoCoordinate>
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,
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<geoCoordinate id="CB05E58D67ADB8797C7CA74B75692507" degrees="83" direction="west" minutes="35" orientation="longitude" precision="15" seconds="33" value="-83.5925">83°35'33"W</geoCoordinate>
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; ridge forest litter;
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<quantity id="5F8C513288D5D80F4320D15F1A07E6AB" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
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<elevation id="E030D404BB8F186D0977F0BBB28B18E9" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
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</quantity>
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elevation;
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<collectingDate id="8C7497E7A73A8EA4F6230537275139B0" value="2001-06-24">24 VI 2001</collectingDate>
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;
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<collectorName id="D73473A5837B3E0CBBDEA57AD4B7D56C">R. Anderson</collectorName>
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leg.; KNHM CR1A01 121
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</materialsCitation>
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</paragraph>
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<paragraph id="7FA49D7CF6828E508CB5CE2F6BE9F652" pageId="0" pageNumber="13">
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<materialsCitation id="B949EC5E4CD6CC9E4927FAE152858E36" collectingDate="1995-05-02" collectorName="Berlese, R. Anderson" country="COSTA RICA" elevation="600" latitude="10.966666" location="Pitilla Biological Station" longLatPrecision="21" longitude="-85.416664" specimenCount="1" specimenCount-female="1" stateProvince="Guanacaste" typeStatus="Paratypes">
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<emphasis id="D1EFF6ED68CAAB9355698097A238660D" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="3E4EEF863A1506BF535C0B2004A12396" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry id="D069ED0C4C321E0CE7C89A8A213EE83A" name="Costa Rica">COSTA RICA</collectingCountry>
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;
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<specimenCount id="C7D21D24D42644BFE5B196D502418071" count="1" type="female">1 female</specimenCount>
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;
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<collectingRegion id="19D674C067E67C1AD99C1B5D50B99370" country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
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;
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||
<location id="5C41B2EEECC4469C63F58359341BBCED" LSID="urn:lsid:plazi:treatment:1FEAD213B864538489F3547034B0111B:5C41B2EEECC4469C63F58359341BBCED" country="COSTA RICA" latitude="10.966666" longLatPrecision="21" longitude="-85.416664" name="Pitilla Biological Station" stateProvince="Guanacaste">Pitilla Biological Station</location>
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;
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<geoCoordinate id="CC35742D9B6E1334F17B9532B1708CE5" degrees="10" direction="north" minutes="58" orientation="latitude" precision="15" seconds="0" value="10.966666">10°58'0"N</geoCoordinate>
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,
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<geoCoordinate id="D79D7DDF0FBE9EC26F4CAB06019ECEF6" degrees="85" direction="west" minutes="25" orientation="longitude" precision="15" seconds="0" value="-85.416664">85°25'0"W</geoCoordinate>
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;
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<quantity id="C020ACF6B4871F5C1FA61FFA056C3C09" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
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<elevation id="302CDD86243C8276EF9E996C218CFEFA" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
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</quantity>
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elevation;
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<collectorName id="658DF382B52F92A31B8EA72819731064">Berlese</collectorName>
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leaf litter;
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<collectingDate id="8DC034651051E265FF02AF10A0FF033E" value="1995-05-02">2 May 1995</collectingDate>
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;
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<collectorName id="B58160AD404CBF7D2E0D208FE1C39E7E">R. Anderson</collectorName>
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leg.; KNHM CR1A95 4B
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D503987604FFD6BA7420985963C51881" pageId="0" pageNumber="13" type="diagnosis">
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<paragraph id="AE23E896EC245D2D059E071E4A51BDF2" pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph id="7FA951269CA531D578401BD706FC1D57" pageId="0" pageNumber="13">
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According to the small dorsal plate of the aedeagus,
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<taxonomicName id="A82B5487BC49AEB09BABD2B425AAB840" lsidName="L. dubia" pageId="0" pageNumber="13" rank="species" species="dubia">
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<emphasis id="B8C12DEBD2E490C302A9A8E3C5943177" italics="true" pageId="0" pageNumber="13">L. dubia</emphasis>
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</taxonomicName>
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must be grouped to the
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<taxonomicName id="881BB75694709A0B801620E3583A952D" lsidName="L. obscura" pageId="0" pageNumber="13" rank="species" species="obscura">
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<emphasis id="ABD117F3F3BA8718B378F078D578E217" italics="true" pageId="0" pageNumber="13">L. obscura</emphasis>
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</taxonomicName>
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-subgroup. In contrast to both other species of the subgroup,
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<taxonomicName id="DDEDDCD40777D03344840DF054FCEBF8" lsidName="L. obscura" pageId="0" pageNumber="13" rank="species" species="obscura">
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<emphasis id="C941EC93BB76789E0E37D1460E8DA5E9" italics="true" pageId="0" pageNumber="13">L. obscura</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="281A9E567360A9D01FE2AB09D98CD4B1" lsidName="L. hansoni" pageId="0" pageNumber="13" rank="species" species="hansoni">
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<emphasis id="A6D47467B46BC0BE35EF39FCDD718E46" italics="true" pageId="0" pageNumber="13">L. hansoni</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="FEB8AEB46D422B702343133839CD8E09" lsidName="L. dubia" pageId="0" pageNumber="13" rank="species" species="dubia">
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<emphasis id="6105E18E49A386681EE441BD35635F23" italics="true" pageId="0" pageNumber="13">L. dubia</emphasis>
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</taxonomicName>
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has no concavely emarginate posterior margin of the head. The sclerotised edges of the orifice are similar as in
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<taxonomicName id="60BECB5905592D32D949541D931D35A6" lsidName="L. hansoni" pageId="0" pageNumber="13" rank="species" species="hansoni">
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<emphasis id="CA089F7D22B180786BB4DB9A9737F01C" italics="true" pageId="0" pageNumber="13">L. hansoni</emphasis>
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</taxonomicName>
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. In the overall habitus,
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<taxonomicName id="CA4682697C189F713E803BA880866D65" lsidName="L. dubia" pageId="0" pageNumber="13" rank="species" species="dubia">
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<emphasis id="41C917E1D0823724837ACF127948DA7C" italics="true" pageId="0" pageNumber="13">L. dubia</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName id="93D10DD1E83C6C6D92C48AD3CFEE6DAB" lsidName="L. nigerrima" pageId="0" pageNumber="13" rank="species" species="nigerrima">
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<emphasis id="B6F1EC754B31426560EE7584906CF447" italics="true" pageId="0" pageNumber="13">L. nigerrima</emphasis>
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</taxonomicName>
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Irmler, 2021 in size, colouration, proportion of the elytra and punctation. As the dorsal plate of the aedeagus is difficult to recognise, the taxonomic situation of the species is not clear.
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</paragraph>
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</subSubSection>
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<subSubSection id="50EB25DDC0D905609A8BBCB079E31847" pageId="0" pageNumber="13" type="description">
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<paragraph id="1A82C3674DDD76F483A30D5EB84D04A9" pageId="0" pageNumber="13">Description.</paragraph>
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<paragraph id="A7503C5325B19AEF32B29025AC73F0F1" pageId="0" pageNumber="13">Length: 4.3 mm; Colouration: black; abdominal tergites II-IV slightly lighter, brown; legs yellow-brown; antennae brown.</paragraph>
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<paragraph id="E63C007AFFE7E4EAD458F7D51B0192EC" pageId="0" pageNumber="13">Head: 0.75 mm long, 0.61 mm wide; eyes moderately large, not prominent; PS:E ratio 3.6; postocular sides nearly parallel; in posterior half semi-circular; interantennal furrows short; setiferous punctation moderately deep and dense; on average, interstices between punctures at least as wide as diameter of punctures; partly wider; surface without microsculpture, polished; antennae with first antennomere elongate; approximately two thirds as long as head; second and third antennomere conical; longer than wide; combined half as long as first antennomere; following antennomeres wider than long; apically increasing in width. Pronotum: 0.87 mm long, 0.53 mm wide; widest in anterior third; 1.2 times as wide as in front of posterior angles; anterior angles widely convex; narrowed to neck; behind middle, abruptly narrower than in front; posterior sides nearly parallel; posterior angles sub-rectangular; posterior margin slightly convex; setiferous punctation deeper and denser than on head; on average, interstices between punctures half as wide as diameter of punctures; laterally partly sparser; wide mid-line impunctate; surface without microsculpture; polished. Elytra: 0.69 mm long, 0.66 mm wide; humeral angles obtuse; sides nearly parallel; posterior angles sub-rectangular; posterior margin retreated to suture; setiferous punctation deeper and denser than on pronotum; on average, interstices between punctures half as wide as diameter of punctures; partly denser; surface without microsculpture; polished. Abdomen with setiferous punctation finer and slightly sparser than on fore-body; surface without microsculpture; polished; sternite VII of male at posterior margin slightly convex with short central emargination; tergite VII of male at posterior margin nearly straight; meso-tibia with 5, meta-tibia with 2 ctenidia. Aedeagus widest in posterior half, in middle, abruptly narrowed to apex; apical part still narrower; long orifice with sclerotised inner edges; endophallus covered by moderately large spines; parameres long; bilobed; inner lobe three fourths as long as central lobe; elongate; slender; straight at base; from middle to apex curved; in basal part, three short setae; in central part with few sensillae; outer lobe transparent; half as long as inner lobe.</paragraph>
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</subSubSection>
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<subSubSection id="42DDD77E356366D123ADEE2B464BBB13" pageId="0" pageNumber="13" type="etymology">
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<paragraph id="316B9A01A682360F2FBC168610DC02A8" pageId="0" pageNumber="13">Etymology.</paragraph>
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<paragraph id="99C4B9AC17F9FB51CF16B57A34D99E7F" pageId="0" pageNumber="13">The species name derived from the same Latin word and means doubtful. It refers to the doubtful taxonomic situation of the species due to the hardly recognisable size of the aedeagal dorsal plate.</paragraph>
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</subSubSection>
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<subSubSection id="160D7F6763B614D73C15C03030A712FF" pageId="0" pageNumber="13" type="geography">
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<paragraph id="47B3B85A735F067546B5BBDE6590CD1A" pageId="0" pageNumber="13">Geography.</paragraph>
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<paragraph id="5BC443915A306DA98326E2C109469355" pageId="0" pageNumber="13">Costa Rica.</paragraph>
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</subSubSection>
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<subSubSection id="35F0935F0C5E5383DE74795DF7915922" pageId="0" pageNumber="13" type="ecology">
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<paragraph id="9FA6BED4223C81320CEE1BF525813E10" pageId="0" pageNumber="13">Ecology.</paragraph>
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<paragraph id="471C30EBCDCF6F72582C117C73E069AC" pageId="0" pageNumber="13">Montane forest litter in May and June.</paragraph>
|
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</subSubSection>
|
||
</treatment>
|
||
</document> |