430 lines
58 KiB
XML
430 lines
58 KiB
XML
<document id="ADE4783106DAD183314593A60F314C63" ID-DOI="10.5281/zenodo.173663" ID-GBIF-Dataset="45e2592f-f329-4cda-bced-0582fee9c38e" ID-ISSN="11755326" ID-Zenodo-Dep="173663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459782631952" checkinUser="plazi" docAuthor="Francke, Oscar F. & Savary, Warren E." docDate="2006" docId="0D67EB43FFCE8063CB4232F67534278F" docLanguage="en" docName="zt01302p030.pdf" docOrigin="Zootaxa 1302" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pseudouroctonus sprousei Francke & Savary, 2006, new species" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="F15E933BFFCC806BCA4A316777252329" masterDocTitle="A new troglobitic Pseudouroctonus Stahnke (Scorpiones: Vaejovidae) from northern México" masterLastPageNumber="30" masterPageNumber="21" pageNumber="23" updateTime="1698217935919" updateUser="plazi">
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<mods:mods id="F11AA94195B0D3C9819D6D0AD24F07D1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F53FD426342D050E79749F10AA2893ED">
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<mods:title id="26050F89F1B19FC5ADEC164C353D0C85">A new troglobitic Pseudouroctonus Stahnke (Scorpiones: Vaejovidae) from northern México</mods:title>
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<mods:name id="39238472EFE9F747092F4211B40C1915" type="personal">
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<mods:roleTerm id="62F2F2E19EE9254EFA131683CC8D9AA9">Author</mods:roleTerm>
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<mods:namePart id="CD7FB8F46BFB60EB5FB6086B03B57656">Francke, Oscar F.</mods:namePart>
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<mods:name id="F6220D81216705F9D7BA2D6B0BE0F04A" type="personal">
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<mods:roleTerm id="D0391B4F5CF5ED6D328D39B9AB5B3841">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="584E1F0E8F642D90B3C0EC4761A63890">Savary, Warren E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1D1B50A484F94014D83233155FD166FB">text</mods:typeOfResource>
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<mods:title id="05EC1EEB6FB3DD540EB03823CCC09A17">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="96D0670F3CDFAF9CEE57C3DD26694963">
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<mods:date id="39DD5A9C1E773FD587C55094A80065A7">2006</mods:date>
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<mods:detail id="9AA13C40D2FB5CEAFA3A5F5954930AD7" type="volume">
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<mods:number id="BA9AE3DCD8AD4B6ACA1847DFCE7894BF">1302</mods:number>
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<mods:classification id="ACB392C3EC00CF411D6A2B654E97F60A">journal article</mods:classification>
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<mods:identifier id="E6B85E72A38D40E05153E64A027A13AD" type="DOI">10.5281/zenodo.173663</mods:identifier>
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<treatment id="0D67EB43FFCE8063CB4232F67534278F" ID-DOI="http://doi.org/10.5281/zenodo.6261576" ID-GBIF-Taxon="119348373" ID-Zenodo-Dep="6261576" LSID="urn:lsid:zoobank.org:act:1E33CE09-74B9-41A0-8E4F-5F0F8CE75CB3" httpUri="http://treatment.plazi.org/id/0D67EB43FFCE8063CB4232F67534278F" lastPageId="8" lastPageNumber="29" pageId="2" pageNumber="23">
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<subSubSection id="CDD409DEFFCE8069CB4232F676B220E7" pageId="2" pageNumber="23" type="nomenclature">
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<paragraph id="85715A55FFCE8069CB4232F675C02082" blockId="2.[264,741,913,974]" box="[264,741,913,939]" pageId="2" pageNumber="23">
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<heading id="DE39ED39FFCE8069CB4232F675C02082" bold="true" box="[264,741,913,939]" fontSize="11" level="1" pageId="2" pageNumber="23" reason="1">
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<emphasis id="B7BA8647FFCE8069CB4232F675C02082" bold="true" box="[264,741,913,939]" pageId="2" pageNumber="23">
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<emphasis id="B7BA8647FFCE8069CB4232F675682082" bold="true" box="[264,589,913,939]" italics="true" pageId="2" pageNumber="23">
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<taxonomicName id="42CE21D6FFCE8069CB4232F675622082" ID-CoL="78HDG" box="[264,583,913,939]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="sprousei" status="sp. nov.">Pseudouroctonus sprousei</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="AC893B3CFFCE8069C81F32F675C02082" box="[597,741,913,939]" pageId="2" pageNumber="23" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="85715A55FFCE8069CB4232D376B220E7" blockId="2.[264,741,913,974]" box="[264,407,948,974]" pageId="2" pageNumber="23">
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(
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<figureCitation id="1DF546D0FFCE8069CB5B32D376AB20E7" box="[273,398,948,974]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 8" captionStart-2="FIGURES 9 – 14" captionStartId-0="4.[264,368,1219,1243]" captionStartId-1="5.[264,382,1651,1675]" captionStartId-2="6.[264,382,1727,1751]" captionTargetBox-0="[539,1047,334,1194]" captionTargetBox-1="[327,1251,486,1629]" captionTargetBox-2="[334,1233,318,1701]" captionTargetId-0="figure@4.[539,1048,334,1194]" captionTargetId-1="figure@5.[327,1251,486,1629]" captionTargetId-2="figure@6.[312,1280,305,1721]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Dorsal view of holotype male of Pseudouroctonus sprousei, sp. nov." captionText-1="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." captionText-2="FIGURES 9 – 14. Morphology of Pseudouroctonus sprousei, sp. nov.: 9. Pedipalp chela, dorsal aspect; 10. Pedipalp chela, external aspect; 11. Pedipalp chela, ventrointernal aspect; 12. Telotarsus III, prolateral aspect showing setation pattern; 13. Telotarsus III, ventral aspect, schematic showing ventromedian spinules and setae; 14. Telotarsus III, retrolateral aspect (scale figs. 9 – 11 = 1 mm, figs. 12 – 14 = 0.5 mm)." httpUri-0="https://zenodo.org/record/173664/files/figure.png" httpUri-1="https://zenodo.org/record/173665/files/figure.png" httpUri-2="https://zenodo.org/record/173666/files/figure.png" pageId="2" pageNumber="23">Figs. 1–14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="CDD409DEFFCE8069CB423563752A2638" pageId="2" pageNumber="23" type="materials_examined">
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<paragraph id="85715A55FFCE8069CB423563752A2638" blockId="2.[264,1323,1028,1297]" pageId="2" pageNumber="23">
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<emphasis id="B7BA8647FFCE8069CB42356376A12737" bold="true" box="[264,388,1028,1054]" pageId="2" pageNumber="23">
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<typeStatus id="5A75E4F7FFCE8069CB42356376602737" box="[264,325,1028,1054]" pageId="2" pageNumber="23">Type</typeStatus>
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data
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</emphasis>
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.
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<typeStatus id="5A75E4F7FFCE8069CBDE356375242737" box="[404,513,1028,1054]" pageId="2" pageNumber="23" type="holotype">Holotype</typeStatus>
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male from
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<collectingCountry id="FDD91AC5FFCE8069C8C0356374272737" box="[650,770,1028,1054]" name="Mexico" pageId="2" pageNumber="23">MEXICO</collectingCountry>
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: Coahuila; El Abra (cave), El Remolino (UTM NAD27: 14R: 267620, 3180825, elev. S
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<specimenCount id="93C891DCFFCE8069C893354874082760" box="[729,813,1071,1097]" pageId="2" pageNumber="23" type="generic">
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<quantity id="4236F7B0FFCE8069C8933548740C2760" box="[729,809,1071,1097]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="23" unit="m" value="530.0">530 m</quantity>
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.
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</specimenCount>
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) [
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<geoCoordinate id="E0FA3C92FFCE8069C90E354874D42760" box="[836,1009,1068,1097]" direction="north" orientation="latitude" pageId="2" pageNumber="23" precision="1" value="28.733473">N 28o 44´0.5´´</geoCoordinate>
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W–101o 22´45´´]
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<date id="F1707C95FFCE8069CE83354876602758" pageId="2" pageNumber="23" value="2001-04-29">29 April 2001</date>
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,
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<date id="F1707C95FFCE8069CB1E353076D62758" box="[340,499,1111,1137]" pageId="2" pageNumber="23" value="2002-02-17">17 Feb. 2002</date>
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(the specimen label bears two dates, on both of which the cave was explored; the collector indicated the second date is probably when the specimen was actually collected). Peter Sprouse. Deposited in the Colección Nacional de Arácnidos (
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<collectionCode id="E3DFC290FFCE8069CB5A35A8764027C0" box="[272,357,1231,1257]" pageId="2" pageNumber="23">CNAN</collectionCode>
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T0202), Departamento de Zoología, Instituto de Biología, Univ. Nacional Autónoma de
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<collectingCountry id="FDD91AC5FFCE8069CBE43590752E2638" box="[430,523,1271,1297]" name="Mexico" pageId="2" pageNumber="23">México</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CDD409DEFFCE8069CB42342074062648" box="[264,803,1351,1377]" pageId="2" pageNumber="23" type="distribution">
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||
<paragraph id="85715A55FFCE8069CB42342074062648" blockId="2.[264,803,1351,1377]" box="[264,803,1351,1377]" pageId="2" pageNumber="23">
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<emphasis id="B7BA8647FFCE8069CB42342076842648" bold="true" box="[264,417,1351,1377]" pageId="2" pageNumber="23">Distribution</emphasis>
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. Known only from the
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<typeStatus id="5A75E4F7FFCE8069C8FE3420743A2648" box="[692,799,1351,1377]" pageId="2" pageNumber="23" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="CDD409DEFFCE8068CB4234F074AA22FF" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23" type="diagnosis">
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<paragraph id="85715A55FFCE8068CB4234F074AA22FF" blockId="2.[264,1324,1431,1897]" lastBlockId="3.[264,1323,284,470]" lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
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<emphasis id="B7BA8647FFCE8069CB4234F076A52698" bold="true" box="[264,384,1431,1457]" pageId="2" pageNumber="23">Diagnosis</emphasis>
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. Differs from all other
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<taxonomicName id="42CE21D6FFCE8069C8E934F074482699" box="[675,877,1431,1456]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="23" phylum="Arthropoda" rank="genus">
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<emphasis id="B7BA8647FFCE8069C8E934F074482699" box="[675,877,1431,1456]" italics="true" pageId="2" pageNumber="23">Pseudouroctonus</emphasis>
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</taxonomicName>
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species by having a distinct patellar internomedian carina and by its large size, with the only adult specimen known measuring
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<quantity id="4236F7B0FFCE8069CB423480767E2528" box="[264,347,1511,1537]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="23" unit="mm" value="74.0">74 mm</quantity>
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in length. Differs from
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<emphasis id="B7BA8647FFCE8069C832348075CA2529" box="[632,751,1511,1536]" italics="true" pageId="2" pageNumber="23">P. l i n d s a y i</emphasis>
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||
and the species in the former
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<emphasis id="B7BA8647FFCE8069CE113480739B2529" box="[1115,1214,1511,1536]" italics="true" pageId="2" pageNumber="23">minimus</emphasis>
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group in having a distinct white patch on the pedipalp chela finger tips, in having prominent teeth ventrally on the fixed and movable fingers of the chelicera, and in having 17 pectinal teeth in males. Further differs from the members of the former
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<emphasis id="B7BA8647FFCE8069C993373873192551" box="[985,1084,1631,1656]" italics="true" pageId="2" pageNumber="23">minimus</emphasis>
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group in lacking a distinctive tooth on the lateral lobe of the lamellar hook of the hemispermatophore, and differs from
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<taxonomicName id="42CE21D6FFCE8069CBEA37C8753925E1" box="[416,540,1711,1736]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="lindsayi">
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<emphasis id="B7BA8647FFCE8069CBEA37C8753925E1" box="[416,540,1711,1736]" italics="true" pageId="2" pageNumber="23">P. lindsayi</emphasis>
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</taxonomicName>
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in having the lamellar hook elevated from the lamella base. It is most closely related to
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<emphasis id="B7BA8647FFCE8069C81F37B075F225D9" box="[597,727,1751,1776]" italics="true" pageId="2" pageNumber="23">P. re d d e l l i</emphasis>
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, the
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<typeStatus id="5A75E4F7FFCE8069C97E37B0744325D8" box="[820,870,1751,1777]" pageId="2" pageNumber="23">type</typeStatus>
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species of the genus, based on hemispermatophore morphology, cheliceral dentition, pedipalps finger terminating in a white patch, and elevated pectinal tooth count. Differs from
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<taxonomicName id="42CE21D6FFCE8069C9B1364073522469" box="[1019,1143,1831,1856]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="reddelli">
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<emphasis id="B7BA8647FFCE8069C9B1364073522469" box="[1019,1143,1831,1856]" italics="true" pageId="2" pageNumber="23">P. reddelli</emphasis>
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</taxonomicName>
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in size, being almost 50% larger; in color, being medium brown without fuscosity (
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<taxonomicName id="42CE21D6FFCE8069CE1E362873EB2441" box="[1108,1230,1871,1896]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="reddelli">
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<emphasis id="B7BA8647FFCE8069CE1E362873EB2441" box="[1108,1230,1871,1896]" italics="true" pageId="2" pageNumber="23">P. reddelli</emphasis>
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</taxonomicName>
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is dark reddish brown and heavily infuscate); in having metasomal segment I clearly longer than wide (subequal in
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<emphasis id="B7BA8647FFCF8068CBA33023757B2274" box="[489,606,324,349]" italics="true" pageId="3" pageNumber="24">P. re d d e l l i</emphasis>
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); in having the pedipalp femur and patella longer than the carapace (shorter or subequal in
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<taxonomicName id="42CE21D6FFCF8068C8CE300B75DE22AC" box="[644,763,364,389]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="reddelli">
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<emphasis id="B7BA8647FFCF8068C8CE300B75DE22AC" box="[644,763,364,389]" italics="true" pageId="3" pageNumber="24">P. reddelli</emphasis>
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</taxonomicName>
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); and in bearing seven rows of denticles clearly defined by six enlarged primary denticles on the movable finger of the pedipalp chela (
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<taxonomicName id="42CE21D6FFCF8068CF5F30F3764722FC" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="reddelli">
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<emphasis id="B7BA8647FFCF8068CF5F30F3764722FC" italics="true" pageId="3" pageNumber="24">P. reddelli</emphasis>
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</taxonomicName>
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has six rows defined by five primary denticles.
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</paragraph>
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</subSubSection>
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<subSubSection id="CDD409DEFFCF806CCB42336C73EE21EF" lastPageId="7" lastPageNumber="28" pageId="3" pageNumber="24" type="description">
|
||
<paragraph id="85715A55FFCF8068CB42336C73F4210F" blockId="3.[264,1324,523,1870]" box="[264,1233,523,550]" pageId="3" pageNumber="24">
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<emphasis id="B7BA8647FFCF8068CB42336C76BC210C" bold="true" box="[264,409,523,549]" pageId="3" pageNumber="24">Description</emphasis>
|
||
. Based on the adult
|
||
<typeStatus id="5A75E4F7FFCF8068C8C3336B75D5210F" box="[649,752,524,550]" pageId="3" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
male and only known specimen (
|
||
<figureCitation id="1DF546D0FFCF8068CE30336B739A210F" box="[1146,1215,524,550]" captionStart="FIGURE 1" captionStartId="4.[264,368,1219,1243]" captionTargetBox="[539,1047,334,1194]" captionTargetId="figure@4.[539,1048,334,1194]" captionTargetPageId="4" captionText="FIGURE 1. Dorsal view of holotype male of Pseudouroctonus sprousei, sp. nov." httpUri="https://zenodo.org/record/173664/files/figure.png" pageId="3" pageNumber="24">Fig. 1</figureCitation>
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||
).
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</paragraph>
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<paragraph id="85715A55FFCF8068CB723353735A2167" blockId="3.[264,1324,523,1870]" box="[312,1151,564,590]" pageId="3" pageNumber="24">Color: Medium brown; venter and legs ochre/yellow, without fuscosity.</paragraph>
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<paragraph id="85715A55FFCF8068CB72333B758D204F" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">
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||
Carapace: Longer than wide. Median eyes on anterior 35%. Ocular tubercle low, without superciliary crests. Median eyes reduced in size,
|
||
<quantity id="4236F7B0FFCF8068C9E433E3732F21B7" box="[942,1034,644,670]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="24" unit="mm" value="0.2">0.2 mm</quantity>
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in diameter. Three pairs of lateral eyes, somewhat reduced in size; posteriormost eye is about ½ diameter of anterior two. Anterior median furrow shallow and broad. Posterior median furrow deep and narrow, the median lobes meeting medially over the furrow. Posterolateral furrows deep and broad. Anterior margin broadly bilobed, with 3 pairs of setae. Entire surface with moderately dense, fine granulation.
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</paragraph>
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<paragraph id="85715A55FFCF8068CB7232137525209F" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">Tergites: With moderately dense, fine granulation. Tergite VII with four denticulate, longitudinal carinae.</paragraph>
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<paragraph id="85715A55FFCF8068CB7232A37438272F" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">Sternum: Subpentagonal, slightly longer than broad; median longitudinal furrow deep and narrow; with seven setae (asymmetrical).</paragraph>
|
||
<paragraph id="85715A55FFCF8068CB7235737575277F" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">Genital operculi: With five setae each; genital papillae well developed, protruding beyond the genital operculi.</paragraph>
|
||
<paragraph id="85715A55FFCF8068CB72350373272757" blockId="3.[264,1324,523,1870]" box="[312,1026,1124,1150]" pageId="3" pageNumber="24">Pectines: With 10–11 middle lamellae; 17 teeth on each side.</paragraph>
|
||
<paragraph id="85715A55FFCF8068CB7235EB7596251F" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">Sternites: Glabrous. Stigmata slitlike, about 3.5 times longer than wide. Sternite VII with one pair of moderately strong, denticulate lateral carinae. Metasoma: Dorsolateral carinae on I–V strong, denticulate/serrate. Lateral supramedian carinae on I–IV strong, denticulate/serrate. Lateral inframedian carinae on I strong, denticulate, complete; on II represented by 3–4 granules distally; on III by one single granule distally; on IV absent. Lateral median carina on V granular on basal onethird. Ventrolateral carinae on I–V, ventral submedian carinae on I–IV and ventral median carina on V strong, denticulate/ serrate. Setation on I–IV: dorsolaterals 0,1,1,2; lateral supramedian 0,1,1,1; lateral inframedian 1,0,0,0; ventrolateral 2,2,2,3; ventral submedian 3,3,3,4. Setation on V: dorsolateral 3, lateromedian 2, ventrolateral 3 and ventromedian 4. Intercarinal spaces densely, minutely granulose (matte).</paragraph>
|
||
<paragraph id="85715A55FFCF8068CB72372373DC25AF" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">
|
||
Telson: Vesicle large and globose (
|
||
<figureCitation id="1DF546D0FFCF8068C89C372374052577" box="[726,800,1604,1630]" captionStart="FIGURES 2 – 8" captionStartId="5.[264,382,1651,1675]" captionTargetBox="[327,1251,486,1629]" captionTargetId="figure@5.[327,1251,486,1629]" captionTargetPageId="5" captionText="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="3" pageNumber="24">Fig. 2</figureCitation>
|
||
), more swollen than that of
|
||
<taxonomicName id="42CE21D6FFCF8068CE3B372373CC2574" box="[1137,1257,1604,1629]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFCF8068CE3B372373CC2574" box="[1137,1257,1604,1629]" italics="true" pageId="3" pageNumber="24">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1DF546D0FFCF8068CEB13723763D25AF" captionStart="FIGURES 2 – 8" captionStartId="5.[264,382,1651,1675]" captionTargetBox="[327,1251,486,1629]" captionTargetId="figure@5.[327,1251,486,1629]" captionTargetPageId="5" captionText="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="3" pageNumber="24">Fig. 3</figureCitation>
|
||
); weakly to faintly granulose, sparsely setose. Aculeus lacking basal microdenticles.
|
||
</paragraph>
|
||
<paragraph id="85715A55FFCF8068CB7237F373362467" blockId="3.[264,1324,523,1870]" pageId="3" pageNumber="24">
|
||
Chelicera: Fixed finger shorter than chela width, movable finger shorter than chela length. Chela with 2 setae dorsally. Fixed finger with basal bicusp symmetrical; ventral margin with 3 prominent accessory denticles (
|
||
<figureCitation id="1DF546D0FFCF8068C975378374A925D7" box="[831,908,1764,1790]" captionStart="FIGURES 2 – 8" captionStartId="5.[264,382,1651,1675]" captionTargetBox="[327,1251,486,1629]" captionTargetId="figure@5.[327,1251,486,1629]" captionTargetPageId="5" captionText="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="3" pageNumber="24">Fig. 4</figureCitation>
|
||
). Movable finger with two dorsal subdistal teeth; with a distinct serrula ventrally, and with ventral carina strongly delineated, bearing three large and several small denticles (
|
||
<figureCitation id="1DF546D0FFCF8068C9F6365373202467" box="[956,1029,1844,1870]" captionStart="FIGURES 2 – 8" captionStartId="5.[264,382,1651,1675]" captionTargetBox="[327,1251,486,1629]" captionTargetId="figure@5.[327,1251,486,1629]" captionTargetPageId="5" captionText="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="3" pageNumber="24">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="D1B10ADDFFC8806FCB4235A4734927F2" box="[264,1132,1219,1243]" httpUri="https://zenodo.org/record/173664/files/figure.png" pageId="4" pageNumber="25" targetBox="[539,1047,334,1194]" targetPageId="4">
|
||
<paragraph id="85715A55FFC8806FCB4235A4734927F2" blockId="4.[264,1132,1219,1243]" box="[264,1132,1219,1243]" pageId="4" pageNumber="25">
|
||
<emphasis id="B7BA8647FFC8806FCB4235A476AF27F2" bold="true" box="[264,394,1219,1243]" pageId="4" pageNumber="25">FIGURE 1.</emphasis>
|
||
Dorsal view of holotype male of
|
||
<emphasis id="B7BA8647FFC8806FC8B835A4733527F3" box="[754,1040,1219,1242]" italics="true" pageId="4" pageNumber="25">
|
||
<taxonomicName id="42CE21D6FFC8806FC8B835A4732927F3" box="[754,1036,1219,1242]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="sprousei" status="sp. nov.">Pseudouroctonus sprousei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="AC893B3CFFC8806FCE5D35A4734927F2" box="[1047,1132,1219,1243]" pageId="4" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="85715A55FFC8806FCB723469751926D9" blockId="4.[264,1324,1294,1880]" pageId="4" pageNumber="25">
|
||
Pedipalp femur: Dorsointernal, dorsoexternal and ventrointernal carinae strong, coarsely granulose; ventroexternal carina obsolete; internomedial carina represented by 4–5 isolated granules which decrease in size distally. Orthobothriotaxia “C” (
|
||
<figureCitation id="1DF546D0FFC8806FCE84343972392651" box="[1230,1308,1374,1400]" captionStart="FIGURES 2 – 8" captionStartId="5.[264,382,1651,1675]" captionTargetBox="[327,1251,486,1629]" captionTargetId="figure@5.[327,1251,486,1629]" captionTargetPageId="5" captionText="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="4" pageNumber="25">Fig. 5</figureCitation>
|
||
). Dorsal face with dense, small granulation; internal face with three submedian setae; external face with four median setae; ventral face with moderately dense granulation, decreasing in size distally.
|
||
</paragraph>
|
||
<paragraph id="85715A55FFC8806FCB72349976CD25C9" blockId="4.[264,1324,1294,1880]" pageId="4" pageNumber="25">
|
||
Pedipalp patella: Internomedian carina represented by a row of 7–8 prominent granules spread on basal twothirds, decreasing in size distally and angling slightly dorsally; dorsointernal carina strong, coarsely granulose; dorsoexternal carina strong, scabrose; ventrointernal carina strong, coarsely granulose; ventroexternal carina strong, scabrose. Orthobothriotaxia “C” (
|
||
<figureCitation id="1DF546D0FFC8806FC8D137F9742A2591" box="[667,783,1694,1720]" captionStart="FIGURES 2 – 8" captionStartId="5.[264,382,1651,1675]" captionTargetBox="[327,1251,486,1629]" captionTargetId="figure@5.[327,1251,486,1629]" captionTargetPageId="5" captionText="FIGURES 2 – 8. Morphology of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 2. Metasomal segment V and telson of P. sprousei, lateral view; 3. Metasomal segment V and telson of P. reddelli, lateral view; 4. Chelicera of P. s p ro u s ei, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of P. s p ro u s e i, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of P. s p ro u s e i, dorsal aspect; 7. pedipalp patella of P. s p ro u s e i, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4 – 8 = 1 mm)." httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="4" pageNumber="25">Figs. 6–8</figureCitation>
|
||
). Intercarinal spaces dorsally matte, matte to glabrous on others.
|
||
</paragraph>
|
||
<paragraph id="85715A55FFC8806ECB7237897406221F" blockId="4.[264,1324,1294,1880]" lastBlockId="5.[264,1323,284,430]" lastPageId="5" lastPageNumber="26" pageId="4" pageNumber="25">
|
||
Pedipalp chela: Carinae scabrous to granulose; intercarinal spaces lustrous. Finger tips with distinctive white patches distally. Fixed finger with 6 rows of granules and 6 inner accessory denticles; movable finger with 7 rows of granules and 7 inner accessory denticles. Orthobothriotaxia “C” (
|
||
<figureCitation id="1DF546D0FFC9806EC8DC307B7430221F" box="[662,789,284,310]" captionStart="FIGURES 9 – 14" captionStartId="6.[264,382,1727,1751]" captionTargetBox="[334,1233,318,1701]" captionTargetId="figure@6.[312,1280,305,1721]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Morphology of Pseudouroctonus sprousei, sp. nov.: 9. Pedipalp chela, dorsal aspect; 10. Pedipalp chela, external aspect; 11. Pedipalp chela, ventrointernal aspect; 12. Telotarsus III, prolateral aspect showing setation pattern; 13. Telotarsus III, ventral aspect, schematic showing ventromedian spinules and setae; 14. Telotarsus III, retrolateral aspect (scale figs. 9 – 11 = 1 mm, figs. 12 – 14 = 0.5 mm)." httpUri="https://zenodo.org/record/173666/files/figure.png" pageId="5" pageNumber="26">Figs. 9–11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="85715A55FFC9806ECB72302373CB2287" blockId="5.[264,1323,284,430]" pageId="5" pageNumber="26">
|
||
Leg III telotarsal armature (
|
||
<figureCitation id="1DF546D0FFC9806EC8D7302374132277" box="[669,822,324,350]" captionStart="FIGURES 9 – 14" captionStartId="6.[264,382,1727,1751]" captionTargetBox="[334,1233,318,1701]" captionTargetId="figure@6.[312,1280,305,1721]" captionTargetPageId="6" captionText="FIGURES 9 – 14. Morphology of Pseudouroctonus sprousei, sp. nov.: 9. Pedipalp chela, dorsal aspect; 10. Pedipalp chela, external aspect; 11. Pedipalp chela, ventrointernal aspect; 12. Telotarsus III, prolateral aspect showing setation pattern; 13. Telotarsus III, ventral aspect, schematic showing ventromedian spinules and setae; 14. Telotarsus III, retrolateral aspect (scale figs. 9 – 11 = 1 mm, figs. 12 – 14 = 0.5 mm)." httpUri="https://zenodo.org/record/173666/files/figure.png" pageId="5" pageNumber="26">Figs. 12–14</figureCitation>
|
||
): two prolateral inframedian setae; two retrolateral inframedian setae; two prolateral infraterminal setae; two retrolateral infraterminal setae. Ventral median spinules: basal cluster, distally with 4–5 spinules.
|
||
</paragraph>
|
||
<caption id="D1B10ADDFFC9806ECB42371476642472" httpUri="https://zenodo.org/record/173665/files/figure.png" pageId="5" pageNumber="26" targetBox="[327,1251,486,1629]" targetPageId="5">
|
||
<paragraph id="85715A55FFC9806ECB42371476642472" blockId="5.[264,1323,1651,1883]" pageId="5" pageNumber="26">
|
||
<emphasis id="B7BA8647FFC9806ECB423714769325A2" bold="true" box="[264,438,1651,1675]" pageId="5" pageNumber="26">FIGURES 2–8.</emphasis>
|
||
Morphology of
|
||
<emphasis id="B7BA8647FFC9806EC822371374A225A2" box="[616,903,1652,1675]" italics="true" pageId="5" pageNumber="26">
|
||
<taxonomicName id="42CE21D6FFC9806EC822371374A625A2" box="[616,899,1652,1675]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="sprousei" status="sp. nov.">Pseudouroctonus sprousei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="AC893B3CFFC9806EC9C5371474C325A2" box="[911,998,1651,1675]" pageId="5" pageNumber="26" rank="species">
|
||
<emphasis id="B7BA8647FFC9806EC9C5371474C325A2" bold="true" box="[911,998,1651,1675]" pageId="5" pageNumber="26">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="42CE21D6FFC9806ECE56371373AC25A2" box="[1052,1161,1652,1675]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFC9806ECE56371373AC25A2" box="[1052,1161,1652,1675]" italics="true" pageId="5" pageNumber="26">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
: 2. Metasomal segment V and telson of
|
||
<taxonomicName id="42CE21D6FFC9806EC86037F075872587" box="[554,674,1687,1710]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFC9806EC86037F075872587" box="[554,674,1687,1710]" italics="true" pageId="5" pageNumber="26">P. sprousei</emphasis>
|
||
</taxonomicName>
|
||
, lateral view; 3. Metasomal segment V and telson of
|
||
<taxonomicName id="42CE21D6FFC9806ECF5D37F0767F25F8" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFC9806ECF5D37F0767F25F8" italics="true" pageId="5" pageNumber="26">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
, lateral view; 4. Chelicera of
|
||
<emphasis id="B7BA8647FFC9806EC8DF37DD742125F8" box="[661,772,1722,1745]" italics="true" pageId="5" pageNumber="26">P. s p ro u s ei</emphasis>
|
||
, ventral view showing diagnostic dentition on both fingers; 5. Pedipalp femur of
|
||
<emphasis id="B7BA8647FFC9806EC80737BB759A25DA" box="[589,703,1756,1779]" italics="true" pageId="5" pageNumber="26">P. s p ro u s e i</emphasis>
|
||
, dorsal view showing trichobothrial pattern; 6. Pedipalp patella of
|
||
<emphasis id="B7BA8647FFC9806ECB30379876CB243F" box="[378,494,1791,1814]" italics="true" pageId="5" pageNumber="26">P. s p ro u s e i</emphasis>
|
||
, dorsal aspect; 7. pedipalp patella of
|
||
<emphasis id="B7BA8647FFC9806EC9D03798732B243F" box="[922,1038,1791,1814]" italics="true" pageId="5" pageNumber="26">P. s p ro u s e i</emphasis>
|
||
, external aspect showing trichobothrial pattern; 8. Pedipalp patella, ventral aspect (Scale figs. 2, 3 = 5 mm, figs. 4–8 = 1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D1B10ADDFFCA806DCB4237D876DB2448" httpUri="https://zenodo.org/record/173666/files/figure.png" pageId="6" pageNumber="27" targetBox="[334,1233,318,1701]" targetPageId="6">
|
||
<paragraph id="85715A55FFCA806DCB4237D876DB2448" blockId="6.[264,1325,1727,1889]" pageId="6" pageNumber="27">
|
||
<emphasis id="B7BA8647FFCA806DCB4237D876E025FE" bold="true" box="[264,453,1727,1751]" pageId="6" pageNumber="27">FIGURES 9–14.</emphasis>
|
||
Morphology of
|
||
<emphasis id="B7BA8647FFCA806DC8CB37A7748625FE" box="[641,931,1728,1751]" italics="true" pageId="6" pageNumber="27">
|
||
<taxonomicName id="42CE21D6FFCA806DC8CB37A774BA25FE" box="[641,927,1728,1751]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="sprousei" status="sp. nov.">Pseudouroctonus sprousei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="AC893B3CFFCA806DC9E437D8732225FE" box="[942,1031,1727,1751]" pageId="6" pageNumber="27" rank="species">
|
||
<emphasis id="B7BA8647FFCA806DC9E437D8732225FE" bold="true" box="[942,1031,1727,1751]" pageId="6" pageNumber="27">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 9. Pedipalp chela, dorsal aspect; 10. Pedipalp chela, external aspect; 11. Pedipalp chela, ventrointernal aspect; 12. Telotarsus III, prolateral aspect showing setation pattern; 13. Telotarsus III, ventral aspect, schematic showing ventromedian spinules and setae; 14. Telotarsus III, retrolateral aspect (scale figs. 9–11 = 1 mm, figs. 12–14 = 0.5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="85715A55FFCB806CCB72307B752A2287" blockId="7.[264,1323,284,430]" pageId="7" pageNumber="28">
|
||
Hemispermatophore: with dorsal hook on basal 28% of lamina (
|
||
<figureCitation id="1DF546D0FFCB806CCE12307B7391221F" box="[1112,1204,284,310]" captionStart="FIGURES 15 – 21" captionStartId="7.[264,382,1623,1647]" captionTargetBox="[302,1288,892,1558]" captionTargetId="figure@7.[264,1323,870,1600]" captionTargetPageId="7" captionText="FIGURES 15 – 21. Hemispermatophores of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 15. Hemispermatophore of P. s p ro u s e i, dorsal aspect; 16, 17. Details of lamellar hook and capsular region of P. sprousei, mating plug removed; 18. Mating plug from the hemispermatophore of P. sprousei; 19. Hemispermatophore of P. reddelli, dorsal aspect; 20, 21. Details of lamellar hook and capsular region of P. reddelli, mating plug in place (scale figs. 15, 19 = 0.67 mm, 16, 17, 20, 21 = 1 mm, 18 = 0, 5 mm)." httpUri="https://zenodo.org/record/173667/files/figure.png" pageId="7" pageNumber="28">Fig. 15</figureCitation>
|
||
). Bears a thickened, acuminate lateral hook connected to the lamella by an elevated lateral flange (
|
||
<figureCitation id="1DF546D0FFCB806CCB5B300B768422AF" box="[273,417,364,390]" captionStart="FIGURES 15 – 21" captionStartId="7.[264,382,1623,1647]" captionTargetBox="[302,1288,892,1558]" captionTargetId="figure@7.[264,1323,870,1600]" captionTargetPageId="7" captionText="FIGURES 15 – 21. Hemispermatophores of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 15. Hemispermatophore of P. s p ro u s e i, dorsal aspect; 16, 17. Details of lamellar hook and capsular region of P. sprousei, mating plug removed; 18. Mating plug from the hemispermatophore of P. sprousei; 19. Hemispermatophore of P. reddelli, dorsal aspect; 20, 21. Details of lamellar hook and capsular region of P. reddelli, mating plug in place (scale figs. 15, 19 = 0.67 mm, 16, 17, 20, 21 = 1 mm, 18 = 0, 5 mm)." httpUri="https://zenodo.org/record/173667/files/figure.png" pageId="7" pageNumber="28">Figs. 16–17</figureCitation>
|
||
); mating plug with distal barb margin smooth (
|
||
<figureCitation id="1DF546D0FFCB806CC987300B730122AF" box="[973,1060,364,390]" captionStart="FIGURES 15 – 21" captionStartId="7.[264,382,1623,1647]" captionTargetBox="[302,1288,892,1558]" captionTargetId="figure@7.[264,1323,870,1600]" captionTargetPageId="7" captionText="FIGURES 15 – 21. Hemispermatophores of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 15. Hemispermatophore of P. s p ro u s e i, dorsal aspect; 16, 17. Details of lamellar hook and capsular region of P. sprousei, mating plug removed; 18. Mating plug from the hemispermatophore of P. sprousei; 19. Hemispermatophore of P. reddelli, dorsal aspect; 20, 21. Details of lamellar hook and capsular region of P. reddelli, mating plug in place (scale figs. 15, 19 = 0.67 mm, 16, 17, 20, 21 = 1 mm, 18 = 0, 5 mm)." httpUri="https://zenodo.org/record/173667/files/figure.png" pageId="7" pageNumber="28">Fig. 18</figureCitation>
|
||
). Resembles that of
|
||
<taxonomicName id="42CE21D6FFCB806CCF5F300B76472284" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFCB806CCF5F300B76472284" italics="true" pageId="7" pageNumber="28">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1DF546D0FFCB806CCB3830F375272287" box="[370,514,404,430]" captionStart="FIGURES 15 – 21" captionStartId="7.[264,382,1623,1647]" captionTargetBox="[302,1288,892,1558]" captionTargetId="figure@7.[264,1323,870,1600]" captionTargetPageId="7" captionText="FIGURES 15 – 21. Hemispermatophores of Pseudouroctonus sprousei, sp. nov. and P. reddelli: 15. Hemispermatophore of P. s p ro u s e i, dorsal aspect; 16, 17. Details of lamellar hook and capsular region of P. sprousei, mating plug removed; 18. Mating plug from the hemispermatophore of P. sprousei; 19. Hemispermatophore of P. reddelli, dorsal aspect; 20, 21. Details of lamellar hook and capsular region of P. reddelli, mating plug in place (scale figs. 15, 19 = 0.67 mm, 16, 17, 20, 21 = 1 mm, 18 = 0, 5 mm)." httpUri="https://zenodo.org/record/173667/files/figure.png" pageId="7" pageNumber="28">Figs. 19–21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="85715A55FFCB806CCB42308473EE21EF" blockId="7.[264,1323,483,710]" pageId="7" pageNumber="28">
|
||
<emphasis id="B7BA8647FFCB806CCB423084751222D4" bold="true" box="[264,567,483,509]" pageId="7" pageNumber="28">Measurements (in mm)</emphasis>
|
||
. Total L 74.1; carapace L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CC9C1308374FC22D7" box="[907,985,484,510]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-8.6">W 8.6</geoCoordinate>
|
||
/7.0; distance from anterior margin to median eyes 3.2; mesosoma L 18.4; metasoma total L 47.1; segment I L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CCE90336B7204210F" box="[1242,1313,524,550]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-5.0">W 5.0</geoCoordinate>
|
||
/ 3.6; II L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CCB3B3353769C2167" box="[369,441,564,590]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-6.0">W 6.0</geoCoordinate>
|
||
/3.4, III L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CC8783353755F2167" box="[562,634,564,590]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-6.3">W 6.3</geoCoordinate>
|
||
/3.2; IV L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CC8BC3353741B2167" box="[758,830,564,590]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-7.8">W 7.8</geoCoordinate>
|
||
/3.0; V L/W/D 11.8/2.8/2.5; telson L/W/D 10.2/3.2/3.3; chelicera L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CC87D333B75A0215F" box="[567,645,604,630]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-3.4">W 3.4</geoCoordinate>
|
||
/1.8, fixed finger L 1.0, movable finger L 2.0; pedipalp femur L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CCB3F33E376E121B7" box="[373,452,644,670]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-9.2">W 9.2</geoCoordinate>
|
||
/2.4; pedipalp patella L/
|
||
<geoCoordinate id="E0FA3C92FFCB806CC8A133E3741C21B7" box="[747,825,644,670]" direction="west" orientation="longitude" pageId="7" pageNumber="28" precision="5555" value="-9.2">W 9.2</geoCoordinate>
|
||
/2.6; pedipalp chela L/W/D 15.6/3.5/4.3, fixed finger L 6.2, movable finger L 7.6. Hemispermatophore L 8.0, lamina L 5.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CDD409DEFFCB806CCB42339B73DD2017" pageId="7" pageNumber="28" type="etymology">
|
||
<paragraph id="85715A55FFCB806CCB42339B73DD2017" blockId="7.[264,1324,764,830]" pageId="7" pageNumber="28">
|
||
<emphasis id="B7BA8647FFCB806CCB42339B76B0203F" bold="true" box="[264,405,764,790]" pageId="7" pageNumber="28">Etymology.</emphasis>
|
||
This species is dedicated to Mr. Peter Sprouse, a tireless explorer of Mexican caves and collector of numerous troglobites, including the
|
||
<typeStatus id="5A75E4F7FFCB806CC9F23243733A2017" box="[952,1055,804,830]" pageId="7" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
of this new taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D1B10ADDFFCB806CCB42373076F22435" httpUri="https://zenodo.org/record/173667/files/figure.png" pageId="7" pageNumber="28" targetBox="[302,1288,892,1558]" targetPageId="7">
|
||
<paragraph id="85715A55FFCB806CCB42373076F22435" blockId="7.[264,1323,1623,1820]" pageId="7" pageNumber="28">
|
||
<emphasis id="B7BA8647FFCB806CCB42373076EA2546" bold="true" box="[264,463,1623,1647]" pageId="7" pageNumber="28">FIGURES 15–21.</emphasis>
|
||
Hemispermatophores of
|
||
<emphasis id="B7BA8647FFCB806CC894373F74D82546" box="[734,1021,1624,1647]" italics="true" pageId="7" pageNumber="28">
|
||
<taxonomicName id="42CE21D6FFCB806CC894373F74DC2546" box="[734,1017,1624,1647]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="sprousei" status="sp. nov.">Pseudouroctonus sprousei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="AC893B3CFFCB806CCE4E3730737F2546" box="[1028,1114,1623,1647]" pageId="7" pageNumber="28" rank="species">
|
||
<emphasis id="B7BA8647FFCB806CCE4E3730737F2546" bold="true" box="[1028,1114,1623,1647]" pageId="7" pageNumber="28">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="42CE21D6FFCB806CCEC4373F73DE2546" box="[1166,1275,1624,1647]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFCB806CCEC4373F73DE2546" box="[1166,1275,1624,1647]" italics="true" pageId="7" pageNumber="28">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
: 15. Hemispermatophore of
|
||
<emphasis id="B7BA8647FFCB806CC846371D75A525B8" box="[524,640,1658,1681]" italics="true" pageId="7" pageNumber="28">P. s p ro u s e i</emphasis>
|
||
, dorsal aspect; 16, 17. Details of lamellar hook and capsular region of
|
||
<taxonomicName id="42CE21D6FFCB806CCB3C37FA76C9259D" box="[374,492,1693,1716]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFCB806CCB3C37FA76C9259D" box="[374,492,1693,1716]" italics="true" pageId="7" pageNumber="28">P. sprousei</emphasis>
|
||
</taxonomicName>
|
||
, mating plug removed; 18. Mating plug from the hemispermatophore of
|
||
<taxonomicName id="42CE21D6FFCB806CCF5D37FA764425FE" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFCB806CCF5D37FA764425FE" italics="true" pageId="7" pageNumber="28">P. sprousei</emphasis>
|
||
</taxonomicName>
|
||
; 19. Hemispermatophore of
|
||
<taxonomicName id="42CE21D6FFCB806CC8DC37A7742725FE" box="[662,770,1728,1751]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFCB806CC8DC37A7742725FE" box="[662,770,1728,1751]" italics="true" pageId="7" pageNumber="28">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
, dorsal aspect; 20, 21. Details of lamellar hook and capsular region of
|
||
<taxonomicName id="42CE21D6FFCB806CCB853785751E25D0" box="[463,571,1762,1785]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFCB806CCB853785751E25D0" box="[463,571,1762,1785]" italics="true" pageId="7" pageNumber="28">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
, mating plug in place (scale figs. 15, 19 = 0.67 mm, 16, 17, 20, 21 = 1 mm, 18 = 0,5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="CDD409DEFFC48063CB42307C7534278F" pageId="8" pageNumber="29" type="discussion">
|
||
<paragraph id="85715A55FFC48063CB42307C7534278F" blockId="8.[264,1323,283,1190]" pageId="8" pageNumber="29">
|
||
<emphasis id="B7BA8647FFC48063CB42307C76AA221C" bold="true" box="[264,399,283,309]" pageId="8" pageNumber="29">Comments</emphasis>
|
||
.
|
||
<taxonomicName id="42CE21D6FFC48063CBEF307B75C4221C" box="[421,737,284,309]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFC48063CBEF307B75C4221C" box="[421,737,284,309]" italics="true" pageId="8" pageNumber="29">Pseudouroctonus sprousei</emphasis>
|
||
</taxonomicName>
|
||
has median and lateral eyes (these somewhat reduced in size), so it has not reached the extreme condition of eyelessness observed in other Mexican cave scorpions (
|
||
<emphasis id="B7BA8647FFC48063C8E9300B75C222AC" box="[675,743,364,389]" italics="true" pageId="8" pageNumber="29">e. g.,</emphasis>
|
||
some
|
||
<taxonomicName id="42CE21D6FFC48063C902300B734C22AF" authority="Mitchell" authorityName="Mitchell" box="[840,1129,364,390]" class="Arachnida" family="Superstitioniidae" genus="Typhlochactas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B7BA8647FFC48063C902300B74D422AC" box="[840,1009,364,389]" italics="true" pageId="8" pageNumber="29">Typhlochactas</emphasis>
|
||
Mitchell
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="42CE21D6FFC48063CECA300B764C2287" authority="Francke" authorityName="Francke" class="Arachnida" family="Superstitioniidae" genus="Sotanochactas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B7BA8647FFC48063CECA300B720E22AC" box="[1152,1323,364,389]" italics="true" pageId="8" pageNumber="29">Sotanochactas</emphasis>
|
||
Francke
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="42CE21D6FFC48063CB3130F375A92287" authority="Francke" authorityName="Francke" box="[379,652,404,430]" class="Arachnida" family="Euscorpiidae" genus="Troglocormus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B7BA8647FFC48063CB3130F3753B2284" box="[379,542,404,429]" italics="true" pageId="8" pageNumber="29">Troglocormus</emphasis>
|
||
Francke
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="42CE21D6FFC48063C89E30F374862287" authority="Francke" authorityName="Francke" box="[724,931,404,430]" class="Arachnida" family="Superstitioniidae" genus="Alacran" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B7BA8647FFC48063C89E30F374162284" box="[724,819,404,429]" italics="true" pageId="8" pageNumber="29">Alacran</emphasis>
|
||
Francke
|
||
</taxonomicName>
|
||
). Although
|
||
<taxonomicName id="42CE21D6FFC48063CE7C30F373962284" box="[1078,1203,404,429]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFC48063CE7C30F373962284" box="[1078,1203,404,429]" italics="true" pageId="8" pageNumber="29">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
has been collected in numerous caves (as well as many surface locations) in Texas and Coahuila, it has no apparent morphological modifications for life in caves. It is hypothesized that
|
||
<taxonomicName id="42CE21D6FFC48063CF5F3083764F210C" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFC48063CF5F3083764F210C" italics="true" pageId="8" pageNumber="29">P. sprousei</emphasis>
|
||
</taxonomicName>
|
||
is troglobitic when compared to
|
||
<emphasis id="B7BA8647FFC48063C947336B74AB210C" box="[781,910,524,549]" italics="true" pageId="8" pageNumber="29">P. re d d e l l i,</emphasis>
|
||
its putative closest relative: it is considerably paler in color; the metasoma is elongated [apparent when the length/width ratio of segment I (close to
|
||
<quantity id="4236F7B0FFC48063C832333B75E7215F" box="[632,706,604,630]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="8" pageNumber="29" unit="in" value="1.5">1.5 in</quantity>
|
||
<taxonomicName id="42CE21D6FFC48063C89B333B747D215C" box="[721,856,604,629]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFC48063C89B333B747D215C" box="[721,856,604,629]" italics="true" pageId="8" pageNumber="29">P. sprousei</emphasis>
|
||
</taxonomicName>
|
||
) is compared to that of
|
||
<taxonomicName id="42CE21D6FFC48063CEE6333B720F215C" box="[1196,1322,604,629]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFC48063CEE6333B720F215C" box="[1196,1322,604,629]" italics="true" pageId="8" pageNumber="29">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
(approximately equal to 1)]; and the pedipalps are attenuated [apparent when the relative lengths of the femur and patella are compared with the carapace length in the two sister species—subequal in
|
||
<taxonomicName id="42CE21D6FFC48063C84333B375A421C4" box="[521,641,724,749]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="reddelli">
|
||
<emphasis id="B7BA8647FFC48063C84333B375A421C4" box="[521,641,724,749]" italics="true" pageId="8" pageNumber="29">P. reddelli</emphasis>
|
||
</taxonomicName>
|
||
and longer in
|
||
<taxonomicName id="42CE21D6FFC48063C96733B3748821C4" box="[813,941,724,749]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="sprousei">
|
||
<emphasis id="B7BA8647FFC48063C96733B3748821C4" box="[813,941,724,749]" italics="true" pageId="8" pageNumber="29">P. sprousei</emphasis>
|
||
</taxonomicName>
|
||
]. This hypothesis awaits further testing by the collection of additional specimens and data on their habitat and biology. Among the
|
||
<taxonomicName id="42CE21D6FFC48063CBE5324375112017" box="[431,564,804,830]" class="Arachnida" family="Vaejovidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="family">Vaejovidae</taxonomicName>
|
||
, three troglophiles (without obvious cave adaptations or troglomorphies) are known:
|
||
<emphasis id="B7BA8647FFC48063C817322B748B204C" box="[605,942,844,869]" italics="true" pageId="8" pageNumber="29">
|
||
<taxonomicName id="42CE21D6FFC48063C817322B75FF204C" box="[605,730,844,869]" class="Arachnida" family="Vaejovidae" genus="Pseudouroctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="reddelli">P. reddelli</taxonomicName>
|
||
,
|
||
<taxonomicName id="42CE21D6FFC48063C8A2322B748B204C" authority="Sissom and Gonzalez-Santillan" authorityName="Sissom and Gonzalez-Santillan" box="[744,942,844,869]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="norteno">Vaejovis norteno</taxonomicName>
|
||
</emphasis>
|
||
Sissom and GonzalezSantillán from Coahuila and Nuevo León,
|
||
<collectingCountry id="FDD91AC5FFC48063C8E33213742320A7" box="[681,774,884,910]" name="Mexico" pageId="8" pageNumber="29">México</collectingCountry>
|
||
, and
|
||
<taxonomicName id="42CE21D6FFC48063C904321373C120A7" authority="Soleglad and Fet" authorityName="Soleglad and Fet" box="[846,1252,884,910]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="davidi">
|
||
<emphasis id="B7BA8647FFC48063C9043213732320A4" box="[846,1030,884,909]" italics="true" pageId="8" pageNumber="29">Vaejovis davidi</emphasis>
|
||
Soleglad and Fet
|
||
</taxonomicName>
|
||
from Puebla,
|
||
<collectingCountry id="FDD91AC5FFC48063CB2032FB76ED209F" box="[362,456,924,950]" name="Mexico" pageId="8" pageNumber="29">México</collectingCountry>
|
||
. An additional troglophile,
|
||
<taxonomicName id="42CE21D6FFC48063C95332FB720F209F" authority="Gertsch and Soleglad" authorityName="Gertsch and Soleglad" box="[793,1322,924,950]" class="Arachnida" family="Vaejovidae" genus="Uroctonites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="sequoia">
|
||
<emphasis id="B7BA8647FFC48063C95332FB732C209C" box="[793,1033,924,949]" italics="true" pageId="8" pageNumber="29">Uroctonites sequoia</emphasis>
|
||
(Gertsch and Soleglad)
|
||
</taxonomicName>
|
||
from California, has the number of lateral eyes reduced in most specimens, but is known from at least one epigean locality. Two other vaejovids,
|
||
<taxonomicName id="42CE21D6FFC48063C9EE328B765D2707" authority="Gertsch and Soleglad" authorityName="Gertsch and Soleglad" class="Arachnida" family="Vaejovidae" genus="Uroctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="B7BA8647FFC48063C9EE328B73AE272C" box="[932,1163,1004,1029]" italics="true" pageId="8" pageNumber="29">Uroctonus grahami</emphasis>
|
||
(Gertsch and Soleglad)
|
||
</taxonomicName>
|
||
from California, and
|
||
<taxonomicName id="42CE21D6FFC48063C8C73573734C2707" authority="Gertsch and Soleglad" authorityName="Gertsch and Soleglad" box="[653,1129,1044,1070]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B7BA8647FFC48063C8C7357374732704" box="[653,854,1044,1069]" italics="true" pageId="8" pageNumber="29">Vaejovis gracilis</emphasis>
|
||
Gertsch and Soleglad
|
||
</taxonomicName>
|
||
from Veracruz, appear to be true troglobites (they are known only from inside of caves) and exhibit distinct cave adaptations (
|
||
<emphasis id="B7BA8647FFC48063C80B350375582754" box="[577,637,1124,1149]" italics="true" pageId="8" pageNumber="29">e. g.,</emphasis>
|
||
reduced eye size and reduced pigmentation compared to their closest relatives).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |