treatments-xml/data/03/EF/F2/03EFF252F31F7F7C14BFA1FBFB06FE12.xml

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<mods:title id="B9A2E5728BEAB11E0E117DBD638AF145">Biodiversity and notes on carabid beetles from Angola with description of new taxa (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="A90588E211DAB9810FD6BD28BF4B057F">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="26A08F9B1747F919721149B641F45E17">Capela, Rúben A.</mods:namePart>
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<treatment id="03EFF252F31F7F7C14BFA1FBFB06FE12" ID-DOI="http://doi.org/10.5281/zenodo.6006037" ID-GBIF-Taxon="137541570" ID-Zenodo-Dep="6006037" LSID="urn:lsid:plazi:treatment:03EFF252F31F7F7C14BFA1FBFB06FE12" httpUri="http://treatment.plazi.org/id/03EFF252F31F7F7C14BFA1FBFB06FE12" lastPageId="31" lastPageNumber="232" pageId="27" pageNumber="228">
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<paragraph id="8BF94344F31F7F7814BFA1FBFCE8FC1D" blockId="27.[151,795,904,963]" box="[151,795,904,930]" pageId="27" pageNumber="228">
<heading id="D0B1F428F31F7F7814BFA1FBFCE8FC1D" bold="true" box="[151,795,904,930]" fontSize="11" level="1" pageId="27" pageNumber="228" reason="1">
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<taxonomicName id="4C4638C7F31F7F7814BFA1FBFD22FC1D" ID-CoL="8MJRW" authority="Serrano &amp; Capela" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[151,721,904,930]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="liboloensis" status="sp. nov.">
<emphasis id="B9329F56F31F7F7814BFA1FBFEF5FC1D" bold="true" box="[151,262,904,930]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B9329F56F31F7F78153EA1FBFEBEFC1D" bold="true" box="[278,333,904,930]" italics="true" pageId="27" pageNumber="228">s. str</emphasis>
.) 
<emphasis id="B9329F56F31F7F78154DA1FBFE15FC1D" bold="true" box="[357,486,904,930]" italics="true" pageId="27" pageNumber="228">liboloensis</emphasis>
Serrano &amp; Capela
</taxonomicName>
<taxonomicNameLabel id="A201222DF31F7F7816F0A1FBFCE8FC1D" box="[728,795,904,930]" pageId="27" pageNumber="228" rank="species">sp. n.</taxonomicNameLabel>
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(
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, 
<figureCitation id="137D5FC1F31F7F781528A1D8FEE3FC7C" box="[256,272,939,963]" captionStart="FIGURE 2" captionStartId="29.[151,250,1733,1755]" captionTargetBox="[163,1418,191,1692]" captionTargetId="figure@29.[151,1436,173,1711]" captionTargetPageId="29" captionText="FIGURE 2. Perigona liboloensis sp. n.: a) Aedeagus, median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view), c) median lobe (right lateral view) and, d) genital ring." httpUri="https://zenodo.org/record/1065128/files/figure.png" pageId="27" pageNumber="228">2</figureCitation>
, 
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)
</paragraph>
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<paragraph id="8BF94344F31F7F7814BFA181FDEBFB24" blockId="27.[151,1437,1010,2010]" pageId="27" pageNumber="228">
<materialsCitation id="3B2E4919F31F7F7814BFA181FCC4FB90" ID-GBIF-Occurrence="1701951635" collectingDate="2015-12-04" collectionCode="ASC" collectorName="Serrano &amp; R. Capela" country="Angola" elevation="1116" latitude="-9.995" location="Calulo" longLatPrecision="21" longitude="14.842222" municipality="Alto Ventura-Col" pageId="27" pageNumber="228" specimenCount="1" specimenCount-male="1" stateProvince="Kwanza Sul" typeStatus="holotype">
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<typeStatus id="54FDFDE6F31F7F7814BFA181FF22FBB4" box="[151,209,1010,1035]" pageId="27" pageNumber="228">Type</typeStatus>
series.
</emphasis>
<typeStatus id="54FDFDE6F31F7F781501A181FE61FBB4" box="[297,402,1010,1035]" pageId="27" pageNumber="228" type="holotype">Holotype</typeStatus>
, 
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; 
<collectingCountry id="F35103D4F31F7F7815EAA181FDE0FBB4" box="[450,531,1010,1035]" name="Angola" pageId="27" pageNumber="228">Angola</collectingCountry>
(
<collectingRegion id="49828DA6F31F7F78160BA180FD5CFBB4" box="[547,687,1010,1035]" country="Angola" name="Kwanza Sul" pageId="27" pageNumber="228">
Kwanza 
<collectingRegion id="49828DA6F31F7F7816AFA181FD5CFBB4" box="[647,687,1010,1035]" country="Tanzania" name="Kusini Unguja" pageId="27" pageNumber="228">Sul</collectingRegion>
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), 
<collectingMunicipality id="6B9DD93EF31F7F7816EBA181FC71FBB4" box="[707,898,1010,1035]" pageId="27" pageNumber="228">Alto Ventura-Col</collectingMunicipality>
(
<location id="8E99159FF31F7F7817BAA181FC2DFBB4" LSID="urn:lsid:plazi:treatment:03EFF252F31F7F7C14BFA1FBFB06FE12:8E99159FF31F7F7817BAA181FC2DFBB4" box="[914,990,1010,1035]" country="Angola" latitude="-9.995" longLatPrecision="21" longitude="14.842222" municipality="Alto Ventura-Col" name="Calulo" pageId="27" pageNumber="228" stateProvince="Kwanza Sul">Calulo</location>
) (
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, 
<geoCoordinate id="EE722583F31F7F7810B0A180FACEFBB5" box="[1176,1341,1010,1035]" direction="east" orientation="longitude" pageId="27" pageNumber="228" precision="15" value="14.842222">14° 50´32´´ E</geoCoordinate>
, 
<quantity id="4CBEEEA1F31F7F781161A181FA6FFBB5" box="[1353,1436,1010,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.116" pageId="27" pageNumber="228" unit="m" value="1116.0">
<elevation id="006BA477F31F7F781161A181FA6FFBB5" box="[1353,1436,1010,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.116" pageId="27" pageNumber="228" unit="m" value="1116.0">1116 m</elevation>
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alt., 128), 
<date id="FFF86584F31F7F78152DA665FE73FB90" box="[261,384,1046,1071]" pageId="27" pageNumber="228" value="2015-12-04">
<collectingDate id="EFBC9C6CF31F7F78152DA665FE73FB90" box="[261,384,1046,1071]" pageId="27" pageNumber="228" value="2015-12-04">4.XII.2015</collectingDate>
</date>
, DO, A. 
<collectorName id="26B32692F31F7F7815C8A664FDC6FB90" box="[480,565,1047,1071]" pageId="27" pageNumber="228">Serrano</collectorName>
&amp; 
<collectorName id="26B32692F31F7F78167FA664FD30FB90" box="[599,707,1046,1071]" pageId="27" pageNumber="228">R. Capela</collectorName>
leg., 
<collectionCode id="ED57DB81F31F7F7816D6A664FCC4FB90" box="[766,823,1047,1071]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="27" pageNumber="228">ASC</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B2E4919F31F7F781769A665FDE7FB24" ID-GBIF-Occurrence="1701951681" collectingDate="2015-12-04" collectionCode="ASC" collectorName="Serrano &amp; R. Capela" country="Angola" county="Derivatio" elevation="1116" latitude="-9.995" location="Libolo" longLatPrecision="21" longitude="14.842222" municipality="This" pageId="27" pageNumber="228" specimenCount="1" specimenCount-female="1" stateProvince="Kwanza Sul Province" typeStatus="allotype">
<typeStatus id="54FDFDE6F31F7F781769A665FC50FB90" box="[833,931,1046,1071]" pageId="27" pageNumber="228" type="allotype">Allotype</typeStatus>
, 
<specimenCount id="9D4088CDF31F7F781786A664FC23FB90" box="[942,976,1047,1071]" pageId="27" pageNumber="228" type="female">1♀</specimenCount>
, same locality and date as 
<typeStatus id="54FDFDE6F31F7F7810DBA665FAA6FB90" box="[1267,1365,1046,1071]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED57DB81F31F7F781148A664FA6AFB90" box="[1376,1433,1047,1071]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15857" name="Northern Arizona University" pageId="27" pageNumber="228">ASC</collectionCode>
. 
<emphasis id="B9329F56F31F7F7814BFA648FE82FBEC" bold="true" box="[151,369,1083,1107]" italics="true" pageId="27" pageNumber="228">
<collectingCounty id="62983BC8F31F7F7814BFA648FEF0FBEC" box="[151,259,1083,1107]" pageId="27" pageNumber="228">Derivatio</collectingCounty>
nominis.
</emphasis>
<collectingMunicipality id="6B9DD93EF31F7F781553A649FE5EFBEC" box="[379,429,1082,1107]" pageId="27" pageNumber="228">This</collectingMunicipality>
species is named after 
<location id="8E99159FF31F7F7816EBA649FCFDFBEC" LSID="urn:lsid:plazi:treatment:03EFF252F31F7F7C14BFA1FBFB06FE12:8E99159FF31F7F7816EBA649FCFDFBEC" box="[707,782,1082,1107]" county="Derivatio" municipality="This" name="Libolo" pageId="27" pageNumber="228" stateProvince="Kwanza Sul Province">Libolo</location>
(noun in apposition), a municipality of the 
<collectingRegion id="49828DA6F31F7F781138A648FF08FBC8" country="Angola" name="Kwanza Sul" pageId="27" pageNumber="228">
<collectingRegion id="49828DA6F31F7F781138A648FA6FFBEC" box="[1296,1436,1082,1107]" inferredBySuffix="true" pageId="27" pageNumber="228">
Kwanza 
<collectingRegion id="49828DA6F31F7F78115FA649FA6FFBEC" box="[1399,1436,1082,1107]" country="Tanzania" inferredBySuffix="true" name="Kusini Unguja" pageId="27" pageNumber="228">Sul</collectingRegion>
</collectingRegion>
Province
</collectingRegion>
where the specimens were found and home of one of the most remarkable 
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: the 
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF31F7F7814EFA6D5FBB8FA94" pageId="27" pageNumber="228" type="diagnosis">
<paragraph id="8BF94344F31F7F7814EFA6D5FBB8FA94" blockId="27.[151,1437,1010,2010]" pageId="27" pageNumber="228">
<emphasis id="B9329F56F31F7F7814EFA6D5FEB2FB00" bold="true" box="[199,321,1190,1215]" pageId="27" pageNumber="228">Diagnosis.</emphasis>
Body elongate, sub-parallel, slightly convex and reddish-brown (
<figureCitation id="137D5FC1F31F7F78104CA6D5FB4EFB00" box="[1124,1213,1190,1215]" captionStart="FIGURE 1" captionStartId="28.[151,250,1253,1275]" captionTargetBox="[163,1412,183,1213]" captionTargetId="figure@28.[151,1436,167,1232]" captionTargetPageId="28" captionText="FIGURE 1. Facies of:a) Perigona liboloensis sp. n. male holotype, b) Lasiocera schuelei sp. n. male holotype." httpUri="https://zenodo.org/record/1065126/files/figure.png" pageId="27" pageNumber="228">Fig. 1a</figureCitation>
); integument with microreticulation, shiny; surface with sparse micropunctures visible at high magnification. Elytral color as 
<taxonomicName id="4C4638C7F31F7F7814BFA69CFDEEFAB8" authority="Castelnau, 1835" authorityName="Laporte de Castelnau" authorityYear="1835" box="[151,541,1262,1287]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9329F56F31F7F7814BFA69CFEABFAB8" box="[151,344,1262,1287]" italics="true" pageId="27" pageNumber="228">Perigona pallida</emphasis>
Castelnau, 1835
</taxonomicName>
. Internal sac of median lobe of aedeagus (
<figureCitation id="137D5FC1F31F7F781024A69DFBA1FAB8" box="[1036,1106,1262,1287]" captionStart="FIGURE 2" captionStartId="29.[151,250,1733,1755]" captionTargetBox="[163,1418,191,1692]" captionTargetId="figure@29.[151,1436,173,1711]" captionTargetPageId="29" captionText="FIGURE 2. Perigona liboloensis sp. n.: a) Aedeagus, median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view), c) median lobe (right lateral view) and, d) genital ring." httpUri="https://zenodo.org/record/1065128/files/figure.png" pageId="27" pageNumber="228">Fig. 2</figureCitation>
) with a conspicuous twisted sclerite closer to the apex and a posterior bone shaped cut half sclerite (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C35C10CFF31F7F7E14EFA745FE39F852" lastPageId="29" lastPageNumber="230" pageId="27" pageNumber="228" type="description">
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<emphasis id="B9329F56F31F7F7814EFA745FEAAFAF0" bold="true" box="[199,345,1334,1359]" pageId="27" pageNumber="228">Description.</emphasis>
Length of 
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: 
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. Length of 
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: 
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.
</paragraph>
<paragraph id="8BF94344F31F7F7814EFA729FAA6F9CD" blockId="27.[151,1437,1010,2010]" pageId="27" pageNumber="228">
<emphasis id="B9329F56F31F7F7814EFA729FEF7FACC" box="[199,260,1370,1395]" italics="true" pageId="27" pageNumber="228">Head</emphasis>
. Slightly wider than long [length: 
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(
<typeStatus id="54FDFDE6F31F7F781728A729FC9BFACC" box="[768,872,1370,1395]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
), 
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(
<typeStatus id="54FDFDE6F31F7F7817D8A729FBA3FACC" box="[1008,1104,1370,1395]" pageId="27" pageNumber="228" type="allotype">allotype</typeStatus>
), width: 
<quantity id="4CBEEEA1F31F7F78109CA728FAEAFACD" box="[1204,1305,1371,1395]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="27" pageNumber="228" unit="mm" value="0.72">0.72 mm</quantity>
(
<typeStatus id="54FDFDE6F31F7F781100A729FA63FACC" box="[1320,1424,1370,1395]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
), 
<quantity id="4CBEEEA1F31F7F7814BFA70DFF09FA29" box="[151,250,1406,1431]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="27" pageNumber="228" unit="mm" value="0.75">0.75 mm</quantity>
(
<typeStatus id="54FDFDE6F31F7F781520A70DFE99FA28" box="[264,362,1406,1431]" pageId="27" pageNumber="228" type="allotype">allotype</typeStatus>
)], darker (black-brownish) than pronotum and elytra, clypeus, labrum, mouth parts and antennae piceous; surface with sparce fine punctuation, microreticulation distinct, isodiametric, rather glossy; eyes large, moderately protruding, labrum moderately notched; clypeal-frontal furrows distinct, oblique, extended to level of anterior supraorbital seta; frons and occiput slightly convex; antenna short just attaining base of pronotum, scape two times longer than wide, 3rd to 9th antennomeres as long as wide, slightly globulose, last antennomere 1.6‒1.8 times longer than wide. Cephalic chaetotaxy (large setae): Labrum with three pairs of setae gradually shorter from exterior to inner sides, one pair on sides of clypeus and two pairs of supraocular setae present over each eye.
</paragraph>
<paragraph id="8BF94344F31F7F7814EFA40EFB58F865" blockId="27.[151,1437,1010,2010]" pageId="27" pageNumber="228">
<emphasis id="B9329F56F31F7F7814EFA40EFEEBF929" box="[199,280,1661,1686]" italics="true" pageId="27" pageNumber="228">Thorax</emphasis>
. Pronotum (
<figureCitation id="137D5FC1F31F7F78158BA40EFE02F929" box="[419,497,1661,1686]" captionStart="FIGURE 1" captionStartId="28.[151,250,1253,1275]" captionTargetBox="[163,1412,183,1213]" captionTargetId="figure@28.[151,1436,167,1232]" captionTargetPageId="28" captionText="FIGURE 1. Facies of:a) Perigona liboloensis sp. n. male holotype, b) Lasiocera schuelei sp. n. male holotype." httpUri="https://zenodo.org/record/1065126/files/figure.png" pageId="27" pageNumber="228">Fig. 1a</figureCitation>
) 1.27‒1.29 times wider than long [length: 
<quantity id="4CBEEEA1F31F7F7817F9A40EFBC6F92A" box="[977,1077,1661,1686]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="27" pageNumber="228" unit="mm" value="0.66">0.66 mm</quantity>
(
<typeStatus id="54FDFDE6F31F7F78106BA40EFB58F929" box="[1091,1195,1661,1686]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
), 
<quantity id="4CBEEEA1F31F7F781096A40DFAD1F92A" box="[1214,1314,1662,1686]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="27" pageNumber="228" unit="mm" value="0.74">0.74 mm</quantity>
(
<typeStatus id="54FDFDE6F31F7F781118A40EFA63F929" box="[1328,1424,1661,1686]" pageId="27" pageNumber="228" type="allotype">allotype</typeStatus>
), width: 
<quantity id="4CBEEEA1F31F7F7814C3A4D2FEA7F906" box="[235,340,1697,1722]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="27" pageNumber="228" unit="mm" value="0.85">0.85 mm</quantity>
(
<typeStatus id="54FDFDE6F31F7F781540A4D2FE23F905" box="[360,464,1697,1722]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
), 
<quantity id="4CBEEEA1F31F7F7815C0A4D1FDA2F906" box="[488,593,1698,1722]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="27" pageNumber="228" unit="mm" value="0.94">0.94 mm</quantity>
(
<typeStatus id="54FDFDE6F31F7F78164DA4D2FD34F905" box="[613,711,1697,1722]" pageId="27" pageNumber="228" type="allotype">allotype</typeStatus>
)], reddish-brown dark, wider than head, barely subcordiform, widest at first third; surface sparcely micropunctate, with microreticulation distinct, consisting of isodiametric meshes in diskal area and transverse meshes and lines laterally, shiny; apical margin not margined, lateral margins with complete and distinct margination continuing in the two thirds of the basal margin after the posterior angle; apical margin arcuate, apical angles roundly acute, protruded; lateral margin slightly curved at first two thirds, followed by a slight sinuosity and ending straight obliquely before the roundly obtuse basal angles, posterior margin distinctly emarginated; median line distinct, not reaching anterior or posterior margins, anterior transverse sulcus indistinct, posterior transverse sulcus slightly conspicuous; anterior marginal setae situated close to the widest width, posterior marginal seta situated at basal angle, none short setae at disk or sides.
</paragraph>
<caption id="DF3913CCF3187F7F14BFA696FAFBFB44" box="[151,1288,1253,1275]" httpUri="https://zenodo.org/record/1065126/files/figure.png" pageId="28" pageNumber="229" targetBox="[163,1412,183,1213]" targetPageId="28">
<paragraph id="8BF94344F3187F7F14BFA696FAFBFB44" blockId="28.[151,1288,1253,1275]" box="[151,1288,1253,1275]" pageId="28" pageNumber="229">
<emphasis id="B9329F56F3187F7F14BFA696FEE0FB45" bold="true" box="[151,275,1253,1275]" pageId="28" pageNumber="229">FIGURE 1.</emphasis>
Facies of: a) 
<taxonomicName id="4C4638C7F3187F7F15B4A695FD9CFB44" authority="Serrano &amp; Capela &amp; Santos, 2017" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[412,623,1253,1275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="liboloensis" status="sp. nov.">
<emphasis id="B9329F56F3187F7F15B4A695FD9CFB44" box="[412,623,1253,1275]" italics="true" pageId="28" pageNumber="229">Perigona liboloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201222DF3187F7F165DA695FD5EFB44" box="[629,685,1254,1275]" pageId="28" pageNumber="229" rank="species">
<emphasis id="B9329F56F3187F7F165DA695FD7EFB44" bold="true" box="[629,653,1254,1275]" pageId="28" pageNumber="229">sp</emphasis>
. 
<emphasis id="B9329F56F3187F7F16B1A695FD5EFB44" bold="true" box="[665,685,1254,1275]" pageId="28" pageNumber="229">n.</emphasis>
</taxonomicNameLabel>
male holotype, b) 
<taxonomicName id="4C4638C7F3187F7F1744A695FBD8FB44" authority="Serrano &amp; Capela &amp; Santos, 2017" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[876,1067,1253,1275]" class="Insecta" family="Carabidae" genus="Lasiocera" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="schuelei" status="sp. nov.">
<emphasis id="B9329F56F3187F7F1744A695FBD8FB44" box="[876,1067,1253,1275]" italics="true" pageId="28" pageNumber="229">Lasiocera schuelei</emphasis>
</taxonomicName>
<emphasis id="B9329F56F3187F7F1019A695FB99FB44" bold="true" box="[1073,1130,1254,1275]" pageId="28" pageNumber="229">
<taxonomicNameLabel id="A201222DF3187F7F1019A695FB99FB44" box="[1073,1130,1254,1275]" pageId="28" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
male holotype.
</paragraph>
</caption>
<paragraph id="8BF94344F3187F7F14EFA750FAEBF9DE" blockId="28.[151,1437,1315,1885]" pageId="28" pageNumber="229">
<emphasis id="B9329F56F3187F7F14EFA750FEFEFA83" box="[199,269,1315,1340]" italics="true" pageId="28" pageNumber="229">Elytra</emphasis>
(
<figureCitation id="137D5FC1F3187F7F1536A750FE9CFA84" box="[286,367,1315,1340]" captionStart="FIGURE 1" captionStartId="28.[151,250,1253,1275]" captionTargetBox="[163,1412,183,1213]" captionTargetId="figure@28.[151,1436,167,1232]" captionTargetPageId="28" captionText="FIGURE 1. Facies of:a) Perigona liboloensis sp. n. male holotype, b) Lasiocera schuelei sp. n. male holotype." httpUri="https://zenodo.org/record/1065126/files/figure.png" pageId="28" pageNumber="229">Fig. 1a</figureCitation>
) 1.61‒1.62 times longer than wide [length: 
<quantity id="4CBEEEA1F3187F7F1745A750FC27FA84" box="[877,980,1315,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="28" pageNumber="229" unit="mm" value="1.9">1.90 mm</quantity>
(
<typeStatus id="54FDFDE6F3187F7F17CCA750FBA1FA83" box="[996,1106,1315,1340]" pageId="28" pageNumber="229" type="holotype">Holotype</typeStatus>
), 
<quantity id="4CBEEEA1F3187F7F1040A750FB3DFA84" box="[1128,1230,1315,1339]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" pageId="28" pageNumber="229" unit="mm" value="1.97">1.97 mm</quantity>
(
<typeStatus id="54FDFDE6F3187F7F10F7A750FACCFA83" box="[1247,1343,1315,1340]" pageId="28" pageNumber="229" type="allotype">allotype</typeStatus>
), width: 
<quantity id="4CBEEEA1F3187F7F14BFA734FF0CFAE0" box="[151,255,1351,1375]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="28" pageNumber="229" unit="mm" value="1.17">1.17 mm</quantity>
(
<typeStatus id="54FDFDE6F3187F7F153AA735FE89FAE0" box="[274,378,1350,1375]" pageId="28" pageNumber="229" type="holotype">holotype</typeStatus>
), 
<quantity id="4CBEEEA1F3187F7F15B9A734FE09FAE0" box="[401,506,1351,1375]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="28" pageNumber="229" unit="mm" value="1.22">1.22 mm</quantity>
(
<typeStatus id="54FDFDE6F3187F7F1625A735FD9CFAE0" box="[525,623,1350,1375]" pageId="28" pageNumber="229" type="allotype">allotype</typeStatus>
)], reddish-brown lighter than pronotum, sutural stripe slightly darker; surface glossy, with sparse very fine punctuation and superficial microreticulation of fine transverse meshes; narrow and elongate, sub-parallel, dorsally moderately convex, very slightly depressed on disk, widest at middle; humeral angles distinct but broadly rounded; mediam marginal punctures arranged in a straight line; marginal channel towards apex widened, depressed, first half scarsely pilose, second half densely pilose; only inner three striae recognizable at two thirds by inconspicuous sulci not marked by punctures, anterior setiferous puncture situated at 3rd striae and at basal quarter, median setiferous puncture situated in 3rd interval close to 2nd striae slightly behind middle, apical setiferous puncture (long setae) situated at 3rd striae just above sub-apical border.
</paragraph>
<paragraph id="8BF94344F3187F7F14EFA41FFDB1F972" blockId="28.[151,1437,1315,1885]" pageId="28" pageNumber="229">Abdominal segments brown-yellowish, covered with dense dressed short hairs; last segment with 3 pairs (male) or 5 pairs (female) of setae near the posterior margin; terminal abdominal sternite in both sexes with fringe of short hairs. Legs brown-testaceous.</paragraph>
<paragraph id="8BF94344F3187F7F14EFA4ABFE76F8E2" blockId="28.[151,1437,1315,1885]" pageId="28" pageNumber="229">
<emphasis id="B9329F56F3187F7F14EFA4ABFEC7F94E" box="[199,308,1752,1777]" italics="true" pageId="28" pageNumber="229">Aedeagus</emphasis>
(
<figureCitation id="137D5FC1F3187F7F156CA4ABFE2EF94E" box="[324,477,1752,1777]" captionStart="FIGURE 2" captionStartId="29.[151,250,1733,1755]" captionTargetBox="[163,1418,191,1692]" captionTargetId="figure@29.[151,1436,173,1711]" captionTargetPageId="29" captionText="FIGURE 2. Perigona liboloensis sp. n.: a) Aedeagus, median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view), c) median lobe (right lateral view) and, d) genital ring." httpUri="https://zenodo.org/record/1065128/files/figure.png" pageId="28" pageNumber="229">Figs 2 a, b, c</figureCitation>
) with median lobe short and stout, ventral margin slightly curved, apex downwards (lateral view); internal sac with a conspicuous twisted sclerite and a bone shaped cut in half sclerite at right and left sides, respectively; genital ring (
<figureCitation id="137D5FC1F3187F7F1624A553FD9BF886" box="[524,616,1824,1849]" captionStart="FIGURE 2" captionStartId="29.[151,250,1733,1755]" captionTargetBox="[163,1418,191,1692]" captionTargetId="figure@29.[151,1436,173,1711]" captionTargetPageId="29" captionText="FIGURE 2. Perigona liboloensis sp. n.: a) Aedeagus, median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view), c) median lobe (right lateral view) and, d) genital ring." httpUri="https://zenodo.org/record/1065128/files/figure.png" pageId="28" pageNumber="229">Fig. 2 d</figureCitation>
) rather asymmetric, with wide, obliquely rounded apex, rather parallel, base evenly rounded.
</paragraph>
<caption id="DF3913CCF3197F7E14BFA4B6FCA1F946" httpUri="https://zenodo.org/record/1065128/files/figure.png" pageId="29" pageNumber="230" targetBox="[163,1418,191,1692]" targetPageId="29">
<paragraph id="8BF94344F3197F7E14BFA4B6FCA1F946" blockId="29.[151,1436,1732,1785]" pageId="29" pageNumber="230">
<emphasis id="B9329F56F3197F7E14BFA4B6FEE7F965" bold="true" box="[151,276,1733,1755]" pageId="29" pageNumber="230">FIGURE 2.</emphasis>
<taxonomicName id="4C4638C7F3197F7E1533A4B5FE1CF965" authority="Serrano &amp; Capela &amp; Santos, 2017" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[283,495,1732,1755]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="liboloensis" status="sp. nov.">
<emphasis id="B9329F56F3197F7E1533A4B5FE1CF965" box="[283,495,1732,1755]" italics="true" pageId="29" pageNumber="230">Perigona liboloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201222DF3197F7E15DEA4B6FDC0F965" box="[502,563,1733,1754]" pageId="29" pageNumber="230" rank="species">
<emphasis id="B9329F56F3197F7E15DEA4B6FDFDF965" bold="true" box="[502,526,1733,1754]" pageId="29" pageNumber="230">sp</emphasis>
. 
<emphasis id="B9329F56F3197F7E1634A4B6FDC0F965" bold="true" box="[540,563,1733,1754]" pageId="29" pageNumber="230">n.</emphasis>
</taxonomicNameLabel>
<emphasis id="B9329F56F3197F7E161BA4B6FDCAF965" bold="true" box="[563,569,1733,1754]" pageId="29" pageNumber="230">:</emphasis>
a) Aedeagus, median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view), c) median lobe (right lateral view) and, d) genital ring.
</paragraph>
</caption>
<paragraph id="8BF94344F3197F7E14EFA553FE39F852" blockId="29.[151,1437,1824,2030]" pageId="29" pageNumber="230">
<emphasis id="B9329F56F3197F7E14EFA553FE76F886" box="[199,389,1824,1849]" italics="true" pageId="29" pageNumber="230">Female genitalia</emphasis>
(
<figureCitation id="137D5FC1F3197F7E15BCA553FE26F886" box="[404,469,1824,1849]" captionStart="FIGURE 3" captionStartId="30.[151,250,1485,1507]" captionTargetBox="[569,1026,211,1420]" captionTargetId="figure@30.[515,1072,193,1464]" captionTargetPageId="30" captionText="FIGURE 3. Perigona liboloensis sp. n., female genitalia: stylomeres 1 and 2 (ventral view)." httpUri="https://zenodo.org/record/1065130/files/figure.png" pageId="29" pageNumber="230">Fig. 3</figureCitation>
) with the ordinary pattern shape of the genus. Stylomere 1 with two ventral large setae on subapical margin. Stylomere 2 with two medium sized ensiform ventro-lateral setae, one quite large ensiform dorso-median seta located about in middle of stylomere, and a single nematiform ventro-seta originating from a groove in apical third of stylomere. Some sensillae (probably chemoreceptors) are dispersed over ventral lateroapical margins of stylomere 1, and other sensillae over basal, medial and sub-apical areas of ventral, lateral and dorsal sides of stylomere 2.
</paragraph>
</subSubSection>
<caption id="DF3913CCF31A7F7D14BFA7BEFBB2FA5C" box="[151,1089,1485,1507]" httpUri="https://zenodo.org/record/1065130/files/figure.png" pageId="30" pageNumber="231" targetBox="[569,1026,211,1420]" targetPageId="30">
<paragraph id="8BF94344F31A7F7D14BFA7BEFBB2FA5C" blockId="30.[151,1089,1485,1507]" box="[151,1089,1485,1507]" pageId="30" pageNumber="231">
<emphasis id="B9329F56F31A7F7D14BFA7BEFEFEFA5D" bold="true" box="[151,269,1485,1507]" pageId="30" pageNumber="231">FIGURE 3</emphasis>
. 
<taxonomicName id="4C4638C7F31A7F7D1531A7BDFE18FA5C" authority="Serrano &amp; Capela &amp; Santos, 2017" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[281,491,1485,1507]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="liboloensis" status="sp. nov.">
<emphasis id="B9329F56F31A7F7D1531A7BDFE18FA5C" box="[281,491,1485,1507]" italics="true" pageId="30" pageNumber="231">Perigona liboloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201222DF31A7F7D15D9A7BDFDD9FA5C" box="[497,554,1486,1507]" pageId="30" pageNumber="231" rank="species">
<emphasis id="B9329F56F31A7F7D15D9A7BDFDFAFA5C" bold="true" box="[497,521,1486,1507]" pageId="30" pageNumber="231">sp</emphasis>
. 
<emphasis id="B9329F56F31A7F7D163EA7BDFDD9FA5C" bold="true" box="[534,554,1486,1507]" pageId="30" pageNumber="231">n.</emphasis>
</taxonomicNameLabel>
, female genitalia: stylomeres 1 and 2 (ventral view).
</paragraph>
</caption>
<subSubSection id="C35C10CFF31A7F7C14EFA479FB06FE12" lastPageId="31" lastPageNumber="232" pageId="30" pageNumber="231" type="discussion">
<paragraph id="8BF94344F31A7F7C14EFA479FAF5FF47" blockId="30.[151,1436,1546,2003]" lastBlockId="31.[151,1436,151,429]" lastPageId="31" lastPageNumber="232" pageId="30" pageNumber="231">
<emphasis id="B9329F56F31A7F7D14EFA479FEC8F99C" bold="true" box="[199,315,1546,1571]" pageId="30" pageNumber="231">Remarks.</emphasis>
The new species belongs to the subgenus 
<emphasis id="B9329F56F31A7F7D1719A47FFC24F99C" box="[817,983,1548,1571]" italics="true" pageId="30" pageNumber="231">
<taxonomicName id="4C4638C7F31A7F7D1719A47FFC6AF99C" authority="Serrano &amp; Capela &amp; Santos, 2017" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[817,921,1548,1571]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="genus" sensu="stricto">Perigona</taxonomicName>
<taxonomicNameLabel id="A201222DF31A7F7D178CA47FFC24F99C" box="[932,983,1548,1571]" pageId="30" pageNumber="231" sensu="stricto">s.str.</taxonomicNameLabel>
</emphasis>
based on the arrangement of the three lateral marginal elytral pores in a straight line and by the presence of distinct clypeo-orbital sulci. Until now there are six species known for continental Africa belonging to this subgenus (
<bibRefCitation id="EFD73EB5F31A7F7D17E8A421FB77F9D4" author="Basilewsky" box="[960,1156,1618,1643]" pageId="30" pageNumber="231" refString="Basilewsky, P. (1989) Revision des Perigonini d`Afrique (Coleoptera Carabidae). Revue de Zoologie Africaine, 103, 413 - 452." type="journal article" year="1989">Basilewsky 1989</bibRefCitation>
, Baehr 2004 and Serrano 2008). 
<taxonomicName id="4C4638C7F31A7F7D14CFA404FE3CF930" authority="Serrano &amp; Capela &amp; Santos, 2017" authorityName="Serrano &amp; Capela &amp; Santos" authorityYear="2017" box="[231,463,1654,1679]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="liboloensis" status="sp. nov.">
<emphasis id="B9329F56F31A7F7D14CFA404FE3CF930" box="[231,463,1654,1679]" italics="true" pageId="30" pageNumber="231">Perigona liboloensis</emphasis>
</taxonomicName>
<emphasis id="B9329F56F31A7F7D15FFA405FDE4F930" bold="true" box="[471,535,1654,1679]" pageId="30" pageNumber="231">
<taxonomicNameLabel id="A201222DF31A7F7D15FFA405FDE4F930" box="[471,535,1654,1679]" pageId="30" pageNumber="231" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is easily segregate from 
<taxonomicName id="4C4638C7F31A7F7D1718A404FB99F930" authority="Chaudoir, 1878" authorityName="Chaudoir" authorityYear="1878" box="[816,1130,1654,1679]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="parallela">
<emphasis id="B9329F56F31A7F7D1718A404FC5CF930" box="[816,943,1654,1679]" italics="true" pageId="30" pageNumber="231">P. parallela</emphasis>
Chaudoir, 1878
</taxonomicName>
, 
<taxonomicName id="4C4638C7F31A7F7D105FA404FF21F90C" authority="Basiewsky, 1953" authorityName="Basilewsky" authorityYear="1953" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="nigrociliata">
<emphasis id="B9329F56F31A7F7D105FA404FAE4F930" box="[1143,1303,1654,1679]" italics="true" pageId="30" pageNumber="231">P. nigrociliata</emphasis>
<bibRefCitation id="EFD73EB5F31A7F7D1135A405FF21F90C" author="Basiewsky" pageId="30" pageNumber="231" refString="Basilewsky, P. (1953) Exploration du Parc National de l'Upemba. Mission G. F. de Witte. 10. Carabidae (Coleoptera Adephaga). Institut des Parcs Nationaux du Congo Belge, Bruxelles, 252 pp." type="book" year="1953">Basiewsky, 1953</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C4638C7F31A7F7D14F7A4EFFDA0F90C" authority="Basilewsky, 1989" authorityName="Basilewsky" authorityYear="1989" box="[223,595,1690,1715]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="mediornata">
<emphasis id="B9329F56F31A7F7D14F7A4EFFE8DF90C" box="[223,382,1690,1715]" italics="true" pageId="30" pageNumber="231">P. mediornata</emphasis>
<bibRefCitation id="EFD73EB5F31A7F7D15A0A4E9FDA0F90C" author="Basilewsky" box="[392,595,1690,1715]" pageId="30" pageNumber="231" refString="Basilewsky, P. (1989) Revision des Perigonini d`Afrique (Coleoptera Carabidae). Revue de Zoologie Africaine, 103, 413 - 452." type="journal article" year="1989">Basilewsky, 1989</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C4638C7F31A7F7D1648A4EFFC82F90C" authority="Baehr, 2004" authorityName="Baehr" authorityYear="2004" box="[608,881,1690,1715]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="wachteli">
<emphasis id="B9329F56F31A7F7D1648A4EFFD2EF90C" box="[608,733,1690,1715]" italics="true" pageId="30" pageNumber="231">P. wachteli</emphasis>
Baehr, 2004
</taxonomicName>
and 
<taxonomicName id="4C4638C7F31A7F7D1784A4EFFB0CF90C" authority="Serrano, 2008" authorityName="Serrano" authorityYear="2008" box="[940,1279,1691,1715]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="principensis">
<emphasis id="B9329F56F31A7F7D1784A4EFFBA0F90C" box="[940,1107,1692,1715]" italics="true" pageId="30" pageNumber="231">P. principensis</emphasis>
Serrano, 2008
</taxonomicName>
, among other characters, by the quite dissimilar aedeagi of all these species. Certainly the new species is next related to 
<taxonomicName id="4C4638C7F31A7F7D111AA4CCFE21F944" authority="Laporte de Castelnau, 1835" authorityName="Laporte de Castelnau" authorityYear="1835" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9329F56F31A7F7D111AA4CCFA68F968" box="[1330,1435,1726,1751]" italics="true" pageId="30" pageNumber="231">P. pallida</emphasis>
Laporte de Castelnau, 1835
</taxonomicName>
with which it shares the general color, the elongate, almost parallel body shape and male median lobe general shape, though differs in its general body length (
<quantity id="4CBEEEA1F31A7F7D1021A575FB41F8A1" box="[1033,1202,1798,1823]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" metricValueMax="3.36" metricValueMin="3.16" pageId="30" pageNumber="231" unit="mm" value="3.26" valueMax="3.36" valueMin="3.16">3.16‒3.36 mm</quantity>
<emphasis id="B9329F56F31A7F7D1094A574FB20F8A1" box="[1212,1235,1799,1822]" italics="true" pageId="30" pageNumber="231">vs</emphasis>
. 
<quantity id="4CBEEEA1F31A7F7D10CDA574FA63F8A0" box="[1253,1424,1799,1823]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="30" pageNumber="231" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.30‒2.70 mm</quantity>
), pronotum conformation (slightly sinuouse before the posterior angles 
<emphasis id="B9329F56F31A7F7D1780A55FFC4CF8FC" box="[936,959,1836,1859]" italics="true" pageId="30" pageNumber="231">vs</emphasis>
. not sinuouse before the posterior angles), elytral index (1.61‒1.62 
<emphasis id="B9329F56F31A7F7D1585A53CFE3FF8D9" box="[429,460,1871,1894]" italics="true" pageId="30" pageNumber="231">vs.</emphasis>
1.47‒1.55), and in the shape of the sclerotized plates in the internal sac of the median lobe of aedeagus (
<emphasis id="B9329F56F31A7F7D154DA500FE72F834" box="[357,385,1907,1931]" italics="true" pageId="30" pageNumber="231">cf.</emphasis>
<figureCitation id="137D5FC1F31A7F7D15A0A501FE04F834" box="[392,503,1906,1931]" captionStart="FIGURE 2" captionStartId="29.[151,250,1733,1755]" captionTargetBox="[163,1418,191,1692]" captionTargetId="figure@29.[151,1436,173,1711]" captionTargetPageId="29" captionText="FIGURE 2. Perigona liboloensis sp. n.: a) Aedeagus, median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view), c) median lobe (right lateral view) and, d) genital ring." httpUri="https://zenodo.org/record/1065128/files/figure.png" pageId="30" pageNumber="231">Figs 2a, b</figureCitation>
<emphasis id="B9329F56F31A7F7D15D6A507FDE6F834" box="[510,533,1908,1931]" italics="true" pageId="30" pageNumber="231">vs</emphasis>
. 
<figureCitation id="137D5FC1F31A7F7D160CA501FD87F834" box="[548,628,1906,1931]" captionStart="FIGURE 5" captionStartId="32.[151,250,1754,1776]" captionTargetBox="[236,1344,993,1722]" captionTargetId="figure@32.[226,1360,978,1732]" captionTargetPageId="32" captionText="FIGURE 5. Lasiocera schuelei sp. n., Aedeagus: a) Median lobe and parameres (left lateral view), b) apex of median lobe (dorsal view)." httpUri="https://zenodo.org/record/1065134/files/figure.png" pageId="30" pageNumber="231">Fig. 5b</figureCitation>
in 
<bibRefCitation id="EFD73EB5F31A7F7D16B1A501FCADF834" author="Basilewsky" box="[665,862,1906,1931]" pageId="30" pageNumber="231" refString="Basilewsky, P. (1989) Revision des Perigonini d`Afrique (Coleoptera Carabidae). Revue de Zoologie Africaine, 103, 413 - 452." type="journal article" year="1989">Basilewsky 1989</bibRefCitation>
). Moreover, 
<taxonomicName id="4C4638C7F31A7F7D17C6A507FBAAF834" authorityName="Laporte de Castelnau" authorityYear="1835" box="[1006,1113,1906,1931]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9329F56F31A7F7D17C6A507FBAAF834" box="[1006,1113,1906,1931]" italics="true" pageId="30" pageNumber="231">P. pallida</emphasis>
</taxonomicName>
presents the pronotal widest situated in the middle while in the new species is in the first third. In the former species the elytral striae are conspicuous with punctuation well marked while in the latter the elytral striae are almost indistinct and without marked punctuation. By the 
<taxonomicName id="4C4638C7F31B7F7C15FEA2EAFD66FF0F" authorityName="Laporte de Castelnau" authorityYear="1835" box="[470,661,151,176]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B9329F56F31B7F7C15FEA2EAFD66FF0F" box="[470,661,151,176]" italics="true" pageId="31" pageNumber="232">Perigona pallida</emphasis>
</taxonomicName>
previous records (e.g. 
<bibRefCitation id="EFD73EB5F31B7F7C17BFA2E4FBAAFF0F" author="Basilewsky" box="[919,1113,151,176]" pageId="31" pageNumber="232" refString="Basilewsky, P. (1989) Revision des Perigonini d`Afrique (Coleoptera Carabidae). Revue de Zoologie Africaine, 103, 413 - 452." type="journal article" year="1989">Basilewsky 1989</bibRefCitation>
) and the new species known locality, the two taxa seem to be sister species with vicariant (allopatric) distribution, a fact referred already for tiger beetle and ant nest beetle species of 
<collectingCountry id="F35103D4F31B7F7C1640A2ACFD48FF47" box="[616,699,223,248]" name="Angola" pageId="31" pageNumber="232">Angola</collectingCountry>
also (Serrano 
<emphasis id="B9329F56F31B7F7C1774A292FC7DFF47" box="[860,910,223,248]" italics="true" pageId="31" pageNumber="232">et al</emphasis>
. 2015, 
<bibRefCitation id="EFD73EB5F31B7F7C17C9A293FB09FF47" author="Serrano" box="[993,1274,223,248]" pageId="31" pageNumber="232" refString="Serrano, A. R. M. &amp; Capela, R. A. (2015 a) Scientific note. New records of ant nest beetles (Coleoptera: Carabidae: Paussinae: Paussini) in Angola and an annotated list of species known from the country. The Coleopterists Bulletin, 69 (3), 525 - 528. https: // doi. org / 10.1649 / 0010 - 065 X- 69.3.525" type="journal article" year="2015" yearSuffix="a">Serrano &amp; Capela 2015a</bibRefCitation>
).
</paragraph>
<paragraph id="8BF94344F31B7F7C14EFA377FB06FE12" blockId="31.[151,1436,151,429]" pageId="31" pageNumber="232">
<emphasis id="B9329F56F31B7F7C14EFA377FE7DFEA2" bold="true" box="[199,398,260,285]" pageId="31" pageNumber="232">Ecological notes.</emphasis>
<collectingRegion id="49828DA6F31B7F7C15BDA377FE5AFEA3" box="[405,425,260,284]" country="Nigeria" name="Rivers" pageId="31" pageNumber="232">A</collectingRegion>
species known until now only from 
<collectingCountry id="F35103D4F31B7F7C176EA377FC6FFEA2" box="[838,924,260,285]" name="Angola" pageId="31" pageNumber="232">Angola</collectingCountry>
. Adults were found under the bark of a fallen tree within an abandoned coffee plantation (
<figureCitation id="137D5FC1F31B7F7C16B9A354FD10FEFF" box="[657,739,295,320]" captionStart="FIGURE 9" captionStartId="46.[151,250,1500,1522]" captionTargetBox="[151,1436,193,1478]" captionTargetId="figure@46.[151,1436,193,1478]" captionTargetPageId="46" captionText="FIGURE 9. Habitats of: a) Perigona liboloensis sp. n., Alto Ventura, Kwanza Sul, b) Lasiocera schuelei sp. n., Cambamba, Malanje, c) Galerita procera capelai ssp. n, Calulo-Cabuta, Kwanza Sul, d) Habrodera nilotica and Lophyra neglecta intermediola destroyed by sand removal (arrow) before Cambambe dan, Kwanza Sul, e) and f) Dromica fredericoi and Trichotaeniapepetela inNovember 2014 and 2015, well perserved and destroyed by fire, respectively, Catota, Bié." httpUri="https://zenodo.org/record/1065142/files/figure.png" pageId="31" pageNumber="232">Fig. 9a</figureCitation>
) with 
<taxonomicName id="4C4638C7F31B7F7C1707A35BFA99FEFF" authority="Laporte de Castelnau, 1835" authorityName="Laporte de Castelnau" authorityYear="1835" box="[815,1386,295,320]" class="Insecta" family="Carabidae" genus="Catascopus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="beauvoisi">
<emphasis id="B9329F56F31B7F7C1707A35BFBD5FEFF" box="[815,1062,295,320]" italics="true" pageId="31" pageNumber="232">Catascopus beauvoisi</emphasis>
Laporte de Castelnau, 1835
</taxonomicName>
and 
<taxonomicName id="4C4638C7F31B7F7C14BFA33FFD06FEDB" authority="Burgeon, 1937" authorityName="Burgeon" authorityYear="1937" box="[151,757,332,357]" class="Insecta" family="Carabidae" genus="Coptodera" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="congolensis" subGenus="Coptoderina">
<emphasis id="B9329F56F31B7F7C14BFA33FFEFCFEDA" box="[151,271,332,357]" italics="true" pageId="31" pageNumber="232">Coptodera</emphasis>
(
<emphasis id="B9329F56F31B7F7C1508A33FFE5EFEDA" box="[288,429,332,357]" italics="true" pageId="31" pageNumber="232">Coptoderina</emphasis>
) 
<emphasis id="B9329F56F31B7F7C1597A33FFDB0FEDA" box="[447,579,332,357]" italics="true" pageId="31" pageNumber="232">congolensis</emphasis>
Burgeon, 1937
</taxonomicName>
. Further adults of 
<taxonomicName id="4C4638C7F31B7F7C17EEA33EFBB9FEDB" box="[966,1098,333,356]" class="Insecta" kingdom="Animalia" order="Dermaptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="order">Dermaptera</taxonomicName>
[
<taxonomicName id="4C4638C7F31B7F7C1074A33FFACDFEDA" baseAuthorityName="Palisot De Beauvois" baseAuthorityYear="1805" box="[1116,1342,332,357]" class="Insecta" family="Apachyidae" genus="Apachyus" kingdom="Animalia" order="Dermaptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B9329F56F31B7F7C1074A33FFACDFEDA" box="[1116,1342,332,357]" italics="true" pageId="31" pageNumber="232">Apachyus depressus</emphasis>
</taxonomicName>
(Palisot de Beauvois, 1805 and 
<taxonomicName id="4C4638C7F31B7F7C15EFA31CFBFDFE37" authority="(Palisot de Beauvois, 1805)" baseAuthorityName="Palisot de Beauvois" baseAuthorityYear="1805" box="[455,1038,367,392]" class="Insecta" family="Pygidicranidae" genus="Echinosoma" kingdom="Animalia" order="Dermaptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="afrum">
<emphasis id="B9329F56F31B7F7C15EFA31CFD55FE37" box="[455,678,367,392]" italics="true" pageId="31" pageNumber="232">Echinosoma afrum</emphasis>
(Palisot de Beauvois, 1805)
</taxonomicName>
] and 
<taxonomicName id="4C4638C7F31B7F7C1049A31CFB2FFE37" box="[1121,1244,367,392]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="4C4638C7F31B7F7C10C5A31CFA6BFE37" authorityName="Leach" authorityYear="1815" box="[1261,1432,367,392]" class="Insecta" family="Endomychidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="family">Endomychidae</taxonomicName>
, 
<taxonomicName id="4C4638C7F31B7F7C14BFA3E7FEC2FE12" authorityName="Latreille" authorityYear="1802" box="[151,305,404,429]" class="Insecta" family="Trogossitidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="family">Trogossitidae</taxonomicName>
, 
<taxonomicName id="4C4638C7F31B7F7C1514A3E7FE2FFE12" authorityName="Latreille" authorityYear="1802" box="[316,476,404,429]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="232" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
(several species) were found together (syntopic) with the new species.
</paragraph>
</subSubSection>
</treatment>
</document>