184 lines
19 KiB
XML
184 lines
19 KiB
XML
<document id="572D3167320EE55F28DA3E5410A09709" ID-DOI="10.13133/2284-4880/1548" ID-ISSN="2284-4880" ID-Zenodo-Dep="12763137" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721241057433" checkinUser="felipe" docAuthor="Perissinotto, Renzo" docDate="2023" docId="1F675E35FFADFFB86E4167C7FBA3F39E" docLanguage="en" docName="FragEntomol.55.2.195-214.pdf" docOrigin="Fragmenta entomologica 55 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/1548" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Leucocelis (Leucocelis) claveaui subsp. claveaui Perissinotto, 2023, sp. et subsp. nov." docType="treatment" docVersion="2" lastPageNumber="197" masterDocId="E35E264DFFAFFFBA6D7E6613FFBBF652" masterDocTitle="New taxa of Leucocelis Burmeister, 1842 from Namibia, and revision of L. (L.) adspersa s. l. (Fabricius, 1801) (Coleoptera: Scarabaeidae, Cetoniinae)" masterLastPageNumber="214" masterPageNumber="195" pageNumber="197" updateTime="1721242607861" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="85E1252B579B3E93B1F71FF88EAC70B9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3439E727F36ED8A2FAD873D4F13803E7">
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<mods:title id="5EB011C67C3DD29C53CBBA8559800A2B">New taxa of Leucocelis Burmeister, 1842 from Namibia, and revision of L. (L.) adspersa s. l. (Fabricius, 1801) (Coleoptera: Scarabaeidae, Cetoniinae)</mods:title>
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<mods:name id="AA5A012F822A52869093A5DCAD3C32A7" type="personal">
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<mods:namePart id="8FD27C45FCE1A3F68A0D02E0EFE548CC">Perissinotto, Renzo</mods:namePart>
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<mods:typeOfResource id="656219DC624273EB396577D88BB12E36">text</mods:typeOfResource>
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<mods:titleInfo id="5EEB0E417F5B22BA6384898D0C393761">
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<mods:title id="715F5B6256F8735768ECE9B1DEC7480E">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="D02328DC7EDA8D2AC00328D19400DF36">
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<mods:date id="FB09F5C5D474BFC2B3B5F81DE153CF21">2023</mods:date>
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<mods:number id="E5B1893278CDF3A5892B25DD4406CFBD">2023-12-21</mods:number>
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<mods:number id="2E9B34BF2EC11F7B45096C5FEDD2F755">55</mods:number>
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<mods:number id="BCC8C3F2E1426596109E0E782D8AB1BC">2</mods:number>
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<mods:start id="69F36BB822B73EAEE2AA99306D653ED6">195</mods:start>
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<mods:url id="59F77AE1B73F4112BDEFA029D6C273FE">http://dx.doi.org/10.13133/2284-4880/1548</mods:url>
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<mods:classification id="2DFE213A3F67CCC691AB8C8DD4E6C61B">journal article</mods:classification>
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<mods:identifier id="5AE1A99F0394E3FCD1395B740FAE723E" type="DOI">10.13133/2284-4880/1548</mods:identifier>
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<mods:identifier id="AF63D7C42F168969163BADCDCE6E19AB" type="ISSN">2284-4880</mods:identifier>
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<treatment id="1F675E35FFADFFB86E4167C7FBA3F39E" ID-DOI="http://doi.org/10.5281/zenodo.12763145" ID-Zenodo-Dep="12763145" LSID="urn:lsid:plazi:treatment:1F675E35FFADFFB86E4167C7FBA3F39E" httpUri="http://treatment.plazi.org/id/1F675E35FFADFFB86E4167C7FBA3F39E" lastPageNumber="197" pageId="2" pageNumber="197">
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<subSubSection id="DFD4BCA8FFADFFB86E4167C7FCD5F45E" pageId="2" pageNumber="197" type="nomenclature">
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<paragraph id="9771EF23FFADFFB86E4167C7FCD5F45E" blockId="2.[831,1452,468,1485]" pageId="2" pageNumber="197">
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<heading id="CC39584FFFADFFB86E4167C7FCD5F45E" bold="true" centered="true" fontSize="10" level="2" pageId="2" pageNumber="197" reason="3">
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<emphasis id="A5BA3331FFADFFB86E4167C7FCD5F45E" bold="true" pageId="2" pageNumber="197">
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<taxonomicName id="50CE94A0FFADFFB86E4167C7FAAAF7BE" authorityName="Perissinotto" authorityYear="2023" box="[831,1297,468,492]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="subSpecies" species="claveaui" status="sp. et subsp. nov." subGenus="Leucocelis" subSpecies="claveaui">
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<emphasis id="A5BA3331FFADFFB86E4167C7FC0FF7BE" bold="true" box="[831,948,468,492]" italics="true" pageId="2" pageNumber="197">Leucocelis</emphasis>
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(
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<emphasis id="A5BA3331FFADFFB86EB967C7FB87F7BE" bold="true" box="[967,1084,468,492]" italics="true" pageId="2" pageNumber="197">Leucocelis</emphasis>
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)
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<emphasis id="A5BA3331FFADFFB8693167C7FAAAF7BE" bold="true" box="[1103,1297,468,492]" italics="true" pageId="2" pageNumber="197">claveaui claveaui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BE898E4AFFADFFB8686567C7FCD5F45E" pageId="2" pageNumber="197" rank="subSpecies">sp. et subsp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DFD4BCA8FFADFFB86E416407FC73F47E" box="[831,968,532,556]" pageId="2" pageNumber="197" type="description">
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<paragraph id="9771EF23FFADFFB86E416407FC73F47E" blockId="2.[831,1452,468,1485]" box="[831,968,532,556]" pageId="2" pageNumber="197">
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(
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<figureCitation id="0FF5F3A6FFADFFB86E396407FC31F47E" box="[839,906,532,556]" captionStart="Fig" captionStartId="3.[817,847,1080,1099]" captionTargetBox="[822,1436,174,1030]" captionTargetId="figure-637@3.[813,1439,166,1033]" captionTargetPageId="3" captionText="Fig. 1 –Leucocelis (Leucocelis) claveaui claveaui sp. et subsp. nov.: A, HT♂ dorsal habitus; B, HT♂ ventral habitus; C, PT♀ dorsal habitus; D, HT♂ lat- eral habitus; E, HT♂ clypeus; F, HT♂ pygidium; G, HT♂ parameres, dorsal view; H, HT♂ parameres, lateral view. Photos: Lynette Clennell." figureDoi="http://doi.org/10.5281/zenodo.12763139" httpUri="https://zenodo.org/record/12763139/files/figure.png" pageId="2" pageNumber="197">Figs 1</figureCitation>
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,
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<figureCitation id="0FF5F3A6FFADFFB86EE66407FC1CF47E" box="[920,935,532,556]" captionStart="Fig" captionStartId="4.[151,182,1912,1931]" captionTargetBox="[151,1451,1222,1886]" captionTargetId="figure-673@4.[151,1451,1219,1889]" captionTargetPageId="4" captionText="Fig. 2 – Erongoberge at Usakos with the typical habitat of L. claveaui claveaui sp. et subsp. nov. (Photo: https://www.facebook.com/AmeibNamibia/photos)." figureDoi="http://doi.org/10.5281/zenodo.12763143" httpUri="https://zenodo.org/record/12763143/files/figure.png" pageId="2" pageNumber="197">2</figureCitation>
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,
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<figureCitation id="0FF5F3A6FFADFFB86ECC6407FC7BF47E" box="[946,960,532,556]" captionStart="Fig" captionStartId="8.[831,862,952,971]" captionTargetBox="[831,1451,166,922]" captionTargetId="figure-739@8.[831,1451,164,924]" captionTargetPageId="8" captionText="Fig. 7 – Known distribution range of Leucocelis (Leucocelis) claveaui claveaui sp. et subsp. nov., L. (L.) claveaui otavi sp. et subsp. nov. and L. (L.) franki Janson, 1888 in South Africa and Namibia (Map: Google Earth Pro with image from Landsat/Copernicus 12/14/2015)." figureDoi="http://doi.org/10.5281/zenodo.12763160" httpUri="https://zenodo.org/record/12763160/files/figure.png" pageId="2" pageNumber="197">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DFD4BCA8FFADFFB86E416427FBA3F39E" pageId="2" pageNumber="197" type="diagnosis">
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<paragraph id="9771EF23FFADFFB86E416427FBA3F39E" blockId="2.[831,1452,468,1485]" pageId="2" pageNumber="197">
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<emphasis id="A5BA3331FFADFFB86E416427FC0FF41E" bold="true" box="[831,948,564,588]" pageId="2" pageNumber="197">Diagnosis.</emphasis>
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This new species resembles superficially darkblue specimens of
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<taxonomicName id="50CE94A0FFADFFB869766447FA39F43E" authority="Janson, 1888" authorityName="Janson Ph. Antoine" authorityYear="1888" box="[1032,1410,596,620]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
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<emphasis id="A5BA3331FFADFFB869766447FBC1F43E" box="[1032,1146,596,620]" italics="true" pageId="2" pageNumber="197">Leucocelis</emphasis>
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(
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<emphasis id="A5BA3331FFADFFB869F46446FB1BF43E" box="[1162,1184,597,620]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
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)
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<emphasis id="A5BA3331FFADFFB869CE6447FB4BF43E" box="[1200,1264,596,620]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
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<bibRefCitation id="F35F92D2FFADFFB869886446FA39F43E" author="Janson O. E." box="[1270,1410,596,620]" pageId="2" pageNumber="197" pagination="107 - 117" refId="ref18226" refString="Janson O. E. 1888. Descriptions of new Cetoniidae. Notes from the Leyden Museum 10: 107 - 117." type="journal article" year="1888">Janson, 1888</bibRefCitation>
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</taxonomicName>
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but there are clear diagnostic characters differentiating it from the more widespread latter species. In particular,
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<emphasis id="A5BA3331FFADFFB868306486FADFF4FE" box="[1358,1380,661,684]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
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(
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<emphasis id="A5BA3331FFADFFB8680D6486FA32F4FE" box="[1395,1417,661,684]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
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)
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<emphasis id="A5BA3331FFADFFB868E76486FC22F49E" italics="true" pageId="2" pageNumber="197">
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c.
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<taxonomicName id="50CE94A0FFADFFB86E4164A7FC22F49E" authorityName="Perissinotto" authorityYear="2023" box="[831,921,692,716]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="claveaui">claveaui</taxonomicName>
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</emphasis>
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exhibits a consistently black pronotum and reddish-orange pygidium, while in the dark specimens of
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<taxonomicName id="50CE94A0FFADFFB868E864C6FC14F55E" authorityName="Janson Ph. Antoine" authorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
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<emphasis id="A5BA3331FFADFFB868E864C6FA1EF4BE" box="[1430,1445,725,748]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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. (
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<emphasis id="A5BA3331FFADFFB86E3664E6FCECF55E" box="[840,855,757,780]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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.)
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<emphasis id="A5BA3331FFADFFB86E1164E7FC14F55E" box="[879,943,756,780]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
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</taxonomicName>
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the pronotum is rather dark blue or dark green and the pygidium is always black to dark brown. Light green, blue, orange or polychromatic specimens of
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<taxonomicName id="50CE94A0FFADFFB868196526FCC4F53E" authorityName="Janson Ph. Antoine" authorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
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<emphasis id="A5BA3331FFADFFB868196526FACDF51E" box="[1383,1398,821,844]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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. (
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<emphasis id="A5BA3331FFADFFB868F36526FA27F51E" box="[1421,1436,821,844]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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.)
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<emphasis id="A5BA3331FFADFFB86E416547FCC4F53E" box="[831,895,852,876]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
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</taxonomicName>
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also have a reddish-orange pygidium, but they differ drastically in many other ways from the new species that only dark specimens can potentially be confused with it. Specimens of
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<emphasis id="A5BA3331FFADFFB86E8665A6FBBCF59E" box="[1016,1031,949,972]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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. (
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<emphasis id="A5BA3331FFADFFB8696165A6FB95F59E" box="[1055,1070,949,972]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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.)
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<emphasis id="A5BA3331FFADFFB8693B65A6FB01F59E" box="[1093,1210,948,972]" italics="true" pageId="2" pageNumber="197">
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c.
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<taxonomicName id="50CE94A0FFADFFB8691E65A7FB01F59E" authorityName="Perissinotto" authorityYear="2023" box="[1120,1210,948,972]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="claveaui">claveaui</taxonomicName>
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</emphasis>
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also have their dorsal white maculae well rounded and reduced in diameter, by comparison with those of
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<taxonomicName id="50CE94A0FFADFFB8692A65E6FB66F25E" authorityName="Janson Ph. Antoine" authorityYear="1888" box="[1108,1245,1012,1036]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
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<emphasis id="A5BA3331FFADFFB8692A65E6FBD2F25E" box="[1108,1129,1013,1036]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
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(
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<emphasis id="A5BA3331FFADFFB8690665E6FB3CF25E" box="[1144,1159,1013,1036]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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.)
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<emphasis id="A5BA3331FFADFFB869E365E7FB66F25E" box="[1181,1245,1012,1036]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
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</taxonomicName>
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, which are particularly large around the apical and postero-lateral margins of their elytra. The ventral surface is markedly less setose and the mesosternal process narrower and more constrained in
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<emphasis id="A5BA3331FFADFFB86E416266FCF5F2DE" box="[831,846,1141,1164]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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||
. (
|
||
<emphasis id="A5BA3331FFADFFB86E1B6266FCCFF2DE" box="[869,884,1141,1164]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
.)
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<emphasis id="A5BA3331FFADFFB86EF56266FC2CF2DE" box="[907,919,1141,1164]" italics="true" pageId="2" pageNumber="197">c</emphasis>
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.
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<taxonomicName id="50CE94A0FFADFFB86EDA6267FC45F2DE" authorityName="Perissinotto" authorityYear="2023" box="[932,1022,1140,1164]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="claveaui">
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<emphasis id="A5BA3331FFADFFB86EDA6267FC45F2DE" box="[932,1022,1140,1164]" italics="true" pageId="2" pageNumber="197">claveaui</emphasis>
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</taxonomicName>
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than in
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<taxonomicName id="50CE94A0FFADFFB869296266FB5EF2DE" authorityName="Janson Ph. Antoine" authorityYear="1888" box="[1111,1253,1140,1164]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
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<emphasis id="A5BA3331FFADFFB869296266FBD6F2DE" box="[1111,1133,1141,1164]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
|
||
(
|
||
<emphasis id="A5BA3331FFADFFB869036266FB37F2DE" box="[1149,1164,1141,1164]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
.)
|
||
<emphasis id="A5BA3331FFADFFB869DC6267FB5EF2DE" box="[1186,1253,1140,1164]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
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</taxonomicName>
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<emphasis id="A5BA3331FFADFFB869986267FB51F2DE" box="[1254,1258,1140,1164]" italics="true" pageId="2" pageNumber="197">.</emphasis>
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The general body shape of
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<taxonomicName id="50CE94A0FFADFFB86EDB6286FB8FF2FE" authorityName="Janson Ph. Antoine" authorityYear="1888" box="[933,1076,1172,1196]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
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<emphasis id="A5BA3331FFADFFB86EDB6286FC00F2FE" box="[933,955,1173,1196]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
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(
|
||
<emphasis id="A5BA3331FFADFFB86EB26286FC60F2FE" box="[972,987,1173,1196]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
.)
|
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<emphasis id="A5BA3331FFADFFB86E8A6287FB8FF2FE" box="[1012,1076,1172,1196]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
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</taxonomicName>
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is notably stockier than that of
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<emphasis id="A5BA3331FFADFFB868E86286FA1EF2FE" box="[1430,1445,1173,1196]" italics="true" pageId="2" pageNumber="197">L</emphasis>
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. (
|
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<emphasis id="A5BA3331FFADFFB86E3662A6FCECF29E" box="[840,855,1205,1228]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
.)
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<emphasis id="A5BA3331FFADFFB86E1162A6FCC0F29E" box="[879,891,1205,1228]" italics="true" pageId="2" pageNumber="197">c</emphasis>
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.
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<emphasis id="A5BA3331FFADFFB86EF562A7FC57F29E" box="[907,1004,1204,1228]" italics="true" pageId="2" pageNumber="197">
|
||
<taxonomicName id="50CE94A0FFADFFB86EF562A7FC53F29E" authorityName="Perissinotto" authorityYear="2023" box="[907,1000,1204,1228]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="claveaui">claveaui</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with the former exhibiting a total length to maximum width ratio (TL:MW) of ≈ 2 and the latter of> 2. Concerning aedeagal parameres, in
|
||
<emphasis id="A5BA3331FFADFFB8698E62E6FB44F35E" box="[1264,1279,1269,1292]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
. (
|
||
<emphasis id="A5BA3331FFADFFB8686A62E6FA98F35E" box="[1300,1315,1269,1292]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
.)
|
||
<emphasis id="A5BA3331FFADFFB8684662E6FAFFF35E" box="[1336,1348,1269,1292]" italics="true" pageId="2" pageNumber="197">c</emphasis>
|
||
.
|
||
<taxonomicName id="50CE94A0FFADFFB8682F62E7FA10F35E" authorityName="Perissinotto" authorityYear="2023" box="[1361,1451,1268,1292]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="claveaui">
|
||
<emphasis id="A5BA3331FFADFFB8682F62E7FA10F35E" box="[1361,1451,1268,1292]" italics="true" pageId="2" pageNumber="197">claveaui</emphasis>
|
||
</taxonomicName>
|
||
the ventral lobes are laterally parallel and project forward forming a tubercle of medium size at the apex with setation on its inner declivity, while in
|
||
<taxonomicName id="50CE94A0FFADFFB869D06346FA8DF33E" authorityName="Janson Ph. Antoine" authorityYear="1888" box="[1198,1334,1364,1388]" class="Insecta" family="Scarabaeidae" genus="Leucocelis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="franki" subGenus="Leucocelis">
|
||
<emphasis id="A5BA3331FFADFFB869D06346FB7FF33E" box="[1198,1220,1365,1388]" italics="true" pageId="2" pageNumber="197">L.</emphasis>
|
||
(
|
||
<emphasis id="A5BA3331FFADFFB869AC6346FB5AF33E" box="[1234,1249,1365,1388]" italics="true" pageId="2" pageNumber="197">L</emphasis>
|
||
.)
|
||
<emphasis id="A5BA3331FFADFFB869886347FA8DF33E" box="[1270,1334,1364,1388]" italics="true" pageId="2" pageNumber="197">franki</emphasis>
|
||
</taxonomicName>
|
||
the ventral lobes are short and smoothly rounded, projecting forward barely above the dorsal lobes at the apex, which does not exhibit any setation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |