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<document id="029C205A77F2B53FF5901E6618A4C215" ID-DOI="10.11646/zootaxa.4743.4.1" ID-GBIF-Dataset="44ef21b3-550b-4f82-950c-3d7db7e04de0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3690502" ID-ZooBank="4F90419E-E06F-4990-84F7-3D0B1BA9AF0D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582874459487" checkinUser="plazi" docAuthor="Ariyama, Hiroyuki" docDate="2020" docId="03DD878B5865CC73D3824CB4FCB6F810" docLanguage="en" docName="zootaxa.4743.4.1.pdf" docOrigin="Zootaxa 4743 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orientomaera incisa Ariyama 2020, sp. nov." docType="treatment" docUuid="2B14D59C-BA56-4FF4-BDD6-7BB8A3F4C824" docUuidSource="ZooBank" docVersion="7" lastPageNumber="472" masterDocId="FFE4FFF35871CC66D3154D6CFFD7FFDD" masterDocTitle="Species of the Maera - clade collected from Japan. Part 3: genera Maera Leach, 1814, Meximaera Barnard, 1969 and Orientomaera Ariyama, 2018 (addendum), with a key to Japanese species of the clade (Crustacea: Amphipoda: Maeridae)" masterLastPageNumber="479" masterPageNumber="451" pageNumber="471" updateTime="1698723202199" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6B41049F9494D1A7227D54DC00C506B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A64F30314C409A1EA4769CFD3ACB2549">
<mods:title id="2DCEB3012561B22C13A8BCA6881BD2BE">Species of the Maera - clade collected from Japan. Part 3: genera Maera Leach, 1814, Meximaera Barnard, 1969 and Orientomaera Ariyama, 2018 (addendum), with a key to Japanese species of the clade (Crustacea: Amphipoda: Maeridae)</mods:title>
</mods:titleInfo>
<mods:name id="29781D3EC8E2B2DBD2A9DB5F75320E62" type="personal">
<mods:role id="3C039725D44662D5AE20642D07EAD183">
<mods:roleTerm id="FE2D08A3D5A67178AAF4FC6A6DB146D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="48FEC6E34A127CBE912906074FE29EE7">Ariyama, Hiroyuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="73EA20354C10764AB8A5E9995D8266A0">text</mods:typeOfResource>
<mods:relatedItem id="D6B708E0DD250FEA3C61B3096B0CDDE2" type="host">
<mods:titleInfo id="32591DCE5AB7B461FFE761E942232B9B">
<mods:title id="1F292C2EBBB0B54EE20AAC9DDC3975AD">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="421C6F74A9A9C3760F41DBF14C93D2D5">
<mods:date id="D5F1C14D658C7ADDA50ED2CC0F41C53E">2020</mods:date>
<mods:detail id="08780976ACEEC85BBF574C0FCBD76C7A" type="pubDate">
<mods:number id="C7585590829D32A297AFD6DDDC19E604">2020-02-27</mods:number>
</mods:detail>
<mods:detail id="DB082359D2149CF295D31FC50FDB6396" type="volume">
<mods:number id="99DB91E362D3EDD30FF4A3A538BE56E4">4743</mods:number>
</mods:detail>
<mods:detail id="105360B975DA20A8008AFF3201A4B295" type="issue">
<mods:number id="371DDDFFB4B7ACED26C2AC7376ECA881">4</mods:number>
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<mods:start id="A15968F33F8E8DD476AE139FAAB403C5">451</mods:start>
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<mods:classification id="3A3AB81EAE1400F76F80D9C516E0A3AB">journal article</mods:classification>
<mods:identifier id="EB82321698EEE029D25C135721B15622" type="DOI">10.11646/zootaxa.4743.4.1</mods:identifier>
<mods:identifier id="C0361D88B6C3CE78D36B869CBC2A9B50" type="GBIF-Dataset">44ef21b3-550b-4f82-950c-3d7db7e04de0</mods:identifier>
<mods:identifier id="DE98570EA4609AC7C0AFA7055819AD66" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="874EB69C2EB9F45632FDFC756848814A" type="Zenodo-Dep">3690502</mods:identifier>
<mods:identifier id="994C3B217310F760CCCA7E16ABCB20E9" type="ZooBank">4F90419E-E06F-4990-84F7-3D0B1BA9AF0D</mods:identifier>
</mods:mods>
<treatment id="03DD878B5865CC73D3824CB4FCB6F810" ID-DOI="http://doi.org/10.5281/zenodo.3691406" ID-GBIF-Taxon="162361448" ID-Zenodo-Dep="3691406" LSID="urn:lsid:zoobank.org:act:2B14D59C-BA56-4FF4-BDD6-7BB8A3F4C824" httpUri="http://treatment.plazi.org/id/03DD878B5865CC73D3824CB4FCB6F810" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471">
<subSubSection id="C36E65165865CC72D3824CB4FD69FDCB" pageId="20" pageNumber="471" type="nomenclature">
<paragraph id="8BCB369D5865CC72D3824CB4FE22FE2F" blockId="20.[151,702,472,571]" box="[151,501,472,498]" pageId="20" pageNumber="471">
<heading id="D08381F15865CC72D3824CB4FE22FE2F" bold="true" box="[151,501,472,498]" fontSize="11" level="1" pageId="20" pageNumber="471" reason="1">
<emphasis id="B900EA8F5865CC72D3824CB4FE22FE2F" bold="true" box="[151,501,472,498]" pageId="20" pageNumber="471">
<taxonomicName id="4C744D1E5865CC72D3824CB4FE47FE2F" ID-CoL="7ZKSY" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[151,400,472,498]" class="Malacostraca" family="Maeridae" genus="Orientomaera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="incisa" status="sp. nov.">
<emphasis id="B900EA8F5865CC72D3824CB4FE47FE2F" bold="true" box="[151,400,472,498]" italics="true" pageId="20" pageNumber="471">Orientomaera incisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357F45865CC72D2824CB4FE22FE2F" box="[407,501,472,498]" pageId="20" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCB369D5865CC72D3824C91FD69FDCB" blockId="20.[151,702,472,571]" box="[151,702,507,535]" pageId="20" pageNumber="471">[Japanese name: Kirekomi-sunnariyokoebi, new]</paragraph>
</subSubSection>
<subSubSection id="C36E65165865CC72D3824F4CFEF0FDE6" box="[151,295,544,571]" pageId="20" pageNumber="471" type="description">
<paragraph id="8BCB369D5865CC72D3824F4CFEF0FDE6" blockId="20.[151,702,472,571]" box="[151,295,544,571]" pageId="20" pageNumber="471">
(
<figureCitation id="134F2A185865CC72D38A4F4CFEC8FDE6" box="[159,287,544,571]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStart-4="FIGURE 20" captionStartId-0="21.[151,250,963,988]" captionStartId-1="22.[151,250,1945,1970]" captionStartId-2="23.[151,250,1988,2013]" captionStartId-3="24.[151,250,1970,1995]" captionStartId-4="25.[151,250,1080,1105]" captionTargetBox-0="[158,1430,187,929]" captionTargetBox-1="[195,1394,189,1904]" captionTargetBox-2="[191,1388,187,1948]" captionTargetBox-3="[194,1387,193,1934]" captionTargetBox-4="[151,1436,181,1055]" captionTargetId-0="figure@21.[151,1436,181,937]" captionTargetId-1="figure@22.[184,1404,181,1919]" captionTargetId-2="figure@23.[182,1406,181,1962]" captionTargetId-3="figure@24.[184,1404,181,1945]" captionTargetId-4="figure@25.[151,1436,181,1055]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="25" captionText-0="FIGURE 16. Orientomaera incisa sp. nov. Holotype, male, 8.5 mm (OMNH-Ar-10737). Habitus." captionText-1="FIGURE 17. Orientomaera incisa sp. nov. All but M, holotype, male, 8.5 mm (OMNH-Ar-10737); M, paratype, male, 6.4 mm (OMNH-Ar-10739). Scales: 0.05 mm." captionText-2="FIGURE 18. Orientomaera incisa sp. nov. Holotype, male, 8.5 mm (OMNH-Ar-10737). Scales: 0.2 mm." captionText-3="FIGURE 19. Orientomaera incisa sp. nov. All except for F and SM, holotype, male, 8.5 mm (OMNH-Ar-10737); F, paratype, ovigerous female, 7.5 mm (OMNH-Ar-10738); SM, paratype, male, 5.6 mm (OMNH-Ar-10740). Scales: 0.1 mm." captionText-4="FIGURE 20. Orientomaera incisa sp. nov. Holotype, male, 6.4 mm (OMNH-Ar-10739). Live specimen, photograph by H. Ariyama." figureDoi-0="http://doi.org/10.5281/zenodo.3690534" figureDoi-1="http://doi.org/10.5281/zenodo.3690536" figureDoi-2="http://doi.org/10.5281/zenodo.3690538" figureDoi-3="http://doi.org/10.5281/zenodo.3690540" figureDoi-4="http://doi.org/10.5281/zenodo.3690542" httpUri-0="https://zenodo.org/record/3690534/files/figure.png" httpUri-1="https://zenodo.org/record/3690536/files/figure.png" httpUri-2="https://zenodo.org/record/3690538/files/figure.png" httpUri-3="https://zenodo.org/record/3690540/files/figure.png" httpUri-4="https://zenodo.org/record/3690542/files/figure.png" pageId="20" pageNumber="471">Figs 16–20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E65165865CC72D3824F04FC25FCCF" pageId="20" pageNumber="471" type="materials_examined">
<paragraph id="8BCB369D5865CC72D3824F04FED6FD33" blockId="20.[151,1437,615,2011]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3824F04FE90FD5F" bold="true" box="[151,327,615,642]" pageId="20" pageNumber="471">Type material.</emphasis>
<materialsCitation id="3B1C3CC05865CC72D25A4F04FADCFD7B" ID-GBIF-Occurrence="2572153714" collectingDate="2018-06-17" collectorName="H. Ariyama." latitude="34.266666" location="City" longLatPrecision="19" longitude="135.06084" municipality="Tagurazaki" pageId="20" pageNumber="471" specimenCount="1" stateProvince="Wakayama" typeStatus="holotype">
<typeStatus id="54CF883F5865CC72D25A4F04FE6BFD5F" box="[335,444,616,642]" pageId="20" pageNumber="471" type="holotype">Holotype</typeStatus>
: male,
<quantity id="4C8C9B785865CC72D11B4F0BFDB2FD5E" box="[526,613,615,643]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="471" unit="mm" value="8.5">8.5 mm</quantity>
(OMNH-Ar-10737),
<collectingMunicipality id="6BAFACE75865CC72D04F4F04FC01FD5F" box="[858,982,616,642]" pageId="20" pageNumber="471">Tagurazaki</collectingMunicipality>
coast,
<collectingRegion id="49B0F87F5865CC72D73D4F04FB73FD5F" box="[1064,1188,616,642]" country="Japan" name="Wakayama" pageId="20" pageNumber="471">Wakayama</collectingRegion>
<location id="8EAB60465865CC72D7B94F04FB0AFD5F" LSID="urn:lsid:plazi:treatment:03DD878B5865CC73D3824CB4FCB6F810:8EAB60465865CC72D7B94F04FB0AFD5F" box="[1196,1245,616,642]" latitude="34.266666" longLatPrecision="19" longitude="135.06084" municipality="Tagurazaki" name="City" pageId="20" pageNumber="471" stateProvince="Wakayama">City</location>
,
<collectingRegion id="49B0F87F5865CC72D7FC4F04FF3FFD7B" country="Japan" name="Wakayama" pageId="20" pageNumber="471">Wakayama Prefecture</collectingRegion>
,
<geoCoordinate id="EE40505A5865CC72D3E14FE0FEABFD7A" box="[244,380,651,679]" degrees="34" direction="north" minutes="16" orientation="latitude" pageId="20" pageNumber="471" precision="15" seconds="00" value="34.266666">34°16ʹ00ʹʹN</geoCoordinate>
,
<geoCoordinate id="EE40505A5865CC72D29D4FE7FDC4FD7A" box="[392,531,651,679]" degrees="135" direction="east" minutes="03" orientation="longitude" pageId="20" pageNumber="471" precision="15" seconds="39" value="135.06084">135°03ʹ39ʹE</geoCoordinate>
(
<figureCitation id="134F2A185865CC72D1364FE0FDADFD7B" box="[547,634,652,678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1627,1652]" captionTargetBox="[187,1400,182,1601]" captionTargetId="figure@2.[186,1400,181,1601]" captionTargetPageId="2" captionText="FIGURE 1. Map showing collecting sites of the examined specimens. A, Sagami Bay, off Jōgashima Island in Miura City, Kanagawa Prefecture; B, Tagurazaki coast, Wakayama City in Wakayama Prefecture; C, Sea of Japan, north of Masuda City in Yamaguchi Prefecture." figureDoi="http://doi.org/10.5281/zenodo.3690504" httpUri="https://zenodo.org/record/3690504/files/figure.png" pageId="20" pageNumber="471">Fig. 1B</figureCitation>
), intertidal, under stones,
<date id="FFCA105D5865CC72D0B44FE0FBECFD7B" box="[929,1083,652,679]" pageId="20" pageNumber="471" value="2018-06-17">
<collectingDate id="EF8EE9B55865CC72D0B44FE0FBECFD7B" box="[929,1083,652,679]" pageId="20" pageNumber="471" value="2018-06-17">17 June 2018</collectingDate>
</date>
, coll.
<collectorName id="2681534B5865CC72D7954FE1FADCFD7B" box="[1152,1291,652,679]" pageId="20" pageNumber="471">H. Ariyama.</collectorName>
</materialsCitation>
<materialsCitation id="3B1C3CC05865CC72D6074FE1FF2AFD33" ID-GBIF-Occurrence="2572153712" collectingDate="2018-06-17" collectorName="H. Ariyama." latitude="34.266666" location="City" longLatPrecision="19" longitude="135.06084" municipality="Tagurazaki" pageId="20" pageNumber="471" specimenCount="2" specimenCount-male="2" stateProvince="Wakayama" typeStatus="paratype">
<typeStatus id="54CF883F5865CC72D6074FE1FA54FD7A" box="[1298,1411,653,679]" pageId="20" pageNumber="471" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D72FD145865CC72D69B4FE0FF0CFD17" pageId="20" pageNumber="471" type="male">2 males</specimenCount>
, 6.4 and
<quantity id="4C8C9B785865CC72D2534FC3FE49FD16" box="[326,414,687,715]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="20" pageNumber="471" unit="mm" value="5.6">5.6 mm</quantity>
(OMNH-Ar-10739–10740); ovigerous female,
<quantity id="4C8C9B785865CC72D0A94FC3FBC3FD16" box="[956,1044,687,715]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="471" unit="mm" value="7.5">7.5 mm</quantity>
(OMNH-Ar-10738), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BCB369D5865CC72D3D24F94FC25FCCF" blockId="20.[151,1437,615,2011]" box="[199,1010,759,787]" pageId="20" pageNumber="471">
<materialsCitation id="3B1C3CC05865CC72D3D24F94FD65FCCF" ID-GBIF-Occurrence="2572153711" box="[199,690,759,787]" location="Tagurazaki" pageId="20" pageNumber="471" specimenCount="1" stateProvince="Wakayama" typeStatus="holotype">
<emphasis id="B900EA8F5865CC72D3D24F94FEB1FCCF" bold="true" box="[199,358,759,786]" pageId="20" pageNumber="471">
<typeStatus id="54CF883F5865CC72D3D24F94FED5FCCF" box="[199,258,760,786]" pageId="20" pageNumber="471">Type</typeStatus>
locality.
</emphasis>
<location id="8EAB60465865CC72D2794F94FE3FFCCF" LSID="urn:lsid:plazi:treatment:03DD878B5865CC73D3824CB4FCB6F810:8EAB60465865CC72D2794F94FE3FFCCF" box="[364,488,760,786]" name="Tagurazaki" pageId="20" pageNumber="471" stateProvince="Wakayama">Tagurazaki</location>
coast,
<collectingRegion id="49B0F87F5865CC72D1234F94FD65FCCF" box="[566,690,760,786]" country="Japan" name="Wakayama" pageId="20" pageNumber="471">Wakayama</collectingRegion>
</materialsCitation>
City,
<collectingRegion id="49B0F87F5865CC72D1E04F94FC38FCCF" box="[757,1007,760,786]" country="Japan" name="Wakayama" pageId="20" pageNumber="471">Wakayama Prefecture</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165865CC72D3D24E70FB5CFCEB" box="[199,1163,796,823]" pageId="20" pageNumber="471" type="etymology">
<paragraph id="8BCB369D5865CC72D3D24E70FB5CFCEB" blockId="20.[151,1437,615,2011]" box="[199,1163,796,823]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3D24E70FE99FCEB" bold="true" box="[199,334,796,822]" pageId="20" pageNumber="471">Etymology.</emphasis>
From the Latin
<emphasis id="B900EA8F5865CC72D1114E71FD85FCEB" box="[516,594,797,822]" italics="true" pageId="20" pageNumber="471">incisus</emphasis>
,
<emphasis id="B900EA8F5865CC72D14A4E71FDA1FCEB" box="[607,630,797,822]" italics="true" pageId="20" pageNumber="471">-a</emphasis>
,
<emphasis id="B900EA8F5865CC72D1904E71FD67FCEB" box="[645,688,797,822]" italics="true" pageId="20" pageNumber="471">-um</emphasis>
(=incised), referring to the shape of palm.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165865CC72D3D24E53FE1EFC1B" pageId="20" pageNumber="471" type="diagnosis">
<paragraph id="8BCB369D5865CC72D3D24E53FE1EFC1B" blockId="20.[151,1437,615,2011]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3D24E53FE95FC87" bold="true" box="[199,322,831,858]" pageId="20" pageNumber="471">Diagnosis.</emphasis>
<emphasis id="B900EA8F5865CC72D25D4E2CFE65FC87" box="[328,434,832,858]" italics="true" pageId="20" pageNumber="471">Antennae</emphasis>
, peduncles slender.
<emphasis id="B900EA8F5865CC72D1844E2CFCF3FC87" box="[657,804,832,858]" italics="true" pageId="20" pageNumber="471">Gnathopod 1</emphasis>
, coxa anteroventral corner produced acutely.
<emphasis id="B900EA8F5865CC72D60B4E2CFF72FCA3" italics="true" pageId="20" pageNumber="471">Gnathopod 2</emphasis>
, propodus subrectangular, palm slightly oblique, palmar margin with 2 large triangular teeth anteriorly and many robust setae.
<emphasis id="B900EA8F5865CC72D2384EE4FE02FC7E" box="[301,469,904,931]" italics="true" pageId="20" pageNumber="471">Pereopods 5–7</emphasis>
, bases subovate, posterior margins convex.
<emphasis id="B900EA8F5865CC72D0DF4EE4FBC5FC7F" box="[970,1042,904,930]" italics="true" pageId="20" pageNumber="471">Telson</emphasis>
wider than long, robust seta about 0.25 times length of telson.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165865CC73D3D24EA3FC48F8C5" lastPageId="21" lastPageNumber="472" pageId="20" pageNumber="471" type="description">
<paragraph id="8BCB369D5865CC72D3D24EA3FDF7FBD3" blockId="20.[151,1437,615,2011]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3D24EA3FE8EFC37" bold="true" box="[199,345,975,1002]" pageId="20" pageNumber="471">Description.</emphasis>
Based on
<typeStatus id="54CF883F5865CC72D2C44EBCFDE1FC37" box="[465,566,976,1002]" pageId="20" pageNumber="471" type="holotype">holotype</typeStatus>
, male,
<quantity id="4C8C9B785865CC72D1924EA3FD09FC36" box="[647,734,975,1003]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="20" pageNumber="471" unit="mm" value="8.5">8.5 mm</quantity>
(OMNH-Ar-10736) and
<typeStatus id="54CF883F5865CC72D0EE4EBDFB89FC36" box="[1019,1118,977,1003]" pageId="20" pageNumber="471" type="paratype">paratype</typeStatus>
, male,
<quantity id="4C8C9B785865CC72D7BA4EBCFAD1FC36" box="[1199,1286,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="20" pageNumber="471" unit="mm" value="6.4">6.4 mm</quantity>
(OMNH-Ar- 10739) for lower lip and maxilla 1.
</paragraph>
<paragraph id="8BCB369D5865CC72D3D24974FDF1FA47" blockId="20.[151,1437,615,2011]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3D24974FED0FBEF" bold="true" box="[199,263,1048,1074]" pageId="20" pageNumber="471">Head</emphasis>
(
<figureCitation id="134F2A185865CC72D2024974FE4EFBEE" box="[279,409,1048,1075]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="21.[151,250,963,988]" captionStartId-1="22.[151,250,1945,1970]" captionTargetBox-0="[158,1430,187,929]" captionTargetBox-1="[195,1394,189,1904]" captionTargetId-0="figure@21.[151,1436,181,937]" captionTargetId-1="figure@22.[184,1404,181,1919]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURE 16. Orientomaera incisa sp. nov. Holotype, male, 8.5 mm (OMNH-Ar-10737). Habitus." captionText-1="FIGURE 17. Orientomaera incisa sp. nov. All but M, holotype, male, 8.5 mm (OMNH-Ar-10737); M, paratype, male, 6.4 mm (OMNH-Ar-10739). Scales: 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3690534" figureDoi-1="http://doi.org/10.5281/zenodo.3690536" httpUri-0="https://zenodo.org/record/3690534/files/figure.png" httpUri-1="https://zenodo.org/record/3690536/files/figure.png" pageId="20" pageNumber="471">Figs 16–17</figureCitation>
).
<emphasis id="B900EA8F5865CC72D2A44974FDF2FBEF" box="[433,549,1048,1074]" italics="true" pageId="20" pageNumber="471">Antenna 1</emphasis>
about 45% body length, setose; peduncle slender, article 2 about 8.5 times as long as wide, ratio of lengths of articles 1–3 1:1.1:0.35, article 1 with 4 robust setae posteriorly; accessory flagellum with 8 articles, terminal article minute, length about half of primary flagellum; primary flagellum with 19 articles, terminal article minute.
<emphasis id="B900EA8F5865CC72D2B849E8FDF5FB43" box="[429,546,1156,1182]" italics="true" pageId="20" pageNumber="471">Antenna 2</emphasis>
about 0.65 times length of antenna 1, slender, setose; peduncle with ratio of lengths of articles 3–5 1:2.2:1.65; flagellum with 8 articles, terminal article minute.
<emphasis id="B900EA8F5865CC72D72949C4FB70FB1F" box="[1084,1191,1192,1218]" italics="true" pageId="20" pageNumber="471">Mandible</emphasis>
, left incisor bearing 5 cusps, right incisor with 4 cusps, left lacinia mobilis 5-dentate, right lacinia mobilis 4-dentate, number of accessory setae 9 on each side; palp article length ratio 1:2.5
<emphasis id="B900EA8F5865CC72D1AB499DFD1BFAD7" box="[702,716,1265,1290]" italics="true" pageId="20" pageNumber="471">–</emphasis>
2.6:1.7, articles 2–3 setose.
<emphasis id="B900EA8F5865CC72D0EB499CFBBEFAD7" box="[1022,1129,1264,1290]" italics="true" pageId="20" pageNumber="471">Maxilla 1</emphasis>
, inner plate with 3 plumose setae on tip, medial margin with several feeble setae; distal margin of outer plate with 8 robust setae; palp article 1 bearing many long feeble setae distolaterally.
<emphasis id="B900EA8F5865CC72D1A14854FCF6FA8F" box="[692,801,1336,1362]" italics="true" pageId="20" pageNumber="471">Maxilla 2</emphasis>
, medial margin of inner plate with a few long and short setae distally.
<emphasis id="B900EA8F5865CC72D22C4830FE65FAAB" box="[313,434,1372,1398]" italics="true" pageId="20" pageNumber="471">Maxilliped</emphasis>
, distomedial corner of inner plate with 3 robust setae; outer plate with 17 long-to-short robust setae on distomedial margin.
</paragraph>
<paragraph id="8BCB369D5865CC72D3D248C8FB90F966" blockId="20.[151,1437,615,2011]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3D248C8FECDFA63" bold="true" box="[199,282,1444,1470]" pageId="20" pageNumber="471">Pereon</emphasis>
(
<figureCitation id="134F2A185865CC72D23F48C8FEABFA63" box="[298,380,1444,1470]" captionStart="FIGURE 18" captionStartId="23.[151,250,1988,2013]" captionTargetBox="[191,1388,187,1948]" captionTargetId="figure@23.[182,1406,181,1962]" captionTargetPageId="23" captionText="FIGURE 18. Orientomaera incisa sp. nov. Holotype, male, 8.5 mm (OMNH-Ar-10737). Scales: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3690538" httpUri="https://zenodo.org/record/3690538/files/figure.png" pageId="20" pageNumber="471">Fig. 18</figureCitation>
).
<emphasis id="B900EA8F5865CC72D28648C8FDF1FA63" box="[403,550,1444,1470]" italics="true" pageId="20" pageNumber="471">Gnathopod 1</emphasis>
, coxa subquadrate, anteroventral corner produced acutely; basis with posterior margin bearing 5 long setae, proximomedial surface with 2 long setae; propodus palm defined by 3 short robust setae.
<emphasis id="B900EA8F5865CC72D3F44880FEA0F9DB" box="[225,375,1516,1542]" italics="true" pageId="20" pageNumber="471">Gnathopod 2</emphasis>
, coxa subtrapezoidal; basis short, anterodistal corner acutely projected, anteroproximal and posterior margins with 1 and 4 long setae, respectively; propodus subrectangular, slightly widened distally, length 1.36 times width; palm slightly oblique (angle of palmar corner: 92°), palmar margin with 2 large triangular teeth anteriorly (making incision) and 2 small teeth at posterior end, and with 4 lateral and 4 medial robust setae on anterior side of incision and 8 lateral and 6 medial robust setae on posterior side of incision; dactylus curved. Right gnathopod 2 similar to left one, but slightly smaller (ratio of propodus length: 96%).
</paragraph>
<paragraph id="8BCB369D5865CC72D3D24BA8FCD6F806" blockId="20.[151,1437,615,2011]" pageId="20" pageNumber="471">
<emphasis id="B900EA8F5865CC72D3D24BA8FEB8F903" box="[199,367,1732,1758]" italics="true" pageId="20" pageNumber="471">Pereopods 3–4</emphasis>
subequal in shape, pereopod 3 about 1.1 times length of pereopod 4; coxae subquadrate; bases posterior margins each bearing 3 long setae.
<emphasis id="B900EA8F5865CC72D19C4B84FCDEF8DE" box="[649,777,1768,1795]" italics="true" pageId="20" pageNumber="471">Pereopod 5</emphasis>
about 1.15 times length of pereopod 4; coxa with posterior lobe bearing robust seta; basis subovate, length about 1.9 times width, anterior margin with 7 short robust setae, posterior margin weakly convex.
<emphasis id="B900EA8F5865CC72D11C4A5CFD5FF897" box="[521,648,1840,1866]" italics="true" pageId="20" pageNumber="471">Pereopod 6</emphasis>
slightly longer than pereopod 5; coxa with 3 ventral robust setae on anterior lobe; basis subovate, length about 1.7 times width, anterior margin with 7 short robust setae, posterior margin weakly convex, bearing sparse short setae.
<emphasis id="B900EA8F5865CC72D16B4A14FCD7F84E" box="[638,768,1912,1939]" italics="true" pageId="20" pageNumber="471">Pereopod 7</emphasis>
about 1.2 times length of pereopod 6; coxa subtrapezoidal, anterior margin with robust seta; basis subovate, length about 1.6 times width, anterior margin with 9 short robust setae, posterior margin convex, with sparse short setae.
</paragraph>
<caption id="DF0B66155864CC73D3824EAFFBA9FC01" ID-DOI="http://doi.org/10.5281/zenodo.3690534" ID-Zenodo-Dep="3690534" box="[151,1150,963,988]" httpUri="https://zenodo.org/record/3690534/files/figure.png" pageId="21" pageNumber="472" startId="21.[151,250,963,988]" targetBox="[158,1430,187,929]" targetPageId="21">
<paragraph id="8BCB369D5864CC73D3824EAFFBA9FC01" blockId="21.[151,1150,963,988]" box="[151,1150,963,988]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3824EAFFEF7FC01" bold="true" box="[151,288,963,988]" pageId="21" pageNumber="472">FIGURE 16.</emphasis>
<taxonomicName id="4C744D1E5864CC73D2334EA8FE2EFC06" authority="Ariyama, 2020" authorityName="Ariyama" authorityYear="2020" box="[294,505,964,987]" class="Malacostraca" family="Maeridae" genus="Orientomaera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="incisa" status="sp. nov.">
<emphasis id="B900EA8F5864CC73D2334EA8FE2EFC06" box="[294,505,964,987]" italics="true" pageId="21" pageNumber="472">Orientomaera incisa</emphasis>
</taxonomicName>
<emphasis id="B900EA8F5864CC73D2EA4EA8FD98FC01" bold="true" box="[511,591,964,988]" pageId="21" pageNumber="472">
<taxonomicNameLabel id="A23357F45864CC73D2EA4EA8FD98FC01" box="[511,591,964,988]" pageId="21" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, male, 8.5 mm (OMNH-Ar-10737). Habitus.
</paragraph>
</caption>
<paragraph id="8BCB369D5864CC73D3D24966FB2EFA25" blockId="21.[151,1437,1034,1997]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24966FEDDFBF9" bold="true" box="[199,266,1034,1060]" pageId="21" pageNumber="472">Pleon</emphasis>
(
<figureCitation id="134F2A185864CC73D20F4966FEBBFBF9" box="[282,364,1034,1060]" captionStart="FIGURE 19" captionStartId="24.[151,250,1970,1995]" captionTargetBox="[194,1387,193,1934]" captionTargetId="figure@24.[184,1404,181,1945]" captionTargetPageId="24" captionText="FIGURE 19. Orientomaera incisa sp. nov. All except for F and SM, holotype, male, 8.5 mm (OMNH-Ar-10737); F, paratype, ovigerous female, 7.5 mm (OMNH-Ar-10738); SM, paratype, male, 5.6 mm (OMNH-Ar-10740). Scales: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3690540" httpUri="https://zenodo.org/record/3690540/files/figure.png" pageId="21" pageNumber="472">Fig. 19</figureCitation>
).
<emphasis id="B900EA8F5864CC73D2964967FDC2FBF9" box="[387,533,1034,1060]" italics="true" pageId="21" pageNumber="472">Epimera 1–3</emphasis>
, each posteroventral corner slightly projected (indistinct in epimeron 1), ventral margins with 2, 3 and 4 robust setae respectively; epimera 1–2 each with oblique lateral ridge.
<emphasis id="B900EA8F5864CC73D7A94942FA83FB95" box="[1212,1364,1070,1096]" italics="true" pageId="21" pageNumber="472">Pleopods 1–3</emphasis>
, inner rami with 7, 7, 6 articles and outer rami with 10, 9, 9 articles respectively; pleopod 3 shortest, peduncle bearing 4 and 2 setae on lateral margin and at posterodistal corner, respectively.
<emphasis id="B900EA8F5864CC73D0B1491AFBD8FB4D" box="[932,1039,1142,1168]" italics="true" pageId="21" pageNumber="472">Uropod 1</emphasis>
, peduncle with 8 dorsolateral and 6 dorsomedial robust setae; outer ramus slightly shorter than inner ramus, about 0.65 times length of peduncle, with 2 lateral, 3 medial and 5 terminal robust setae; inner ramus with 2 lateral, 5 medial and 5 terminal robust setae.
<emphasis id="B900EA8F5864CC73D65349D2FF72FB21" italics="true" pageId="21" pageNumber="472">Uropod 2</emphasis>
about 0.6 times length of uropod 1; peduncle bearing 2 dorsolateral and 2 dorsomedial robust setae, distolateral corner with robust seta; rami subequal, outer ramus almost same length as peduncle, with 5 lateral and 5 terminal robust setae; inner ramus with 2 lateral, 5 medial and 5 terminal robust setae.
<emphasis id="B900EA8F5864CC73D7174846FBB9FA99" box="[1026,1134,1322,1348]" italics="true" pageId="21" pageNumber="472">Uropod 3</emphasis>
about 0.75 times length of uropod 2; distolateral corner of peduncle with 3 robust setae; rami subequal, about 1.3 times length of peduncle; outer ramus bearing 3 groups of robust setae laterally and 2 single robust setae medially, inner ramus with 2 medial robust setae, longest distal seta about 1.15 times length of rami.
<emphasis id="B900EA8F5864CC73D04D48FAFC77FA6D" box="[856,928,1430,1456]" italics="true" pageId="21" pageNumber="472">Telson</emphasis>
wider than long, lateral margins convex, each with 2 sensory setae, distal margins of lobes each with 2 processes, inner process larger than outer, bearing single robust seta and sensory seta between processes, robust seta short, about 0.25 times as long as telson.
</paragraph>
<paragraph id="8BCB369D5864CC73D3D24B6DFE6AF955" blockId="21.[151,1437,1034,1997]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24B6DFE19F9C1" bold="true" box="[199,462,1537,1564]" pageId="21" pageNumber="472">Description of female.</emphasis>
Ovigerous female,
<typeStatus id="54CF883F5864CC73D1B94B6FFCD8F9C0" box="[684,783,1539,1565]" pageId="21" pageNumber="472" type="paratype">paratype</typeStatus>
,
<quantity id="4C8C9B785864CC73D00E4B6DFCA6F9C0" box="[795,881,1537,1565]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="21" pageNumber="472" unit="mm" value="7.5">7.5 mm</quantity>
(OMNH-Ar-10738).
<emphasis id="B900EA8F5864CC73D7714B6EFB20F9C1" box="[1124,1271,1538,1564]" italics="true" pageId="21" pageNumber="472">Gnathopod 2</emphasis>
(
<figureCitation id="134F2A185864CC73D6124B6EFA81F9C1" box="[1287,1366,1538,1564]" captionStart="FIGURE 19" captionStartId="24.[151,250,1970,1995]" captionTargetBox="[194,1387,193,1934]" captionTargetId="figure@24.[184,1404,181,1945]" captionTargetPageId="24" captionText="FIGURE 19. Orientomaera incisa sp. nov. All except for F and SM, holotype, male, 8.5 mm (OMNH-Ar-10737); F, paratype, ovigerous female, 7.5 mm (OMNH-Ar-10738); SM, paratype, male, 5.6 mm (OMNH-Ar-10740). Scales: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3690540" httpUri="https://zenodo.org/record/3690540/files/figure.png" pageId="21" pageNumber="472">Fig. 19</figureCitation>
F-G2) subsimilar to that of
<typeStatus id="54CF883F5864CC73D26B4B4AFE34F99D" box="[382,483,1574,1600]" pageId="21" pageNumber="472" type="holotype">holotype</typeStatus>
, male, palm with 4 lateral and 3 medial robust setae on anterior side of incision and 8 lateral and 5 medial robust setae on posterior side of incision; left gnathopod 2 slightly smaller than right one (ratio of propodus length: 97%).
</paragraph>
<paragraph id="8BCB369D5864CC73D3D24BFDFEE1F929" blockId="21.[151,1437,1034,1997]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24BFDFEE8F971" bold="true" box="[199,319,1681,1708]" pageId="21" pageNumber="472">Variation.</emphasis>
Male
<typeStatus id="54CF883F5864CC73D29D4BFFFE3CF970" box="[392,491,1683,1709]" pageId="21" pageNumber="472" type="paratype">paratype</typeStatus>
,
<quantity id="4C8C9B785864CC73D2ED4BFDFD98F970" box="[504,591,1681,1709]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="21" pageNumber="472" unit="mm" value="5.6">5.6 mm</quantity>
(OMNH-Ar-10740).
<emphasis id="B900EA8F5864CC73D0514BFEFC0FF971" box="[836,984,1682,1708]" italics="true" pageId="21" pageNumber="472">Gnathopod 2</emphasis>
(
<figureCitation id="134F2A185864CC73D0FD4BFEFBEFF971" box="[1000,1080,1682,1708]" captionStart="FIGURE 19" captionStartId="24.[151,250,1970,1995]" captionTargetBox="[194,1387,193,1934]" captionTargetId="figure@24.[184,1404,181,1945]" captionTargetPageId="24" captionText="FIGURE 19. Orientomaera incisa sp. nov. All except for F and SM, holotype, male, 8.5 mm (OMNH-Ar-10737); F, paratype, ovigerous female, 7.5 mm (OMNH-Ar-10738); SM, paratype, male, 5.6 mm (OMNH-Ar-10740). Scales: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3690540" httpUri="https://zenodo.org/record/3690540/files/figure.png" pageId="21" pageNumber="472">Fig. 19</figureCitation>
SM-G2) almost same as that of
<typeStatus id="54CF883F5864CC73D3824BDAFF2BF90D" box="[151,252,1718,1744]" pageId="21" pageNumber="472" type="holotype">holotype</typeStatus>
, palm with 3 lateral and 2 medial robust setae on anterior side and 7 lateral and 6 medial robust setae on posterior side.
</paragraph>
<paragraph id="8BCB369D5864CC73D3D24B91FC48F8C5" blockId="21.[151,1437,1034,1997]" box="[199,927,1789,1817]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24B91FE44F8C5" bold="true" box="[199,403,1789,1816]" pageId="21" pageNumber="472">Coloration in life</emphasis>
(
<figureCitation id="134F2A185864CC73D2B74B92FE23F8C5" box="[418,500,1790,1816]" captionStart="FIGURE 20" captionStartId="25.[151,250,1080,1105]" captionTargetBox="[151,1436,181,1055]" captionTargetId="figure@25.[151,1436,181,1055]" captionTargetPageId="25" captionText="FIGURE 20. Orientomaera incisa sp. nov. Holotype, male, 6.4 mm (OMNH-Ar-10739). Live specimen, photograph by H. Ariyama." figureDoi="http://doi.org/10.5281/zenodo.3690542" httpUri="https://zenodo.org/record/3690542/files/figure.png" pageId="21" pageNumber="472">Fig. 20</figureCitation>
). Eyes dark red, other parts pale pink.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165864CC73D3D24A4EFD5CF875" pageId="21" pageNumber="472" type="discussion">
<paragraph id="8BCB369D5864CC73D3D24A4EFB38F8BD" blockId="21.[151,1437,1034,1997]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24A4EFEECF8E1" bold="true" box="[199,315,1826,1852]" pageId="21" pageNumber="472">Remarks.</emphasis>
This new species apparently has the character of
<taxonomicName id="4C744D1E5864CC73D04E4A4FFC2DF8E0" authorityName="Ariyama" authorityYear="2018" box="[859,1018,1827,1853]" class="Malacostraca" family="Maeridae" genus="Orientomaera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="B900EA8F5864CC73D04E4A4FFC2DF8E0" box="[859,1018,1827,1853]" italics="true" pageId="21" pageNumber="472">Orientomaera</emphasis>
</taxonomicName>
except for the shape of the gnathopod 2 palm.
<taxonomicName id="4C744D1E5864CC73D3E74A2BFE0CF8BD" authorityName="Ariyama" authorityYear="2020" box="[242,475,1863,1889]" class="Malacostraca" family="Maeridae" genus="Orientomaera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="B900EA8F5864CC73D3E74A2BFE0CF8BD" box="[242,475,1863,1889]" italics="true" pageId="21" pageNumber="472">Orientomaera incisa</emphasis>
</taxonomicName>
is different from its congeners in the incised palm of the gnathopod 2.
</paragraph>
<paragraph id="8BCB369D5864CC73D3D24A05FD5CF875" blockId="21.[151,1437,1034,1997]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24A05FEFDF859" bold="true" box="[199,298,1897,1924]" pageId="21" pageNumber="472">Habitat.</emphasis>
Under stones, intertidal. This species is collected from similar habitat of
<taxonomicName id="4C744D1E5864CC73D74F4A07FECDF875" authority="Ariyama, 2018" authorityName="Ariyama" authorityYear="2018" class="Malacostraca" family="Maeridae" genus="Orientomaera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B900EA8F5864CC73D74F4A07FABEF859" box="[1114,1385,1898,1925]" italics="true" pageId="21" pageNumber="472">Orientomaera decipiens</emphasis>
<bibRefCitation id="EFE54B6C5864CC73D67B4A06FECDF875" author="Ariyama, H." pageId="21" pageNumber="472" pagination="201 - 244" refId="ref10982" refString="Ariyama, H. (2018) Species of the Maera - clade collected from Japan. Part 1: genera Maeropsis Chevreux, 1919 and Orientomaera gen. nov. (Crustacea: Amphipoda: Maeridae). Zootaxa, 4433 (2), 201 - 244. https: // doi. org / 10.11646 / zootaxa. 4433.2.1" type="journal article" year="2018">Ariyama, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C744D1E5864CC73D2454AE3FD50F875" authority="Ariyama, 2018" authorityName="Ariyama" authorityYear="2018" box="[336,647,1934,1961]" class="Malacostraca" family="Maeridae" genus="Orientomaera" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Amphipoda" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="obliquua">
<emphasis id="B900EA8F5864CC73D2454AE3FE01F875" box="[336,470,1934,1961]" italics="true" pageId="21" pageNumber="472">O. obliquua</emphasis>
<bibRefCitation id="EFE54B6C5864CC73D2CE4AE2FD50F875" author="Ariyama, H." box="[475,647,1934,1960]" pageId="21" pageNumber="472" pagination="201 - 244" refId="ref10982" refString="Ariyama, H. (2018) Species of the Maera - clade collected from Japan. Part 1: genera Maeropsis Chevreux, 1919 and Orientomaera gen. nov. (Crustacea: Amphipoda: Maeridae). Zootaxa, 4433 (2), 201 - 244. https: // doi. org / 10.11646 / zootaxa. 4433.2.1" type="journal article" year="2018">Ariyama, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E65165864CC73D3D24ADDFCB6F810" box="[199,865,1969,1997]" pageId="21" pageNumber="472" type="distribution">
<paragraph id="8BCB369D5864CC73D3D24ADDFCB6F810" blockId="21.[151,1437,1034,1997]" box="[199,865,1969,1997]" pageId="21" pageNumber="472">
<emphasis id="B900EA8F5864CC73D3D24ADDFEB6F811" bold="true" box="[199,353,1969,1996]" pageId="21" pageNumber="472">Distribution.</emphasis>
<collectingCountry id="F363760D5864CC73D2724ADFFE7CF810" box="[359,427,1971,1997]" name="Japan" pageId="21" pageNumber="472">Japan</collectingCountry>
:
<collectingRegion id="49B0F87F5864CC73D2A04ADEFD7CF811" box="[437,683,1970,1996]" country="Japan" name="Wakayama" pageId="21" pageNumber="472">Wakayama Prefecture</collectingRegion>
(present study).
</paragraph>
</subSubSection>
</treatment>
</document>
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