162 lines
16 KiB
XML
162 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.288.4095" ID-GBIF-Dataset="43b6b64a-b86b-4c0b-afcc-229ccecb6e16" ID-PMC="PMC3690914" ID-Pensoft-Pub="1313-2970-288-1" ID-PubMed="23798897" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-288-1" ModsDocOrigin="ZooKeys 288" ModsDocTitle="Generic revision and species classification of the Microdontinae (Diptera, Syrphidae)" checkinTime="1451247431953" checkinUser="pensoft" docAuthor="Reemer, Menno & Stahls, Gunilla" docDate="2013" docId="C9158C9B964142B07BC1866A43B4F390" docLanguage="en" docName="ZooKeys 288: 1-213" docOrigin="ZooKeys 288" docSource="http://dx.doi.org/10.3897/zookeys.288.4095" docTitle="Ceratrichomyia Seguy, stat. n." docType="treatment" docVersion="3" lastPageNumber="24" masterDocId="582E2572FFD0FFB3884E8864FFF1A15A" masterDocTitle="Generic revision and species classification of the Microdontinae (Diptera, Syrphidae)" masterLastPageNumber="213" masterPageNumber="1" pageNumber="22" updateTime="1668155636847" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Generic revision and species classification of the Microdontinae (Diptera, Syrphidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Reemer, Menno</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Stahls, Gunilla</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2013</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>288</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>213</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.288.4095</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.288.4095</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-288-1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="152042627" LSID="urn:lsid:plazi:treatment:C9158C9B964142B07BC1866A43B4F390" httpUri="http://treatment.plazi.org/id/C9158C9B964142B07BC1866A43B4F390" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
|
||
<subSubSection pageId="21" pageNumber="22" type="nomenclature">
|
||
<paragraph pageId="21" pageNumber="22">
|
||
<taxonomicName LSID="http://species-id.net/wiki/Ceratrichomyia" authority="Seguy" class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
Ceratrichomyia
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="21" pageNumber="22">stat. n.</taxonomicNameLabel>
|
||
Figs 46-58
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="21" pageNumber="22" type="reference_group">
|
||
<paragraph pageId="21" pageNumber="22">
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
, 1951: 14. Type species:
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia behara" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="behara">Ceratrichomyia behara</taxonomicName>
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
, 1951: 14, by original designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
|
||
<paragraph pageId="21" pageNumber="22">Description.</paragraph>
|
||
<paragraph lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
Body length: 7-10 mm. Slender, black flies with yellow markings and a constricted abdomen. Head wider than thorax, face and vertex wider than an eye. Face ventrally produced in profile; wider than an eye. Lateral oral margins not produced. Vertex swollen. Occiput narrow ventrally, strongly widened dorsally. Eye
|
||
<pageBreakToken pageId="22" pageNumber="23" start="start">bare</pageBreakToken>
|
||
. Eyes in male not approaching each other; smallest mutual distance much larger than width of antennal fossa. Antennal fossa about as high as wide. Antenna longer than height of head. Basoflagellomere at least three times as long as scape; with long pilosity. Postpronotum pilose or bare. Mesoscutum with transverse suture complete. Scutellum without calcars. Anepisternum with deep sulcus; entirely pilose. Anepimeron entirely pilose. Katepimeron convex; pilose or bare. Wing: vein R4+5 with posterior appendix; vein M1 straight, perpendicular to R4+5 and M; postero-apical corner of cell r4+5 rectangular, with small appendix; crossvein r-m located around basal 1/4 of cell dm. Abdomen constricted at segment 2. Tergites 3 and 4 not fused, able to articulate independently. Sternite 1 bare. Sternite 4 in male covered by genital capsule, therefore not visible without removing genitalia. Male genitalia: phallus straight or slightly bent dorsad, with spherical base very large, at least as long as remaining part of phallus; phallus furcate near apex; epandrium with or without ventrolateral ridge; surstylus deeply furcate.
|
||
</paragraph>
|
||
<paragraph pageId="22" pageNumber="23">
|
||
Diagnosis. The combination of a complete transverse suture on the mesoscutum and a constricted abdomen is only found in
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Indascia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indascia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Indascia</taxonomicName>
|
||
Keiser, Indascia, 1958,
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Thompsodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thompsodon" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Thompsodon</taxonomicName>
|
||
gen. n. and certain species of
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Paramixogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramixogaster" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Paramixogaster</taxonomicName>
|
||
Brunetti, 1923. Males are easily distinguished from all these taxa by the long pilosity of the basoflagellomere, and also by sternite 4, which is covered by the genital capsule. From
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Paramixogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramixogaster" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Paramixogaster</taxonomicName>
|
||
this genus also differs by the unfused tergites 3 and 4. Females are unknown.
|
||
</paragraph>
|
||
<paragraph pageId="22" pageNumber="23">
|
||
Discussion.
|
||
<bibRefCitation author="Seguy, E" journalOrPublisher="Revue Francaise d'Entomologie" pageId="85" pageNumber="86" pagination="14 - 18" title="Trois nouveaux syrphides de Madagascar (Dipt.)." volume="18" year="1951">
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
(1951)
|
||
</bibRefCitation>
|
||
attributed one species to this genus. He designated a male and a female as
|
||
<normalizedToken originalValue="‘types’">'types'</normalizedToken>
|
||
, and another male as
|
||
<normalizedToken originalValue="‘cotype’">'cotype'</normalizedToken>
|
||
. These are here all considered as syntypes. Examination of these three specimens made clear that they belong to three different species, which makes it necessary to designate a lectotype. The male with the following label data is here designated as lectotype. Label 1: "Madagascar, Behara"; label 2 (blue): "Museum Paris, III-38, A. Seyrig"; label 3 (red):
|
||
<normalizedToken originalValue="“Type”">"Type"</normalizedToken>
|
||
; label 4: "
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia behara" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="behara">Ceratrichomyia behara</taxonomicName>
|
||
type du genre [male symbol]
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
50"; coll. MNHN. A redescription of the lecotype is given in the next section of the present paper. By this lectotype designation, the other two syntypes become paralectotypes. The male collected in Bekily (Madagascar) belongs to a new species of
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
, which is described in the present paper as
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia bullabucca" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bullabucca">Ceratrichomyia bullabucca</taxonomicName>
|
||
spec. n. The female paralectotype, collected in Bekily, is here considered to belong to a previously undescribed species of
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Paramixogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramixogaster" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Paramixogaster</taxonomicName>
|
||
, because it possesses all characters described as diagnostic for that genus (see genus account). A description of that species is given under the name
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Paramixogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramixogaster piptotus" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="piptotus">Paramixogaster piptotus</taxonomicName>
|
||
sp. n. A third species attributed to this genus,
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia angolensis" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="angolensis">Ceratrichomyia angolensis</taxonomicName>
|
||
sp. n., is described from Angola.
|
||
</paragraph>
|
||
<paragraph lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
The long pilosity of the male basoflagellomere was used by
|
||
<bibRefCitation author="Seguy, E" journalOrPublisher="Revue Francaise d'Entomologie" pageId="85" pageNumber="86" pagination="14 - 18" title="Trois nouveaux syrphides de Madagascar (Dipt.)." volume="18" year="1951">
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
(1951)
|
||
</bibRefCitation>
|
||
as a character to set his African genus
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
apart from other
|
||
<taxonomicName lsidName="" pageId="22" pageNumber="23" rank="subFamily" subFamily="Microdontinae">Microdontinae</taxonomicName>
|
||
. This character is also present in
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ptilobactrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptilobactrum" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Ptilobactrum</taxonomicName>
|
||
Bezzi, 1915, another African taxon. Apparently
|
||
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
|
||
was not aware of this, as he did not refer to
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ptilobactrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptilobactrum" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">Ptilobactrum</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Cheng, X-Y" journalOrPublisher="Zootaxa" pageId="82" pageNumber="83" pagination="21 - 48" title="A generic conspectus of the Microdontinae (Diptera: Syrphidae) with the description of two new genera from Africa and China." volume="1879" year="2008">Cheng and Thompson (2008)</bibRefCitation>
|
||
did notice the similarity in antennal structure in both taxa and,
|
||
<pageBreakToken pageId="23" pageNumber="24" start="start">based</pageBreakToken>
|
||
on the descriptions, proposed to regard
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
as a subjective junior synonym of
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ptilobactrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptilobactrum" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ptilobactrum</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="23" pageNumber="24">
|
||
Study of the type specimens of
|
||
<taxonomicName genus="Ceratricomyia" lsidName="Ceratricomyia" pageId="23" pageNumber="24" rank="genus">Ceratricomyia</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ptilobactrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptilobactrum" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ptilobactrum</taxonomicName>
|
||
revealed that these taxa are in fact very different. While
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
has, for instance, a constricted abdomen with unfused tergites 3 and 4,
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ptilobactrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptilobactrum" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ptilobactrum</taxonomicName>
|
||
has a conical abdomen with fused tergites 3 and 4. The structures of the male genitalia are also very different (compare Figs 56-58 with 326), e.g. with a deeply furcate surstylus in
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
and an unfurcate one in
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ptilobactrum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptilobactrum" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ptilobactrum</taxonomicName>
|
||
. Considering these morphological differences, and supported by the phylogenetic results of
|
||
<bibRefCitation pageId="23" pageNumber="24">
|
||
Reemer and
|
||
<normalizedToken originalValue="Ståhls">Stahls</normalizedToken>
|
||
(in press)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName class="Insecta" family="Syrphidae" genus="Ceratrichomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratrichomyia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Ceratrichomyia</taxonomicName>
|
||
is here re-instated as a valid genus.
|
||
</paragraph>
|
||
<paragraph pageId="23" pageNumber="24">Diversity and distribution. Described species: 3. Two species are known from Madagascar, one from the African mainland (Angola).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |