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22 KiB
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211 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.906.47978" ID-GBIF-Dataset="a8cc42c7-e9ad-42fc-ba79-78daea513354" ID-PMC="PMC6989643" ID-Pensoft-Pub="1313-2970-906-13" ID-Pensoft-UUID="33B523D44A1E5FECBC706B74B9397519" ID-PubMed="32021555" ID-ZooBank="EF6D38B4B1A3402BB7642DACCA490AC7" ModsDocID="1313-2970-906-13" checkinTime="1579803439732" checkinUser="pensoft" docAuthor="Zamani, Alireza, Marusik, Yuri M. & Sestakova, Anna" docDate="2020" docId="F0071CFFB12051E28A5E683A1F9DE49A" docLanguage="en" docName="ZooKeys 906: 13-40" docOrigin="ZooKeys 906" docSource="http://dx.doi.org/10.3897/zookeys.906.47978" docTitle="Neoscona theisi *" docType="treatment" docVersion="4" id="33B523D44A1E5FECBC706B74B9397519" lastPageNumber="13" masterDocId="33B523D44A1E5FECBC706B74B9397519" masterDocTitle="On Araniella and Neoscona (Araneae, Araneidae) of the Caucasus, Middle East and Central Asia" masterLastPageNumber="40" masterPageNumber="13" pageNumber="13" updateTime="1668168350213" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>On Araniella and Neoscona (Araneae, Araneidae) of the Caucasus, Middle East and Central Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sestakova, Anna</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>906</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>13</mods:start>
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<mods:end>40</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.906.47978</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.906.47978</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-906-13</mods:identifier>
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<mods:identifier type="ZooBank">EF6D38B4B1A3402BB7642DACCA490AC7</mods:identifier>
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<mods:identifier type="Pensoft-UUID">33B523D44A1E5FECBC706B74B9397519</mods:identifier>
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<treatment ID-GBIF-Taxon="161838079" LSID="urn:lsid:plazi:treatment:F0071CFFB12051E28A5E683A1F9DE49A" httpUri="http://treatment.plazi.org/id/F0071CFFB12051E28A5E683A1F9DE49A" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection pageId="0" pageNumber="13" type="nomenclature">
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<paragraph pageId="0" pageNumber="13">
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<taxonomicName LSID="F0071CFF-B120-51E2-8A5E-683A1F9DE49A" authority="(Walckenaer, 1841) *" authorityName="*" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona theisi" order="Araneae" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="theisi">Neoscona theisi (Walckenaer, 1841)*</taxonomicName>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Dorsal habitus of Neoscona theisi (A, B), N. spasskyi (C) and N. isatis sp. nov. (D, E). A, C, D Females B, E males." figureDoi="10.3897/zookeys.906.47978.figure11" httpUri="https://binary.pensoft.net/fig/374413" pageId="0" pageNumber="13">Figs 11A, B</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Neoscona spasskyi (A, B) and N. theisi (C, D). A-C Dorsal D ventral C, D showing variations in comparison to specimens depicted in Figure 11. Photos C, D courtesy of A. Seropian." figureDoi="10.3897/zookeys.906.47978.figure12" httpUri="https://binary.pensoft.net/fig/374414" pageId="0" pageNumber="13">; 12C, D</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Females of Neoscona theisi (A, B), N. spasskyi (C, D) and N. isatis sp. nov. (E, F). A, C, E Prosoma, ventral B, D, F abdomen, ventral. Photos A, B courtesy of A. Seropian." figureDoi="10.3897/zookeys.906.47978.figure13" httpUri="https://binary.pensoft.net/fig/374415" pageId="0" pageNumber="13">; 13A, B</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Male palps and tibiae II of Neoscona theisi (A, D), N. isatis sp. nov. (B, F) and N. spasskyi (C, E). A-C Male palp, prolateral D-F male tibia II, ventral. Abbreviations: Co conductor, La lamella, Ma median apophysis, Me extension of median apophysis, Ta terminal apophysis, Te tegulum. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.906.47978.figure14" httpUri="https://binary.pensoft.net/fig/374416" pageId="0" pageNumber="13">; 14A, D</figureCitation>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Epigynes of Neoscona theisi (A-C), N. spasskyi (D-F) and N. isatis sp. nov. (G-I). A, D, G Ventral B, E, H posterior C, F, I lateral. Abbreviations: Le lateral extension, Sc scape. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.906.47978.figure17" httpUri="https://binary.pensoft.net/fig/374419" pageId="0" pageNumber="13">; 17A-C</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution records of Araniella mithra sp. nov. (blue circle), A. villanii sp. nov. (violet star), Neoscona isatis sp. nov. (green pentagon), N. spasskyi (black square), N. theisi (gray triangle, only new records) and N. tedgenica (brown asterisk)." figureDoi="10.3897/zookeys.906.47978.figure18" httpUri="https://binary.pensoft.net/fig/374420" pageId="0" pageNumber="13">; 18</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="reference_group">
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<paragraph pageId="0" pageNumber="13">
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<taxonomicName authorityName="*" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona theisi" order="Araneae" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="theisi">Neoscona theisi</taxonomicName>
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:
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<bibRefCitation author="Grasshoff, M" journalOrPublisher="Annalen Zoologische Wetenschappen" pageId="0" pageNumber="13" pagination="1 - 123" refId="B10" refString="Grasshoff, M, 1986. Die Radnetzspinnen-Gattung Neoscona in Afrika (Arachnida: Araneae). Annalen Zoologische Wetenschappen 250: 1 - 123" title="Die Radnetzspinnen-Gattung Neoscona in Afrika (Arachnida: Araneae)." volume="250" year="1986">Grasshoff 1986</bibRefCitation>
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: 69, fig. 90-100 (♂♀);
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<bibRefCitation DOI="https://doi.org/10.2476/asjaa.47.133" author="Tanikawa, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="13" pagination="133 - 168" refId="B22" refString="Tanikawa, A, 1998. A revision of the Japanese spiders of the genus Neoscona (Araneae: Araneidae). Acta Arachnologica 47: 133 - 168, DOI: https://doi.org/10.2476/asjaa.47.133" title="A revision of the Japanese spiders of the genus Neoscona (Araneae: Araneidae)." url="https://doi.org/10.2476/asjaa.47.133" volume="47" year="1998">Tanikawa 1998</bibRefCitation>
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: 137, fig. 1-8, 11-17 (♂♀);
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<bibRefCitation author="Tanikawa, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="13" refId="B23" refString="Tanikawa, A, 2007. An identification guide to the Japanese spiders of the families Araneidae, Nephilidae and Tetragnathidae. Arachnological Society of Japan, 121 pp." title="An identification guide to the Japanese spiders of the families Araneidae, Nephilidae and Tetragnathidae. Arachnological Society of Japan, 121 pp." year="2007">Tanikawa 2007</bibRefCitation>
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: 67, fig. 150-159, 572-574 (♂♀).
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</paragraph>
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<paragraph pageId="0" pageNumber="13">
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<taxonomicName authorityName="Levy" authorityYear="1998" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona sodom" order="Araneae" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="sodom">Neoscona sodom</taxonomicName>
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Levy, 1998: 340, fig. 117-126 (♂♀).
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<emphasis bold="true" pageId="0" pageNumber="13">syn. nov.</emphasis>
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</paragraph>
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<paragraph pageId="0" pageNumber="13">
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<taxonomicName authorityName="Levy" authorityYear="1998" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona sodom" order="Araneae" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="sodom">Neoscona sodom</taxonomicName>
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:
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<bibRefCitation author="Bosmans, R" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="13" refId="B8" refString="Bosmans, R, Van Keer, J, Russell-Smith, A, Hadjiconstantis, M, Komnenov, M, Bosselaers, J, Huber, S, Mccowan, D, Snazell, R, Decae, A, Zoumides, C, Kielhorn, K-H, Oger, P, 2019. Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Arachnological Contributions. Newsletter of the Belgian Arachnological Society, 34(suppl.): 1-197." title="Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Arachnological Contributions. Newsletter of the Belgian Arachnological Society, 34 (suppl.): 1 - 197." year="2019">Bosmans et al. 2019</bibRefCitation>
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: 9, fig. 1a-e (♂).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="materials_examined">
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<paragraph pageId="0" pageNumber="13">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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Iran: 2♂ 5♀ (MHNG), Mazandaran Province: around Amol,
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<geoCoordinate degrees="36" direction="north" minutes="18" orientation="latitude" precision="925" value="36.3">36°18'N</geoCoordinate>
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,
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<geoCoordinate degrees="52" direction="east" minutes="21" orientation="longitude" precision="925" value="52.35">52°21'E</geoCoordinate>
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, 18.07.1973 (A. Senglet); 3♂ 6♀ (MHNG), Babol,
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<geoCoordinate degrees="36" direction="north" minutes="33" orientation="latitude" precision="925" value="36.55">36°33'N</geoCoordinate>
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,
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<geoCoordinate degrees="52" direction="east" minutes="42" orientation="longitude" precision="925" value="52.7">52°42'E</geoCoordinate>
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, 19.07.1973 (A. Senglet); 5♀ (MHNG), Gilan Province: Rudbar,
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<geoCoordinate degrees="36" direction="north" minutes="49" orientation="latitude" precision="925" value="36.816666">36°49'N</geoCoordinate>
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,
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<geoCoordinate degrees="49" direction="east" minutes="25" orientation="longitude" precision="925" value="49.416668">49°25'E</geoCoordinate>
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, 4.09.1973 (A. Senglet). Russia: Daghestan: 1♂ (PPC), Sergokalinski Dist., Sergokala Vil., 31.07.2008 (A.V. Alieva); 1♀ (PPC), Makhachkala, 08.2009 (S.V. Alieva); 1♀ (PPC), same locality, 08.2008 (S.V. Alieva); 3♀ (PPC), Magaramkentski Dist., Tselegyun Vil., 8.08.2008 (S.V. Alieva); 1♀ (PPC), Kizilyurtovski Dist., Sultan-Yangiyurt Vil., 18.05.2009 (M.A. Aliev, Z.A. Shavlukov); 1♀ (PPC), Karabudakhkentsky Dist., 07.2008. (N.M. Gasanova). Georgia: 1♀ (photographed specimen), Tbilisi, 41.767986N, 44.767779E, 17.09.2019 (A. Seropian). India: 1♀ (MMUE), Himachal Pradesh State: Patlikuhl Town,
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<geoCoordinate degrees="32" direction="north" minutes="07" orientation="latitude" precision="925" value="32.116665">32°07'N</geoCoordinate>
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,
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<geoCoordinate degrees="77" direction="east" minutes="08" orientation="longitude" precision="925" value="77.13333">77°08'E</geoCoordinate>
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, 1200 m, 28-29.5.1999 (Y.M. Marusik); 4♂ 2♀ (MMUE), Punjab State: Patiala, University campus,
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<geoCoordinate degrees="30" direction="north" minutes="21" orientation="latitude" precision="925" value="30.35">30°21'N</geoCoordinate>
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,
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<geoCoordinate degrees="76" direction="east" minutes="27" orientation="longitude" precision="925" value="76.45">76°27'E</geoCoordinate>
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, 24-25.6.1999 (Y.M. Marusik); 4♂ 1♀ (MMUE) and 5♀ (MMUE), same data.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="diagnosis">
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<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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<taxonomicName authorityName="*" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona theisi" order="Araneae" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="theisi">
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<emphasis italics="true" pageId="0" pageNumber="13">Neoscona theisi</emphasis>
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</taxonomicName>
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differs from the congeners occurring in the region by the presence of a wide black median band on the venter of abdomen and thin white lateral stripes (Fig.
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitus of Neoscona spasskyi (A, B) and N. theisi (C, D). A-C Dorsal D ventral C, D showing variations in comparison to specimens depicted in Figure 11. Photos C, D courtesy of A. Seropian." figureDoi="10.3897/zookeys.906.47978.figure12" httpUri="https://binary.pensoft.net/fig/374414" pageId="0" pageNumber="13">12D</figureCitation>
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). Males of this species have tibia II with fewer ventral spines (ca. 20) than
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<taxonomicName lsidName="N. spasskyi" order="Aranei" pageId="0" pageNumber="13" rank="species" species="spasskyi">
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<emphasis italics="true" pageId="0" pageNumber="13">N. spasskyi</emphasis>
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</taxonomicName>
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(ca. 40) and
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<taxonomicName lsidName="N. isatis" order="Aranei" pageId="0" pageNumber="13" rank="species" species="isatis">
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<emphasis italics="true" pageId="0" pageNumber="13">N. isatis</emphasis>
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</taxonomicName>
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sp. nov. (ca. 90) and more than in
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<taxonomicName lsidName="N. adianta" order="Aranei" pageId="0" pageNumber="13" rank="species" species="adianta">
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<emphasis italics="true" pageId="0" pageNumber="13">N. adianta</emphasis>
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</taxonomicName>
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(ca. 10). Males of
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<taxonomicName lsidName="N. theisi" order="Aranei" pageId="0" pageNumber="13" rank="species" species="theisi">
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<emphasis italics="true" pageId="0" pageNumber="13">N. theisi</emphasis>
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</taxonomicName>
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can be recognized also by the palp with pointed dorsal extension/projection of the tibia (Fig.
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Male palps and tibiae II of Neoscona theisi (A, D), N. isatis sp. nov. (B, F) and N. spasskyi (C, E). A-C Male palp, prolateral D-F male tibia II, ventral. Abbreviations: Co conductor, La lamella, Ma median apophysis, Me extension of median apophysis, Ta terminal apophysis, Te tegulum. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="10.3897/zookeys.906.47978.figure14" httpUri="https://binary.pensoft.net/fig/374416" pageId="0" pageNumber="13">14A</figureCitation>
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) (vs. absent), distinct ventral conical projection of the tegulum (
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<emphasis italics="true" pageId="0" pageNumber="13">Te</emphasis>
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) lacking in other species, broad and well sclerotized lamella and wide conductor (vs. lamella thin and weakly sclerotized, conductor club-like), and long prolateral extension of median apophysis, longer than spur of median apophysis (vs. extension absent or as long as spur). The epigyne of
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<taxonomicName lsidName="N. theisi" order="Aranei" pageId="0" pageNumber="13" rank="species" species="theisi">
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<emphasis italics="true" pageId="0" pageNumber="13">N. theisi</emphasis>
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</taxonomicName>
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differs from those of
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<taxonomicName lsidName="N. isatis" order="Aranei" pageId="0" pageNumber="13" rank="species" species="isatis">
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<emphasis italics="true" pageId="0" pageNumber="13">N. isatis</emphasis>
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</taxonomicName>
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sp. nov. and
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<taxonomicName lsidName="N. spasskyi" order="Aranei" pageId="0" pageNumber="13" rank="species" species="spasskyi">
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<emphasis italics="true" pageId="0" pageNumber="13">N. spasskyi</emphasis>
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</taxonomicName>
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by the lack of constriction. Females of
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<taxonomicName lsidName="N. theisi" order="Aranei" pageId="0" pageNumber="13" rank="species" species="theisi">
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<emphasis italics="true" pageId="0" pageNumber="13">N. theisi</emphasis>
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</taxonomicName>
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well differ from those of
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<taxonomicName lsidName="N. adianta" order="Aranei" pageId="0" pageNumber="13" rank="species" species="adianta">
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<emphasis italics="true" pageId="0" pageNumber="13">N. adianta</emphasis>
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</taxonomicName>
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by having a white median band on carapace, darker abdominal pattern and the epigyne being almost twice longer than wide (vs. white band absent, epigyne almost as wide as long).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="description">
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<paragraph pageId="0" pageNumber="13">Description.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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See
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<bibRefCitation author="Grasshoff, M" journalOrPublisher="Annalen Zoologische Wetenschappen" pageId="0" pageNumber="13" pagination="1 - 123" refId="B10" refString="Grasshoff, M, 1986. Die Radnetzspinnen-Gattung Neoscona in Afrika (Arachnida: Araneae). Annalen Zoologische Wetenschappen 250: 1 - 123" title="Die Radnetzspinnen-Gattung Neoscona in Afrika (Arachnida: Araneae)." volume="250" year="1986">Grasshoff (1986)</bibRefCitation>
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and
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<bibRefCitation DOI="https://doi.org/10.2476/asjaa.47.133" author="Tanikawa, A" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="13" pagination="133 - 168" refId="B22" refString="Tanikawa, A, 1998. A revision of the Japanese spiders of the genus Neoscona (Araneae: Araneidae). Acta Arachnologica 47: 133 - 168, DOI: https://doi.org/10.2476/asjaa.47.133" title="A revision of the Japanese spiders of the genus Neoscona (Araneae: Araneidae)." url="https://doi.org/10.2476/asjaa.47.133" volume="47" year="1998">Tanikawa (1998)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="comments">
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<paragraph pageId="0" pageNumber="13">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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<taxonomicName authorityName="*" baseAuthorityName="Walckenaer" baseAuthorityYear="1841" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona theisi" order="Araneae" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="theisi">
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<emphasis italics="true" pageId="0" pageNumber="13">Neoscona theisi</emphasis>
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</taxonomicName>
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is a widely distributed species, with a current natural range covering Pakistan to Japan.
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<bibRefCitation author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="13" pagination="311 - 365" refId="B11" refString="Levy, G, 1998. Twelve genera of orb-weaver spiders (Araneae, Araneidae) from Israel. Israel Journal of Zoology 43: 311 - 365" title="Twelve genera of orb-weaver spiders (Araneae, Araneidae) from Israel." volume="43" year="1998">Levy (1998)</bibRefCitation>
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described
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<taxonomicName lsidName="N. sodom" order="Aranei" pageId="0" pageNumber="13" rank="species" species="sodom">
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<emphasis italics="true" pageId="0" pageNumber="13">N. sodom</emphasis>
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</taxonomicName>
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on the basis of both sexes from Israel. Judging by the figures provided in the original description, there are no significant differences in the copulatory organs and habitus of
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<taxonomicName lsidName="N. sodom" order="Aranei" pageId="0" pageNumber="13" rank="species" species="sodom">
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<emphasis italics="true" pageId="0" pageNumber="13">N. sodom</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="N. theisi" order="Aranei" pageId="0" pageNumber="13" rank="species" species="theisi">
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<emphasis italics="true" pageId="0" pageNumber="13">N. theisi</emphasis>
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</taxonomicName>
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. Therefore, the former name is synonymized with the latter.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="distribution">
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<paragraph pageId="0" pageNumber="13">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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Pakistan, India, Philippines, China to Indonesia, Japan. Introduced to Seychelles, Pacific Is. (
|
|
<bibRefCitation author="WSC" journalOrPublisher="Sjoestedts Kilimandjaro-Meru Expedition" pageId="0" pageNumber="13" publicationUrl="http://wsc.nmbe.ch" refId="B25" refString="WSC, 2019. World Spider Catalog. Version 20.5. Natural History Museum Bern. http://wsc.nmbe.ch" title="World Spider Catalog. Version 20.5. Natural History Museum Bern." url="http://wsc.nmbe.ch" year="2019">WSC 2019</bibRefCitation>
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|
). The westernmost localities of this species (sub
|
|
<taxonomicName lsidName="N. sodom" order="Aranei" pageId="0" pageNumber="13" rank="species" species="sodom">
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|
<emphasis italics="true" pageId="0" pageNumber="13">N. sodom</emphasis>
|
|
</taxonomicName>
|
|
) are Cyprus (
|
|
<bibRefCitation author="Bosmans, R" journalOrPublisher="Zoologica Scripta" pageId="0" pageNumber="13" refId="B8" refString="Bosmans, R, Van Keer, J, Russell-Smith, A, Hadjiconstantis, M, Komnenov, M, Bosselaers, J, Huber, S, Mccowan, D, Snazell, R, Decae, A, Zoumides, C, Kielhorn, K-H, Oger, P, 2019. Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Arachnological Contributions. Newsletter of the Belgian Arachnological Society, 34(suppl.): 1-197." title="Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Arachnological Contributions. Newsletter of the Belgian Arachnological Society, 34 (suppl.): 1 - 197." year="2019">Bosmans et al. 2019</bibRefCitation>
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|
) and Israel (
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<bibRefCitation author="Levy, G" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="13" pagination="311 - 365" refId="B11" refString="Levy, G, 1998. Twelve genera of orb-weaver spiders (Araneae, Araneidae) from Israel. Israel Journal of Zoology 43: 311 - 365" title="Twelve genera of orb-weaver spiders (Araneae, Araneidae) from Israel." volume="43" year="1998">Levy 1998</bibRefCitation>
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). New records for Iran, Georgia, and Russia.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |