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<document ID-DOI="10.11646/zootaxa.5270.2.8" ID-GBIF-Dataset="f2385569-01e1-40b0-8bc2-7f4d6b4f4272" ID-ISSN="1175-5326" ID-Zenodo-Dep="7865018" ID-ZooBank="02AE384D-AE40-4DA9-813B-66D3453FDF50" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1681979585876" checkinUser="plazi" docAuthor="Li, Bing, Zhao, Zhe &amp; Li, Shuqiang" docDate="2023" docId="DA0CB62D4B294D239AFA3FFB49D9262C" docLanguage="en" docName="zootaxa.5270.2.8.pdf" docOrigin="Zootaxa 5270 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5270.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vappolotes tianjiayu Li &amp; Zhao &amp; Li 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="332" masterDocId="2635CE554B2C4D249A6D3B354A462749" masterDocTitle="Three new species of the genus Vappolotes Zhao &amp; Li, 2019 (Araneae, Agelenidae) from southwest China" masterLastPageNumber="336" masterPageNumber="325" pageNumber="330" updateTime="1682453639316" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Three new species of the genus Vappolotes Zhao &amp; Li, 2019 (Araneae, Agelenidae) from southwest China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Bing</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7106-4680</mods:nameIdentifier>
<mods:affiliation>College of Life Science, Langfang Normal University, Langfang 065000, China &amp; libing @ lfnu. edu. cn; https: // orcid. org / 0000 - 0002 - 7106 - 4680</mods:affiliation>
<mods:nameIdentifier type="email">libing@lfnu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Zhe</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0781-0204</mods:nameIdentifier>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; zhaozhe @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 0781 - 0204</mods:affiliation>
<mods:nameIdentifier type="email">zhaozhe@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Shuqiang</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5270</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:end>336</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5270.2.8</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5270.2.8</mods:identifier>
<mods:identifier type="GBIF-Dataset">f2385569-01e1-40b0-8bc2-7f4d6b4f4272</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">7865018</mods:identifier>
<mods:identifier type="ZooBank">02AE384D-AE40-4DA9-813B-66D3453FDF50</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7865158" ID-Zenodo-Dep="7865158" LSID="urn:lsid:plazi:treatment:DA0CB62D4B294D239AFA3FFB49D9262C" httpUri="http://treatment.plazi.org/id/DA0CB62D4B294D239AFA3FFB49D9262C" lastPageId="7" lastPageNumber="332" pageId="5" pageNumber="330">
<subSubSection box="[151,470,1230,1257]" pageId="5" pageNumber="330" type="nomenclature">
<paragraph blockId="5.[151,1058,1230,1329]" box="[151,470,1230,1257]" pageId="5" pageNumber="330">
<heading bold="true" box="[151,470,1230,1257]" fontSize="11" level="1" pageId="5" pageNumber="330" reason="1">
<emphasis bold="true" box="[151,470,1230,1257]" pageId="5" pageNumber="330">
<taxonomicName authority="Li &amp; Zhao &amp; Li, 2023" authorityName="Li &amp; Zhao &amp; Li" authorityYear="2023" box="[151,397,1230,1257]" class="Arachnida" family="Agelenidae" genus="Vappolotes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="330" phylum="Arthropoda" rank="species" species="tianjiayu" status="sp. n.">
<emphasis bold="true" box="[151,397,1230,1257]" italics="true" pageId="5" pageNumber="330">Vappolotes tianjiayu</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[404,470,1231,1257]" pageId="5" pageNumber="330" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="330" type="description">
<paragraph blockId="5.[151,1058,1230,1329]" box="[151,248,1267,1293]" pageId="5" pageNumber="330">
<figureCitation box="[151,222,1267,1293]" captionStart="FIGURE 4" captionStartId="6.[152,255,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-31@6.[216,1372,181,1880]" captionTargetPageId="6" captionText="FIGURE 4. Epigyne and habitus of Vappolotes tianjiayu sp. n., holotype. A epigyne, ventral view B vulva, dorsal view C female habitus, dorsal view D same, ventral view E same, lateral view. Scale bars: equal for A and B, equal for C, D, and E. Abbreviations:A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head." figureDoi="http://doi.org/10.5281/zenodo.7865032" httpUri="https://zenodo.org/record/7865032/files/figure.png" pageId="5" pageNumber="330">Figs 4</figureCitation>
,
<figureCitation box="[234,248,1267,1293]" captionStart="FIGURE 7" captionStartId="9.[152,255,1953,1977]" captionTargetBox="[151,1435,957,1930]" captionTargetId="figure-393@9.[151,1435,957,1930]" captionTargetPageId="9" captionText="FIGURE 7. Type localities of new (orange) and previously described (blue) species of Vappolotes from China. 1 V. ganlongensis 2 V. jianpingensis 3 V. longshan sp. n., 4 V. tianjiayu sp. n., 5 V. hei sp. n." figureDoi="http://doi.org/10.5281/zenodo.7865042" httpUri="https://zenodo.org/record/7865042/files/figure.png" pageId="5" pageNumber="330">7</figureCitation>
</paragraph>
<paragraph blockId="5.[151,1058,1230,1329]" box="[151,1058,1302,1329]" pageId="5" pageNumber="330">
LSIDurn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="330" type="materials_examined">
<paragraph blockId="5.[151,1437,1374,2013]" pageId="5" pageNumber="330">
<emphasis bold="true" box="[151,332,1374,1401]" pageId="5" pageNumber="330">Type material.</emphasis>
<materialsCitation collectingDate="2016-04-14" collectorName="Y. L. Li &amp; Z. G. Chen" country="China" county="Shimen Co." elevation="448" latitude="29.9065" location="Hupingshan Town" longLatPrecision="6" longitude="110.8087" municipality="Changde City" pageId="5" pageNumber="330" specimenCount="1" specimenCount-female="1" stateProvince="Hunan" typeStatus="holotype">
<emphasis bold="true" box="[346,464,1375,1401]" pageId="5" pageNumber="330">
<typeStatus box="[346,459,1375,1401]" pageId="5" pageNumber="330">Holotype</typeStatus>
:
</emphasis>
<specimenCount box="[479,500,1375,1401]" pageId="5" pageNumber="330" type="female"></specimenCount>
(IZCAS-Ar44376, XQ270):
<collectingCountry box="[848,919,1375,1401]" name="China" pageId="5" pageNumber="330">China</collectingCountry>
:
<collectingRegion box="[936,1088,1375,1401]" country="China" name="Hunan" pageId="5" pageNumber="330">Hunan Prov.</collectingRegion>
:
<collectingMunicipality box="[1105,1272,1375,1401]" pageId="5" pageNumber="330">Changde City</collectingMunicipality>
:
<collectingCounty box="[1289,1432,1375,1401]" pageId="5" pageNumber="330">Shimen Co.</collectingCounty>
:
<location LSID="urn:lsid:plazi:treatment:DA0CB62D4B294D239AFA3FFB49D9262C:577A51E04B294D219AFA3EB64B2022D4" box="[151,358,1411,1437]" country="China" county="Shimen Co." latitude="29.9065" longLatPrecision="6" longitude="110.8087" municipality="Changde City" name="Hupingshan Town" pageId="5" pageNumber="330" stateProvince="Hunan">Hupingshan Town</location>
, Tianjiayu Vil., an unnamed cave, (
<geoCoordinate box="[745,872,1410,1437]" degrees="29.9065" direction="north" orientation="latitude" pageId="5" pageNumber="330" precision="5" value="29.9065">N29.9065°</geoCoordinate>
,
<geoCoordinate box="[882,1017,1411,1437]" degrees="110.8087" direction="east" orientation="longitude" pageId="5" pageNumber="330" precision="5" value="110.8087">E110.8087°</geoCoordinate>
,
<quantity box="[1027,1100,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="4.4799999999999995" pageId="5" pageNumber="330" unit="m" value="448.0">
<elevation box="[1027,1100,1411,1437]" metricMagnitude="2" metricUnit="m" metricValue="4.4799999999999995" pageId="5" pageNumber="330" unit="m" value="448.0">448 m</elevation>
</quantity>
),
<date box="[1119,1245,1411,1437]" pageId="5" pageNumber="330" value="2016-04-14">
<collectingDate box="[1119,1245,1411,1437]" pageId="5" pageNumber="330" value="2016-04-14">14.IV.2016</collectingDate>
</date>
,
<collectorName box="[1254,1333,1411,1437]" pageId="5" pageNumber="330">Y.L. Li</collectorName>
and
<collectorName pageId="5" pageNumber="330">Z.G. Chen</collectorName>
leg.
</materialsCitation>
<materialsCitation collectingDate="2016-04-12" country="China" elevation="442" latitude="29.9869" location="Qingshanxi Vil." longLatPrecision="6" longitude="110.863" pageId="5" pageNumber="330" specimenCount="1" specimenCount-female="1" stateProvince="Hunan" typeStatus="paratype">
<emphasis bold="true" box="[267,396,1447,1473]" pageId="5" pageNumber="330">
<typeStatus box="[267,391,1447,1473]" pageId="5" pageNumber="330">Paratypes</typeStatus>
:
</emphasis>
<specimenCount box="[404,439,1447,1473]" pageId="5" pageNumber="330" type="female">1♀</specimenCount>
(IZCAS-Ar44382, XQ263): same town as holotype,
<location LSID="urn:lsid:plazi:treatment:DA0CB62D4B294D239AFA3FFB49D9262C:577A51E04B294D219E7C3E924E832288" box="[1041,1221,1447,1473]" country="China" latitude="29.9869" longLatPrecision="6" longitude="110.863" name="Qingshanxi Vil." pageId="5" pageNumber="330" stateProvince="Hunan">Qingshanxi Vil.</location>
, an unnamed cave, (
<geoCoordinate box="[159,286,1483,1509]" degrees="29.9869" direction="north" orientation="latitude" pageId="5" pageNumber="330" precision="5" value="29.9869">N29.9869°</geoCoordinate>
,
<geoCoordinate box="[297,433,1483,1509]" degrees="110.8630" direction="east" orientation="longitude" pageId="5" pageNumber="330" precision="5" value="110.863">E110.8630°</geoCoordinate>
,
<quantity box="[444,518,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="5" pageNumber="330" unit="m" value="442.0">
<elevation box="[444,518,1483,1509]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="5" pageNumber="330" unit="m" value="442.0">442 m</elevation>
</quantity>
),
<date box="[539,665,1483,1509]" pageId="5" pageNumber="330" value="2016-04-12">
<collectingDate box="[539,665,1483,1509]" pageId="5" pageNumber="330" value="2016-04-12">12.IV.2016</collectingDate>
</date>
, same leg
</materialsCitation>
.;
<materialsCitation collectingDate="2016-04-14" collectorName="Y. L. Li &amp; Z. G. Chen" country="China" county="Shimen Co." elevation="448" latitude="29.9065" location="Hupingshan Town" longLatPrecision="6" longitude="110.8087" municipality="Changde City" pageId="5" pageNumber="330" specimenCount="5" specimenCount-female="5" stateProvince="Hunan" typeStatus="paratype">
<specimenCount box="[798,854,1482,1509]" pageId="5" pageNumber="330" type="female">5♀♀</specimenCount>
(IZCAS-Ar44377Ar44381, XQ270), same data as holotype
</materialsCitation>
;
<specimenCount box="[265,300,1519,1545]" pageId="5" pageNumber="330" type="female">1♀</specimenCount>
(IZCAS-Ar44383, XQ256): same vil. as holotype, another unnamed cave, (
<geoCoordinate box="[1158,1285,1519,1545]" degrees="29.9066" direction="north" orientation="latitude" pageId="5" pageNumber="330" precision="5" value="29.9066">N29.9066°</geoCoordinate>
,
<geoCoordinate box="[1297,1432,1519,1545]" degrees="110.8088" direction="east" orientation="longitude" pageId="5" pageNumber="330" precision="5" value="110.8088">E110.8088°</geoCoordinate>
,
<quantity box="[151,225,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="4.55" pageId="5" pageNumber="330" unit="m" value="455.0">455 m</quantity>
),
<date box="[244,371,1555,1581]" pageId="5" pageNumber="330" value="2016-04-14">14.IV.2016</date>
, same leg.;
<materialsCitation collectingDate="2016-04-14" country="China" elevation="395" latitude="29.9362" location="Guluoping Vil." longLatPrecision="6" longitude="110.8574" municipality="Guluo Cave" pageId="5" pageNumber="330" specimenCount="3" specimenCount-female="3" stateProvince="Hunan" typeStatus="paratype">
<specimenCount box="[501,556,1555,1581]" pageId="5" pageNumber="330" type="female">3♀♀</specimenCount>
(IZCAS-Ar44384Ar44386, XQ245): same town as holotype,
<location LSID="urn:lsid:plazi:treatment:DA0CB62D4B294D239AFA3FFB49D9262C:577A51E04B294D219E833D264FDE2164" box="[1262,1432,1555,1581]" country="China" latitude="29.9362" longLatPrecision="6" longitude="110.8574" municipality="Guluo Cave" name="Guluoping Vil." pageId="5" pageNumber="330" stateProvince="Hunan">Guluoping Vil.</location>
,
<collectingMunicipality box="[151,288,1591,1617]" pageId="5" pageNumber="330">Guluo Cave</collectingMunicipality>
, (
<geoCoordinate box="[308,435,1591,1617]" degrees="29.9362" direction="north" orientation="latitude" pageId="5" pageNumber="330" precision="5" value="29.9362">N29.9362°</geoCoordinate>
,
<geoCoordinate box="[445,581,1590,1617]" degrees="110.8574" direction="east" orientation="longitude" pageId="5" pageNumber="330" precision="5" value="110.8574">E110.8574°</geoCoordinate>
,
<quantity box="[592,666,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="3.95" pageId="5" pageNumber="330" unit="m" value="395.0">
<elevation box="[592,666,1590,1617]" metricMagnitude="2" metricUnit="m" metricValue="3.95" pageId="5" pageNumber="330" unit="m" value="395.0">395 m</elevation>
</quantity>
),
<date box="[686,812,1591,1617]" pageId="5" pageNumber="330" value="2016-04-14">
<collectingDate box="[686,812,1591,1617]" pageId="5" pageNumber="330" value="2016-04-14">14.IV.2016</collectingDate>
</date>
, same leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,1110,1627,1653]" pageId="5" pageNumber="330" type="etymology">
<paragraph blockId="5.[151,1437,1374,2013]" box="[199,1110,1627,1653]" pageId="5" pageNumber="330">
<emphasis bold="true" box="[199,334,1627,1653]" pageId="5" pageNumber="330">Etymology.</emphasis>
The new species is named after the
<typeStatus box="[740,788,1627,1653]" pageId="5" pageNumber="330">type</typeStatus>
locality; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="330" type="diagnosis">
<paragraph blockId="5.[151,1437,1374,2013]" pageId="5" pageNumber="330">
<emphasis bold="true" box="[199,322,1662,1688]" pageId="5" pageNumber="330">Diagnosis.</emphasis>
The female of
<taxonomicName authority="Li &amp; Zhao &amp; Li, 2023" authorityName="Li &amp; Zhao &amp; Li" authorityYear="2023" box="[486,715,1663,1689]" class="Arachnida" family="Agelenidae" genus="Vappolotes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="330" phylum="Arthropoda" rank="species" species="tianjiayu" status="sp. n.">
<emphasis box="[486,715,1663,1689]" italics="true" pageId="5" pageNumber="330">Vappolotes tianjiayu</emphasis>
</taxonomicName>
<emphasis bold="true" box="[720,782,1663,1689]" pageId="5" pageNumber="330">
<taxonomicNameLabel box="[720,782,1663,1689]" pageId="5" pageNumber="330" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from other species of the genus by the small copulatory ducts and the large spermathecae (
<figureCitation box="[728,816,1699,1725]" captionStart="FIGURE 4" captionStartId="6.[152,255,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-31@6.[216,1372,181,1880]" captionTargetPageId="6" captionText="FIGURE 4. Epigyne and habitus of Vappolotes tianjiayu sp. n., holotype. A epigyne, ventral view B vulva, dorsal view C female habitus, dorsal view D same, ventral view E same, lateral view. Scale bars: equal for A and B, equal for C, D, and E. Abbreviations:A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head." figureDoi="http://doi.org/10.5281/zenodo.7865032" httpUri="https://zenodo.org/record/7865032/files/figure.png" pageId="5" pageNumber="330">Fig. 4B</figureCitation>
). The length of copulatory ducts far less than the length of spermathecae (
<figureCitation box="[349,437,1735,1761]" captionStart="FIGURE 4" captionStartId="6.[152,255,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-31@6.[216,1372,181,1880]" captionTargetPageId="6" captionText="FIGURE 4. Epigyne and habitus of Vappolotes tianjiayu sp. n., holotype. A epigyne, ventral view B vulva, dorsal view C female habitus, dorsal view D same, ventral view E same, lateral view. Scale bars: equal for A and B, equal for C, D, and E. Abbreviations:A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head." figureDoi="http://doi.org/10.5281/zenodo.7865032" httpUri="https://zenodo.org/record/7865032/files/figure.png" pageId="5" pageNumber="330">Fig. 4B</figureCitation>
), but longer than 1.5 times in others (
<figureCitation box="[856,945,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[152,255,1702,1726]" captionTargetBox="[151,1435,306,1674]" captionTargetId="figure-91@2.[151,1435,306,1676]" captionTargetPageId="2" captionText="FIGURE 1. Male palp and epigyne of Vappolotes ganlongensis Zhao &amp; Li, 2019 A male palp, prolateral view B same, ventral view C same, retrolateral view D epigyne, ventral view E vulva, dorsal view. Scale bar: equal for A, B, and C, equal for D and E. Abbreviations: A = atrium; AC = anterior conductor; CD = copulatory duct; CF = cymbial furrow; CO = copulatory opening; DC = dorsal conductor; E = embolus; EB = embolic base; FD = fertilization duct; H = hood; LC = lamella of conductor; LTA = lateral tibial apophysis; PA = patellar apophysis; PC = posterior conductor; PMA = posterior margin of atrium; RTA = retrolateral tibial apophysis; S = spermatheca; ST = subtegulum; T = tegulum; TS = tegulum sclerite." figureDoi="http://doi.org/10.5281/zenodo.7865024" httpUri="https://zenodo.org/record/7865024/files/figure.png" pageId="5" pageNumber="330">Figs 1E</figureCitation>
,
<figureCitation box="[958,993,1735,1761]" captionStart="FIGURE 3" captionStartId="4.[152,255,1894,1918]" captionTargetBox="[225,1368,181,1870]" captionTargetId="figure-31@4.[219,1368,181,1870]" captionTargetPageId="4" captionText="FIGURE 3. Epigyne and habitus of Vappolotes longshan sp. n. A epigyne, ventral view B vulva, dorsal view C male habitus, dorsal view D female habitus, dorsal view E same, ventral view. Scale bars: equal for A and B, equal for D and E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca." figureDoi="http://doi.org/10.5281/zenodo.7865030" httpUri="https://zenodo.org/record/7865030/files/figure.png" pageId="5" pageNumber="330">3B</figureCitation>
,
<figureCitation box="[1004,1041,1735,1761]" captionStart="FIGURE 6" captionStartId="8.[152,255,1894,1918]" captionTargetBox="[219,1368,181,1870]" captionTargetId="figure-31@8.[219,1368,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Epigyne and habitus of Vappolotes hei sp. n. A epigyne, ventral view B vulva, dorsal view C male habitus, dorsal view D female habitus, dorsal view E same, ventral view. Scale bars: equal for A and B, equal for D and E. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head." figureDoi="http://doi.org/10.5281/zenodo.7865040" httpUri="https://zenodo.org/record/7865040/files/figure.png" pageId="5" pageNumber="330">6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="332" pageId="5" pageNumber="330" type="description">
<paragraph blockId="5.[151,1437,1374,2013]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="332" pageId="5" pageNumber="330">
<emphasis bold="true" box="[199,580,1770,1797]" pageId="5" pageNumber="330">
Description. Female (
<typeStatus box="[457,567,1770,1796]" pageId="5" pageNumber="330">holotype</typeStatus>
):
</emphasis>
Abdomen light brownish gray, four chevrons present, rest as for the genus (
<figureCitation box="[159,285,1807,1833]" captionStart="FIGURE 4" captionStartId="6.[152,255,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-31@6.[216,1372,181,1880]" captionTargetPageId="6" captionText="FIGURE 4. Epigyne and habitus of Vappolotes tianjiayu sp. n., holotype. A epigyne, ventral view B vulva, dorsal view C female habitus, dorsal view D same, ventral view E same, lateral view. Scale bars: equal for A and B, equal for C, D, and E. Abbreviations:A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head." figureDoi="http://doi.org/10.5281/zenodo.7865032" httpUri="https://zenodo.org/record/7865032/files/figure.png" pageId="5" pageNumber="330">Fig. 4CE</figureCitation>
). Total length 7.16. Carapace 3.08 long, 2.04 wide. Abdomen 4.08 long, 2.81 wide. Eye sizes and inter-distances: AME 0.06, ALE 0.11, PME 0.09, PLE 0.10; AMEAME 0.03, AMEALE 0.04, AMEPME 0.03, ALEPLE 0.01, PMEPME 0.04, PMEPLE 0.06. Leg measurements: leg I 9.23 (2.51, 0.79, 2.39, 1.80, 1.74); II 8.59 (2.33, 0.73, 2.03, 1.86, 1.64); III 8.09 (2.49, 0.78, 1.86, 1.94, 1.02); IV 11.62 (3.13, 0.84, 2.97, 2.92, 1.76). Epigyne as in
<figureCitation box="[314,431,1951,1977]" captionStart="FIGURE 4" captionStartId="6.[152,255,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-31@6.[216,1372,181,1880]" captionTargetPageId="6" captionText="FIGURE 4. Epigyne and habitus of Vappolotes tianjiayu sp. n., holotype. A epigyne, ventral view B vulva, dorsal view C female habitus, dorsal view D same, ventral view E same, lateral view. Scale bars: equal for A and B, equal for C, D, and E. Abbreviations:A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head." figureDoi="http://doi.org/10.5281/zenodo.7865032" httpUri="https://zenodo.org/record/7865032/files/figure.png" pageId="5" pageNumber="330">Fig 4A, B</figureCitation>
: ca. 1.4 times wider than long; atrium large, occupying ca. 3/5 of epigynal field, as long as wide; its posterior margin narrowed in the middle, ca. 1/7 the length of the atrium, at least 1.4 times longer than the width of its lateral margin; copulatory ducts cumuliform, located posteriorly, occupying less than one tenth of spermathecae, covered by spermathecae in dorsal view; spermathecae enlarged, occupying 1/2 of epigynal field; fertilization ducts thin, ca. 5 times longer than wide.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7865032" ID-Zenodo-Dep="7865032" httpUri="https://zenodo.org/record/7865032/files/figure.png" pageId="6" pageNumber="331" startId="6.[152,255,1904,1928]" targetBox="[216,1372,181,1880]" targetPageId="6" targetType="figure">
<paragraph blockId="6.[151,1436,1904,2036]" pageId="6" pageNumber="331">
<emphasis bold="true" box="[152,282,1904,1928]" pageId="6" pageNumber="331">FIGURE 4.</emphasis>
Epigyne and habitus of
<taxonomicName authority="Li &amp; Zhao &amp; Li, 2023" authorityName="Li &amp; Zhao &amp; Li" authorityYear="2023" box="[540,750,1904,1928]" class="Arachnida" family="Agelenidae" genus="Vappolotes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="331" phylum="Arthropoda" rank="species" species="tianjiayu" status="sp. n.">
<emphasis box="[540,750,1904,1928]" italics="true" pageId="6" pageNumber="331">Vappolotes tianjiayu</emphasis>
</taxonomicName>
<emphasis bold="true" box="[758,817,1904,1928]" pageId="6" pageNumber="331">
<taxonomicNameLabel box="[758,817,1904,1928]" pageId="6" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype.
<emphasis bold="true" box="[935,953,1904,1928]" pageId="6" pageNumber="331">A</emphasis>
epigyne, ventral view
<emphasis bold="true" box="[1194,1211,1904,1928]" pageId="6" pageNumber="331">B</emphasis>
vulva, dorsal view
<emphasis bold="true" box="[1418,1436,1904,1928]" pageId="6" pageNumber="331">C</emphasis>
female habitus, dorsal view
<emphasis bold="true" box="[440,458,1940,1964]" pageId="6" pageNumber="331">D</emphasis>
same, ventral view
<emphasis bold="true" box="[666,683,1940,1964]" pageId="6" pageNumber="331">E</emphasis>
same, lateral view. Scale bars: equal for
<emphasis bold="true" box="[1106,1124,1940,1964]" pageId="6" pageNumber="331">A</emphasis>
and
<emphasis bold="true" box="[1175,1192,1940,1964]" pageId="6" pageNumber="331">B</emphasis>
, equal for
<emphasis bold="true" box="[1305,1323,1940,1964]" pageId="6" pageNumber="331">C</emphasis>
,
<emphasis bold="true" box="[1337,1355,1940,1964]" pageId="6" pageNumber="331">D</emphasis>
, and
<emphasis bold="true" box="[1413,1430,1940,1964]" pageId="6" pageNumber="331">E</emphasis>
. Abbreviations: A = atrium; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; H = hood; PMA = posterior margin of atrium; S = spermatheca; SH = spermathecal head.
</paragraph>
</caption>
<paragraph blockId="7.[151,1436,151,357]" box="[199,392,259,285]" pageId="7" pageNumber="332">
<emphasis bold="true" box="[199,267,259,285]" pageId="7" pageNumber="332">Male.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="7.[151,1436,151,357]" box="[199,800,294,321]" pageId="7" pageNumber="332">
<emphasis bold="true" box="[199,319,294,320]" pageId="7" pageNumber="332">Variation.</emphasis>
Total length of females: 6.527.16 (n=11).
</paragraph>
</subSubSection>
<subSubSection box="[199,927,330,357]" pageId="7" pageNumber="332" type="distribution">
<paragraph blockId="7.[151,1436,151,357]" box="[199,927,330,357]" pageId="7" pageNumber="332">
<emphasis bold="true" box="[199,353,330,356]" pageId="7" pageNumber="332">Distribution.</emphasis>
Known only from
<collectingRegion box="[567,751,331,357]" country="China" name="Hunan" pageId="7" pageNumber="332">Hunan Province</collectingRegion>
,
<collectingCountry box="[762,829,331,357]" name="China" pageId="7" pageNumber="332">China</collectingCountry>
(
<figureCitation box="[844,913,331,357]" captionStart="FIGURE 7" captionStartId="9.[152,255,1953,1977]" captionTargetBox="[151,1435,957,1930]" captionTargetId="figure-393@9.[151,1435,957,1930]" captionTargetPageId="9" captionText="FIGURE 7. Type localities of new (orange) and previously described (blue) species of Vappolotes from China. 1 V. ganlongensis 2 V. jianpingensis 3 V. longshan sp. n., 4 V. tianjiayu sp. n., 5 V. hei sp. n." figureDoi="http://doi.org/10.5281/zenodo.7865042" httpUri="https://zenodo.org/record/7865042/files/figure.png" pageId="7" pageNumber="332">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>