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<mods:title id="35002EB3CC9E526C0575B16A844A0544">Five new species of the Afrotropical dark sac spider genus Messapus Simon, 1898 (Araneae: Corinnidae)</mods:title>
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Genus
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<emphasis id="BEB7EA94FFB2FF92FF337B35551FFE4A" bold="true" box="[237,356,359,385]" italics="true" pageId="2" pageNumber="387">Messapus</emphasis>
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<emphasis id="BEB7EA94FFB2FF92FF497BE25480FE0E" box="[151,251,432,453]" italics="true" pageId="2" pageNumber="387">Messapus</emphasis>
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<bibRefCitation id="E8524B77FFB2FF92FEDF7BFC55FDFE0F" author="Simon" box="[257,390,430,452]" pageId="2" pageNumber="387" refString="Simon, E. (1898) Histoire naturelle des araignees. Tom 2. Librairie Encyclopedique de Roret, Paris, pp. 193 - 380." type="journal article" year="1898">Simon, 1898</bibRefCitation>
: 214
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;
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<bibRefCitation id="E8524B77FFB2FF92FE1D7BFC561BFE0F" author="Reiskind" box="[451,608,430,452]" pageId="2" pageNumber="387" refString="Reiskind, J. (1969) The spider subfamily Castianeirinae of North and Central America (Araneae, Clubionidae). Bulletin of the Museum of Comparative Zoology, 138, 163 - 325." type="journal article" year="1969">Reiskind, 1969</bibRefCitation>
: 166
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;
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<bibRefCitation id="E8524B77FFB2FF92FD437BFC57C8FE0F" author="Bosselaers" box="[669,947,430,452]" pageId="2" pageNumber="387" refString="Bosselaers, J. &amp; Jocque, R. (2000) Studies in Corinnidae: transfer of four genera and descriptions of the female of Lessertina mutica Lawrence 1942. Tropical Zoology, 13, 305 - 325. http: // dx. doi. org / 10.1080 / 03946975.2000.10531138" type="journal article" year="2000">Bosselaers &amp; Jocqué, 2000</bibRefCitation>
: 307
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;
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<bibRefCitation id="E8524B77FFB2FF92FC2E7BFC50FAFE0F" author="Haddad" box="[1008,1153,430,452]" pageId="2" pageNumber="387" refString="Haddad, C. R. (2013) Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae. Zootaxa, 3688 (1), 1 - 79. http: // dx. doi. org / 10.11646 / zootaxa. 3688.1.1" type="journal article" year="2013">Haddad, 2013</bibRefCitation>
: 16
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.
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species:
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<taxonomicName id="4BC34D05FFB2FF92FEF57B9C5611FE28" ID-CoL="427TN" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[299,618,461,483]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB2FF92FEF57B9C559BFE28" box="[299,480,462,483]" italics="true" pageId="2" pageNumber="387">Messapus martini</emphasis>
<bibRefCitation id="E8524B77FFB2FF92FE387B9F5611FE28" author="Simon" box="[486,618,461,483]" pageId="2" pageNumber="387" refString="Simon, E. (1898) Histoire naturelle des araignees. Tom 2. Librairie Encyclopedique de Roret, Paris, pp. 193 - 380." type="journal article" year="1898">Simon, 1898</bibRefCitation>
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, by original designation.
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<paragraph id="8C7C3686FFB2FF92FF49785D5138FD73" blockId="2.[151,1436,527,696]" pageId="2" pageNumber="387">
<emphasis id="BEB7EA94FFB2FF92FF49785D56BAFDE3" bold="true" box="[151,705,527,552]" pageId="2" pageNumber="387">Remarks and supplementary descriptive data:</emphasis>
<bibRefCitation id="E8524B77FFB2FF92FD17785D5709FDE3" author="Haddad" box="[713,882,527,552]" pageId="2" pageNumber="387" refString="Haddad, C. R. (2013) Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae. Zootaxa, 3688 (1), 1 - 79. http: // dx. doi. org / 10.11646 / zootaxa. 3688.1.1" type="journal article" year="2013">Haddad (2013)</bibRefCitation>
provided a detailed description and diagnosis for the genus. The presence of a lobe found on the distal side of the cheliceral paturon in front of the fang base (cheliceral promarginal pronounced mound, CPPM,
<figureCitation id="14F82A03FFB2FF92FD3D7805575DFDBB" box="[739,806,599,624]" captionStart="FIGURES 3 10" captionStartId="2.[151,264,1661,1683]" captionTargetBox="[151,1436,749,1639]" captionTargetId="figure@2.[151,1436,749,1640]" captionTargetPageId="2" captionText="FIGURES 3 10. Digital microscope photographs of Messapus species, female right chelicerae in posterior view (3) and dorsal habitus (4 10): 3, 8 M. tigris sp. n., female; 4 M. megae sp. n., female; 5 Same, male; 6 M. meridionalis sp. n., female; 7 M. seiugatus sp. n., female; 9 M. tropicus sp. n., female; 10 Same, male. Scale bars: 3 = 1 mm, 4 10 = 2 mm. Abbreviations: CPPM—cheliceral promarginal pronounced mound; F—fang; P—paturon; PMT—promarginal teeth; RMT—retromarginal teeth." httpUri="https://zenodo.org/record/238362/files/figure.png" pageId="2" pageNumber="387">Fig. 3</figureCitation>
; also see
<bibRefCitation id="E8524B77FFB2FF92FC4C780A504FFDBB" author="Ramirez" box="[914,1076,600,624]" pageId="2" pageNumber="387" refString="Ramirez, M. J. (2014) The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374." type="journal article" year="2014">Ramírez 2014</bibRefCitation>
: 53) is a structure reported from the genus for the first time here. A CPPM is found in four of the
<taxonomicName id="4BC34D05FFB2FF92FC5C782F578BFD5F" box="[898,1008,637,660]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="BEB7EA94FFB2FF92FC5C782F578BFD5F" box="[898,1008,637,660]" italics="true" pageId="2" pageNumber="387">Messapus</emphasis>
</taxonomicName>
species described here, as well as
<taxonomicName id="4BC34D05FFB2FF92FAA3782F5498FD73" class="Arachnida" family="Corinnidae" genus="Corinna" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="natalis">
<emphasis id="BEB7EA94FFB2FF92FAA3782F5498FD73" italics="true" pageId="2" pageNumber="387">M. natalis</emphasis>
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, but is absent from
<taxonomicName id="4BC34D05FFB2FF92FE1F78F35642FD7C" box="[449,569,672,696]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB2FF92FE1F78F35642FD7C" box="[449,569,672,696]" italics="true" pageId="2" pageNumber="387">M. martini</emphasis>
</taxonomicName>
and
<taxonomicName id="4BC34D05FFB2FF92FDAE78F35758FD73" box="[624,803,672,696]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FDAE78F35758FD73" box="[624,803,672,696]" italics="true" pageId="2" pageNumber="387">M. meridionalis</emphasis>
</taxonomicName>
<emphasis id="BEB7EA94FFB2FF92FCF478CD5712FD73" bold="true" box="[810,873,671,696]" pageId="2" pageNumber="387">
<taxonomicNameLabel id="A58457EFFFB2FF92FCF478CD5712FD73" box="[810,873,671,696]" pageId="2" pageNumber="387" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and may be of phylogenetic significance.
</paragraph>
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<paragraph id="8C7C3686FFB2FF92FF497C2F54B4F8C5" blockId="2.[151,1437,1661,1806]" pageId="2" pageNumber="387">
<emphasis id="BEB7EA94FFB2FF92FF497C2F553DF958" bold="true" box="[151,326,1661,1683]" pageId="2" pageNumber="387">FIGURES 310.</emphasis>
Digital microscope photographs of
<taxonomicName id="4BC34D05FFB2FF92FD6C7C2C576DF958" box="[690,790,1662,1683]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="BEB7EA94FFB2FF92FD6C7C2C576DF958" box="[690,790,1662,1683]" italics="true" pageId="2" pageNumber="387">Messapus</emphasis>
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species, female right chelicerae in posterior view (3) and dorsal habitus (410): 3, 8
<taxonomicName id="4BC34D05FFB2FF92FEB97CCF55C4F979" box="[359,447,1693,1714]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FEB97CCF55C4F979" box="[359,447,1693,1714]" italics="true" pageId="2" pageNumber="387">M. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A58457EFFFB2FF92FE187CCE567BF979" box="[454,512,1692,1714]" pageId="2" pageNumber="387" rank="species">
<emphasis id="BEB7EA94FFB2FF92FE187CCE567BF979" bold="true" box="[454,512,1692,1714]" pageId="2" pageNumber="387">sp. n.</emphasis>
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, female; 4
<taxonomicName id="4BC34D05FFB2FF92FDAB7CCF56A2F979" box="[629,729,1693,1714]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="megae" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FDAB7CCF56A2F979" box="[629,729,1693,1714]" italics="true" pageId="2" pageNumber="387">M. megae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A58457EFFFB2FF92FD3F7CCE5760F979" box="[737,795,1692,1714]" pageId="2" pageNumber="387" rank="species">
<emphasis id="BEB7EA94FFB2FF92FD3F7CCE5760F979" bold="true" box="[737,795,1692,1714]" pageId="2" pageNumber="387">sp. n.</emphasis>
</taxonomicNameLabel>
, female; 5 Same, male; 6
<taxonomicName id="4BC34D05FFB2FF92FBF67CCF50B0F979" box="[1064,1227,1692,1714]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FBF67CCF50B0F979" box="[1064,1227,1692,1714]" italics="true" pageId="2" pageNumber="387">M. meridionalis</emphasis>
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<emphasis id="BEB7EA94FFB2FF92FB0C7CCE5177F979" bold="true" box="[1234,1292,1692,1714]" pageId="2" pageNumber="387">sp. n.</emphasis>
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, female; 7
<taxonomicName id="4BC34D05FFB2FF92FA5E7CCF548CF91A" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="seiugatus" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FA5E7CCF548CF91A" italics="true" pageId="2" pageNumber="387">M. seiugatus</emphasis>
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<emphasis id="BEB7EA94FFB2FF92FF217CE95541F91A" bold="true" box="[255,314,1723,1745]" pageId="2" pageNumber="387">sp. n.</emphasis>
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, female; 9
<taxonomicName id="4BC34D05FFB2FF92FE6C7CEE5653F91A" box="[434,552,1724,1745]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="tropicus" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FE6C7CEE5653F91A" box="[434,552,1724,1745]" italics="true" pageId="2" pageNumber="387">M. tropicus</emphasis>
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<emphasis id="BEB7EA94FFB2FF92FDEF7CE95617F91A" bold="true" box="[561,620,1723,1745]" pageId="2" pageNumber="387">sp. n.</emphasis>
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, female; 10 Same, male. Scale bars: 3 = 1 mm, 410 = 2 mm. Abbreviations: CPPM—cheliceral promarginal pronounced mound; F—fang; P—paturon; PMT—promarginal teeth; RMT—retromarginal teeth.
</paragraph>
</caption>
<subSubSection id="C4D9650DFFB2FF94FF197D325530F8D0" lastPageId="4" lastPageNumber="389" pageId="2" pageNumber="387" type="reference_group">
<paragraph id="8C7C3686FFB2FF93FF197D325621FE67" blockId="2.[151,1437,1886,2020]" lastBlockId="3.[151,1437,151,429]" lastPageId="3" lastPageNumber="388" pageId="2" pageNumber="387">
<taxonomicName id="4BC34D05FFB2FF92FF197D3255F5F8BC" ID-CoL="427TN" box="[199,398,1888,1911]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB2FF92FF197D3255F5F8BC" box="[199,398,1888,1911]" italics="true" pageId="2" pageNumber="387">Messapus martini</emphasis>
</taxonomicName>
possesses feathery setae (FS) on the body and legs (
<figureCitation id="14F82A03FFB2FF92FC0A7D0C5028F8BC" box="[980,1107,1886,1911]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="2" pageNumber="387">Figs 1118</figureCitation>
) typical for
<taxonomicName id="4BC34D05FFB2FF92FB027D0C5127F8BC" ID-CoL="8KG" box="[1244,1372,1886,1911]" class="Arachnida" family="Corinnidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="family">Corinnidae</taxonomicName>
. Here we report on the presence of squamate setae with brachia (SSB;
<emphasis id="BEB7EA94FFB2FF92FCAA7DD157C9F851" box="[884,946,1923,1946]" italics="true" pageId="2" pageNumber="387">sensu</emphasis>
<bibRefCitation id="E8524B77FFB2FF92FC647DD05022F850" author="Murphy" box="[954,1113,1922,1947]" pageId="2" pageNumber="387" refString="Murphy, J. (2007) Gnaphosid genera of the world. Vol. 1 &amp; 2. British Arachnological Society, St Neots, Cambridge, 605 pp. [pp. i - xii + 1 - 92 &amp; pp. i - ii + 93 - 605]" type="book chapter" year="2007">Murphy 2007</bibRefCitation>
: 30, fig. k; see also
<bibRefCitation id="E8524B77FFB2FF92FAE37DD054AFF874" author="Bonaldo" pageId="2" pageNumber="387" refString="Bonaldo, A. B. (2000) Taxonomia da subfamilia Corinninae (Araneae, Corinnidae) nas Regioes Neotropical e Neartica. Iheringia, Serie Zoologia, 89, 3 - 148. http: // dx. doi. org / 10.1590 / s 0073 - 47212007000300015" type="journal article" year="2000">Bonaldo 2000</bibRefCitation>
: figs 44, 46, 48 and
<bibRefCitation id="E8524B77FFB2FF92FE687DF55622F874" author="Ramirez" box="[438,601,1959,1983]" pageId="2" pageNumber="387" refString="Ramirez, M. J. (2014) The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374." type="journal article" year="2014">Ramírez 2014</bibRefCitation>
: figs 92H, I, 93D, G for possible homologs) for the first time in the genus. These setae cover most of the body and legs in
<taxonomicName id="4BC34D05FFB2FF92FD6D7D9E576FF828" ID-CoL="427TT" box="[691,788,1996,2019]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="BEB7EA94FFB2FF92FD6D7D9E576FF828" box="[691,788,1996,2019]" italics="true" pageId="2" pageNumber="387">M. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A58457EFFFB2FF92FCC27D995726F82F" box="[796,861,1995,2020]" pageId="2" pageNumber="387" rank="species">
<emphasis id="BEB7EA94FFB2FF92FCC27D995726F82F" bold="true" box="[796,861,1995,2020]" pageId="2" pageNumber="387">sp. n.</emphasis>
</taxonomicNameLabel>
but are not reported in the other
<taxonomicName id="4BC34D05FFB2FF92FB0A7D9E5139F828" ID-CoL="5Q3W" box="[1236,1346,1996,2019]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="BEB7EA94FFB2FF92FB0A7D9E5139F828" box="[1236,1346,1996,2019]" italics="true" pageId="2" pageNumber="387">Messapus</emphasis>
</taxonomicName>
species treated here. In
<taxonomicName id="4BC34D05FFB3FF93FE977ACB55D0FF64" ID-CoL="427TT" box="[329,427,152,176]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="BEB7EA94FFB3FF93FE977ACB55D0FF64" box="[329,427,152,176]" italics="true" pageId="3" pageNumber="388">M. tigris</emphasis>
</taxonomicName>
<emphasis id="BEB7EA94FFB3FF93FE6D7AC55588FF7B" bold="true" box="[435,499,151,176]" pageId="3" pageNumber="388">
<taxonomicNameLabel id="A58457EFFFB3FF93FE6D7AC55588FF7B" box="[435,499,151,176]" pageId="3" pageNumber="388" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, SSB are found from the coxae to the proximal region of the tarsal segments (
<figureCitation id="14F82A03FFB3FF93FAB47AC5549BFF1F" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="3" pageNumber="388">Figs 2328</figureCitation>
), with the brachiae of the SSB increasing in length along the length of the tarsi so that the distal setae have a relatively narrow shaft and long brachiae (
<figureCitation id="14F82A03FFB3FF93FDAC7A8D56BEFF33" box="[626,709,223,248]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="3" pageNumber="388">Fig. 27</figureCitation>
), similar to the FS typical of other species in the genus, although with a slightly thicker shaft. While the SSB are the dominant setal
<typeStatus id="53788824FFB3FF93FCA47B5757D6FED6" box="[890,941,261,285]" pageId="3" pageNumber="388">type</typeStatus>
, there are also scattered elongate scales with fine longitudinal striae present (e.g.
<figureCitation id="14F82A03FFB3FF93FDE17B7556BCFE8B" box="[575,711,295,320]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="3" pageNumber="388">Figs 24, 26</figureCitation>
). The FS of
<taxonomicName id="4BC34D05FFB3FF93FCBF7B7B506CFE8B" ID-CoL="427TQ" box="[865,1047,296,320]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="BEB7EA94FFB3FF93FCBF7B7B506CFE8B" box="[865,1047,296,320]" italics="true" pageId="3" pageNumber="388">M. meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A58457EFFFB3FF93FBFC7B75501EFE8B" box="[1058,1125,295,320]" pageId="3" pageNumber="388" rank="species">
<emphasis id="BEB7EA94FFB3FF93FBFC7B75501EFE8B" bold="true" box="[1058,1125,295,320]" pageId="3" pageNumber="388">sp. n.</emphasis>
</taxonomicNameLabel>
are similar to those of
<taxonomicName id="4BC34D05FFB3FF93FAA37B7B5491FEAF" ID-CoL="427TN" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB3FF93FAA37B7B5491FEAF" italics="true" pageId="3" pageNumber="388">M. martini</emphasis>
</taxonomicName>
, but those of the other
<taxonomicName id="4BC34D05FFB3FF93FE327B1F5620FEAF" ID-CoL="5Q3W" box="[492,603,333,356]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="genus">
<emphasis id="BEB7EA94FFB3FF93FE327B1F5620FEAF" box="[492,603,333,356]" italics="true" pageId="3" pageNumber="388">Messapus</emphasis>
</taxonomicName>
species are somewhat intermediate between the extremes presented by
<taxonomicName id="4BC34D05FFB3FF93FAA37B1F5491FE4C" ID-CoL="427TN" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB3FF93FAA37B1F5491FE4C" italics="true" pageId="3" pageNumber="388">M. martini</emphasis>
</taxonomicName>
and
<taxonomicName id="4BC34D05FFB3FF93FEFE7B2355F9FE4C" ID-CoL="427TT" box="[288,386,368,392]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="BEB7EA94FFB3FF93FEFE7B2355F9FE4C" box="[288,386,368,392]" italics="true" pageId="3" pageNumber="388">M. tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A58457EFFFB3FF93FE577B2255BDFE4C" box="[393,454,368,391]" pageId="3" pageNumber="388" rank="species">sp. n.</taxonomicNameLabel>
, and have a thicker shaft and slightly shorter brachia than those seen in
<taxonomicName id="4BC34D05FFB3FF93FB2F7B235112FE4C" ID-CoL="427TN" box="[1265,1385,368,392]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB3FF93FB2F7B235112FE4C" box="[1265,1385,368,392]" italics="true" pageId="3" pageNumber="388">M. martini</emphasis>
</taxonomicName>
. We consider these to be FS rather than SSB.
</paragraph>
<caption id="D8BC660EFFB3FF93FF497D0E5110F81C" httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="3" pageNumber="388" targetBox="[151,1436,483,1862]" targetPageId="3">
<paragraph id="8C7C3686FFB3FF93FF497D0E5110F81C" blockId="3.[151,1436,1884,2007]" pageId="3" pageNumber="388">
<emphasis id="BEB7EA94FFB3FF93FF497D0E552FF8B9" bold="true" box="[151,340,1884,1906]" pageId="3" pageNumber="388">FIGURES 1122.</emphasis>
Scanning electron micrographs of
<taxonomicName id="4BC34D05FFB3FF93FD1F7D0F507FF8B9" ID-CoL="427TN" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[705,1028,1884,1906]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="BEB7EA94FFB3FF93FD1F7D0F5703F8B9" box="[705,888,1885,1906]" italics="true" pageId="3" pageNumber="388">Messapus martini</emphasis>
Simon, 1898
</taxonomicName>
subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 1416 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 2021 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III.
</paragraph>
</caption>
<caption id="D8BC660EFFB4FF94FF497E8F5121FAA5" httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389" targetBox="[151,1436,193,1223]" targetPageId="4">
<paragraph id="8C7C3686FFB4FF94FF497E8F5121FAA5" blockId="4.[151,1436,1245,1390]" pageId="4" pageNumber="389">
<emphasis id="BEB7EA94FFB4FF94FF497E8F552EFB38" bold="true" box="[151,341,1245,1267]" pageId="4" pageNumber="389">FIGURES 2331.</emphasis>
Scanning electron micrographs of
<taxonomicName id="4BC34D05FFB4FF94FD617E8D571BFB3F" ID-CoL="427TT" box="[703,864,1247,1268]" class="Arachnida" family="Corinnidae" genus="Messapus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="389" phylum="Arthropoda" rank="species" species="tigris" status="sp. nov.">
<emphasis id="BEB7EA94FFB4FF94FD617E8D571BFB3F" box="[703,864,1247,1268]" italics="true" pageId="4" pageNumber="389">Messapus tigris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A58457EFFFB4FF94FCB67E8F57D9FB38" box="[872,930,1245,1267]" pageId="4" pageNumber="389" rank="species">
<emphasis id="BEB7EA94FFB4FF94FCB67E8F57D9FB38" bold="true" box="[872,930,1245,1267]" pageId="4" pageNumber="389">sp. n.</emphasis>
</taxonomicNameLabel>
female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum.
</paragraph>
</caption>
<paragraph id="8C7C3686FFB4FF94FF197FEC5530F8D0" blockId="4.[151,1436,1470,1820]" pageId="4" pageNumber="389">
The femora, tibiae and metatarsi are usually armed with relatively narrow, elongate spines with fine barbs (
<figureCitation id="14F82A03FFB4FF94FF417FB1555FFA37" box="[159,292,1507,1532]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Figs 1416</figureCitation>
,
<figureCitation id="14F82A03FFB4FF94FEEA7FB155F9FA37" box="[308,386,1507,1532]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">23, 25</figureCitation>
). Femora, patellae (
<figureCitation id="14F82A03FFB4FF94FDB37FB156B8FA30" box="[621,707,1507,1532]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">Fig. 24</figureCitation>
) and tibiae with dorsal lines of naked rugose cuticle. Patellar indentation narrow, with proximal lyriform organs (
<figureCitation id="14F82A03FFB4FF94FD3B7C545711F9D4" box="[741,874,1542,1567]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Figs 17, 18</figureCitation>
). Tibiae, metatarsi and tarsi with scattered dorsal trichobothria. Metatarsi with distinct dorsal vibration sense organ at distal end (
<figureCitation id="14F82A03FFB4FF94FBC77C795016F98F" box="[1049,1133,1579,1604]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Fig. 16</figureCitation>
). Tarsal claws paired, with several small teeth, claw tufts dense (
<figureCitation id="14F82A03FFB4FF94FD927C1C56DDF9AC" box="[588,678,1614,1639]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Figs 19</figureCitation>
,
<figureCitation id="14F82A03FFB4FF94FD6B7C1D56AFF9AC" box="[693,724,1615,1639]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">28</figureCitation>
); tarsi with tactile setae, chemosensory setae, trichobothria, a single tarsal slit sensilla and several small pores (
<figureCitation id="14F82A03FFB4FF94FCD67C21571EF940" box="[776,869,1651,1676]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">Fig. 27</figureCitation>
); trichobothria with sunken distal plate, tarsal trichobothria with more ridges (
<figureCitation id="14F82A03FFB4FF94FDCE7CC45617F964" box="[528,620,1686,1711]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Figs 20</figureCitation>
,
<figureCitation id="14F82A03FFB4FF94FDA37CC556E1F964" box="[637,666,1687,1711]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">29</figureCitation>
) than metatarsal trichobothria (
<figureCitation id="14F82A03FFB4FF94FBCE7CC45013F964" box="[1040,1128,1686,1711]" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Fig. 21</figureCitation>
); tarsal organ oval, finely wrinkled, only slightly elevated from surrounding integument, with an oval opening containing nerve endings (
<figureCitation id="14F82A03FFB4FF94FAB17CE954CCF93C" captionStart="FIGURES 11 22" captionStartId="3.[151,264,1884,1906]" captionTargetBox="[151,1436,483,1862]" captionTargetId="figure@3.[151,1436,483,1862]" captionTargetPageId="3" captionText="FIGURES 11 22. Scanning electron micrographs of Messapus martini Simon, 1898 subadult female: 11 Eye region, dorsal view; 12 Detail of feathery setae in eye region; 13 Same, anterior dorsal abdomen; 14 16 Spines and feathery setae on femur (14), tibia (15) and metatarsus (16) of leg I; 17 Patella I, patellar indentation (PI); 18 Same, detail of lyriform organ at proximal end of PI; 19 Tip of tarsus III; 20 21 Trichobothrial bases on tarsus I (20) and metatarsus III (21); 22 Tarsal organ, leg III." httpUri="https://zenodo.org/record/238363/files/figure.png" pageId="4" pageNumber="389">Fig. 22</figureCitation>
); tarsal pores sunken from cuticle, forming near circular pit (
<figureCitation id="14F82A03FFB4FF94FCB27C8C57BAF93C" box="[876,961,1758,1783]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">Fig. 30</figureCitation>
); tarsal slit sensillum small, with elevated ridges (
<figureCitation id="14F82A03FFB4FF94FF357D515545F8D0" box="[235,318,1795,1820]" captionStart="FIGURES 23 31" captionStartId="4.[151,264,1245,1267]" captionTargetBox="[151,1436,193,1223]" captionTargetId="figure@4.[151,1436,193,1224]" captionTargetPageId="4" captionText="FIGURES 23 31. Scanning electron micrographs of Messapus tigris sp. n. female, leg III: 23 Femur, spine, elongate striate scales (ESS) and squamose setae with brachia (SSB); 24 Patella, SSB, ESS and rugose longitudinal dorsal lines (RL); 25 Tibia, spine, SSB and ESS; 26 Metatarsus, SSB and ESS; 27 Distal part of tarsus, indicating trichobothriae (TR), tarsal pores (TP), tarsal slit sensillum (TS), tactile setae (TaS) and chemosensory seta (CS); note the much longer brachia and narrower shafts of the SSB in this part of tarsus; 28 Tarsal claw; 29 Proximal tarsal trichobothrium; 30 Tarsal pore; 31 Tarsal slit sensillum." httpUri="https://zenodo.org/record/238364/files/figure.png" pageId="4" pageNumber="389">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>