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<document id="52C8F0D5D519706502D64248EE9561EA" ID-DOI="10.11646/zootaxa.3884.2.7" ID-GBIF-Dataset="877ae302-0eb1-44c9-b7d3-a277a5292ef7" ID-ISSN="1175-5326" ID-Zenodo-Dep="225533" ID-ZooBank="B769B761-CE47-48F9-8AAD-E15EB1973BA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460966694343" checkinUser="plazi" docAuthor="Kazerani, Farzaneh, Khaghaninia, Samad, Talebi, Ali Asghar &amp; Shamshev, Igor" docDate="2014" docId="A50C3C40FFC0AF57FF0AFE0AFBEFFD16" docLanguage="en" docName="zt03884p193.pdf" docOrigin="Zootaxa 3884 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empis (Lissempis) guilanensis Kazerani &amp; Shamshev, sp. nov." docType="treatment" docVersion="9" lastPageNumber="190" masterDocId="59354438FFC2AF52FF9DFFD0FFD1FFBE" masterDocTitle="New data on Empis Linnaeus (Diptera: Empididae) from Iran, with description of a new species of the subgenus Lissempis Bezzi" masterLastPageNumber="193" masterPageNumber="185" pageNumber="187" updateTime="1698557877113" updateUser="plazi">
<mods:mods id="743A0F9D0A82FFBD0BEBB3DD592CC5EE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4FDA267B9BEAB70885D8E74178E3E56D">
<mods:title id="D86985A931ABCBB811054F42C6875DB2">New data on Empis Linnaeus (Diptera: Empididae) from Iran, with description of a new species of the subgenus Lissempis Bezzi</mods:title>
</mods:titleInfo>
<mods:name id="06A902A319DDCE13C8C6BB701E0C01D9" type="personal">
<mods:role id="7D3791A9B8EA23F67AC4EA27A4E15C6A">
<mods:roleTerm id="B45A28EED4F7301C52BD2F0678B6E3EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE4BC68EBBAF88E28793371C63D14281">Kazerani, Farzaneh</mods:namePart>
</mods:name>
<mods:name id="978AB4ADF7164DB76E061A409D4940ED" type="personal">
<mods:role id="8239716163779EA5D49697684CC4DD73">
<mods:roleTerm id="1E9AF24AFD8475B6B35F8FEB228CD787">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E2655FAAD49A60A09F29404D620814F">Khaghaninia, Samad</mods:namePart>
</mods:name>
<mods:name id="71E3778CA79290F13C7D990EC691F846" type="personal">
<mods:role id="BFD84316BCC1356460C3C0354943FE94">
<mods:roleTerm id="3EE41BC411589A90B2B96AB56CFC8EF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="533F1ED949AC3A7CD7C5B6C81847ED3C">Talebi, Ali Asghar</mods:namePart>
</mods:name>
<mods:name id="E96890DA2775FDD6AEF9E6E3739665B4" type="personal">
<mods:role id="986C4CA8F5408E7F2BBC22F380AC2618">
<mods:roleTerm id="87959E42E0F242711FCB79A19A3B9AC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F0A8D7693E1A4F3E88005E057C4836E">Shamshev, Igor</mods:namePart>
</mods:name>
<mods:typeOfResource id="4DF93ED0EC33B058AAB4A33EFB16824A">text</mods:typeOfResource>
<mods:relatedItem id="9C37BB67B8811EAD27CCA9D21E0441D6" type="host">
<mods:titleInfo id="50852277BBFDD4193DAEC0A3D0366C79">
<mods:title id="6FE001DE0E8A49CCD2B6F3F6B61B9A63">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BD8354E6218D0CC145E3E2D8037209EE">
<mods:date id="EE8E72F34F65D7BB7AE105D35CE56F07">2014</mods:date>
<mods:detail id="F7740A0BE0CB8C354096C6A7C5A9667E" type="volume">
<mods:number id="14A01B41E092523220625F64BF302243">3884</mods:number>
</mods:detail>
<mods:detail id="32F1D17BEB86731F9B3E3D62AA4AB3EE" type="issue">
<mods:number id="8805F524675518407957F2CC4A53F317">2</mods:number>
</mods:detail>
<mods:extent id="7D72B1BB30D1F15217DF97D2043D5D60" unit="page">
<mods:start id="8D5D5C54DEF2EC569033DAB9149F7634">185</mods:start>
<mods:end id="5214EC722D6393BB2A89E44881161C90">193</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D599E2E5FE835C54CDC0A464FA9A0577">journal article</mods:classification>
<mods:identifier id="214A6C7AAA2028CF051710CA3FA009C8" type="DOI">10.11646/zootaxa.3884.2.7</mods:identifier>
<mods:identifier id="2907A8B1FB618C9FC3D35E37CE193EA6" type="GBIF-Dataset">877ae302-0eb1-44c9-b7d3-a277a5292ef7</mods:identifier>
<mods:identifier id="708B1ED655A0E485461E4275DEBEDFA5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="34D8961AF51E2F384DB2DC1AB426EA6F" type="Zenodo-Dep">225533</mods:identifier>
<mods:identifier id="76ECB79225DEACCD800AF2F1419DEB94" type="ZooBank">B769B761-CE47-48F9-8AAD-E15EB1973BA5</mods:identifier>
</mods:mods>
<treatment id="A50C3C40FFC0AF57FF0AFE0AFBEFFD16" ID-DOI="http://doi.org/10.5281/zenodo.6141610" ID-GBIF-Taxon="119593387" ID-Zenodo-Dep="6141610" LSID="urn:lsid:plazi:treatment:A50C3C40FFC0AF57FF0AFE0AFBEFFD16" httpUri="http://treatment.plazi.org/id/A50C3C40FFC0AF57FF0AFE0AFBEFFD16" lastPageId="5" lastPageNumber="190" pageId="2" pageNumber="187">
<subSubSection id="65BFDEDDFFC0AF50FF0AFE0AFEDAFDA8" pageId="2" pageNumber="187" type="nomenclature">
<paragraph id="2D1A8D56FFC0AF50FF0AFE0AFC43FE4A" blockId="2.[151,914,474,534]" box="[151,914,474,500]" pageId="2" pageNumber="187">
<heading id="76523A3AFFC0AF50FF0AFE0AFC43FE4A" bold="true" box="[151,914,474,500]" fontSize="11" level="1" pageId="2" pageNumber="187" reason="1">
<emphasis id="1FD15144FFC0AF50FF0AFE0AFC43FE4A" bold="true" box="[151,914,474,500]" pageId="2" pageNumber="187">
<taxonomicName id="EAA5F6D5FFC0AF50FF0AFE0AFCF9FE4A" authority="Kazerani &amp; Shamshev" authorityName="Kazerani &amp; Shamshev" box="[151,808,474,500]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="guilanensis" status="sp. nov." subGenus="Lissempis">
<emphasis id="1FD15144FFC0AF50FF0AFE0AFF35FE4A" bold="true" box="[151,228,474,500]" italics="true" pageId="2" pageNumber="187">Empis</emphasis>
(
<emphasis id="1FD15144FFC0AF50FF69FE0AFEBBFE4A" bold="true" box="[244,362,474,500]" italics="true" pageId="2" pageNumber="187">Lissempis</emphasis>
)
<emphasis id="1FD15144FFC0AF50FEE1FE0AFDD7FE4A" bold="true" box="[380,518,474,500]" italics="true" pageId="2" pageNumber="187">guilanensis</emphasis>
Kazerani &amp; Shamshev
</taxonomicName>
,
<taxonomicNameLabel id="04E2EC3FFFC0AF50FCA8FE0AFC43FE4A" box="[821,914,474,500]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2D1A8D56FFC0AF50FF0AFE2DFEDAFDA8" blockId="2.[151,914,474,534]" box="[151,267,509,534]" pageId="2" pageNumber="187">
(
<figureCitation id="B59E91D3FFC0AF50FF02FE2DFED3FDA8" box="[159,258,509,534]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 6" captionStart-2="FIGURE 7" captionStartId-0="2.[151,263,1907,1930]" captionStartId-1="3.[151,263,1786,1809]" captionStartId-2="4.[151,249,1325,1347]" captionTargetBox-0="[340,1246,839,1884]" captionTargetBox-1="[340,1240,208,1765]" captionTargetBox-2="[344,1237,630,1300]" captionTargetId-0="figure@2.[340,1247,838,1886]" captionTargetId-1="figure@3.[340,1247,193,1765]" captionTargetId-2="figure@4.[340,1247,625,1303]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." captionText-1="FIGURES 5 – 6. Empis (Lissempis) guilanensis sp. nov. 5. antenna; 6. male genitalia (lateral view) (scale: 0.5 mm)." captionText-2="FIGURE 7. Empis (Lissempis) guilanensis sp. nov., male genitalia, lateral view. Abbreviations: cerc—cercus; epandepandrium; hypd— hypandrium; phal—phallus (scale: 0.5 mm)." httpUri-0="https://zenodo.org/record/225534/files/figure.png" httpUri-1="https://zenodo.org/record/225535/files/figure.png" httpUri-2="https://zenodo.org/record/225536/files/figure.png" pageId="2" pageNumber="187">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="65BFDEDDFFC0AF50FF0AFD95FC1AFCAC" pageId="2" pageNumber="187" type="materials_examined">
<paragraph id="2D1A8D56FFC0AF50FF0AFD95FB40FD3C" blockId="2.[151,1437,581,786]" pageId="2" pageNumber="187">
<emphasis id="1FD15144FFC0AF50FF0AFD95FEECFDE0" bold="true" box="[151,317,581,606]" pageId="2" pageNumber="187">
<typeStatus id="F21E33F4FFC0AF50FF0AFD95FF03FDE0" box="[151,210,581,606]" pageId="2" pageNumber="187">Type</typeStatus>
material
</emphasis>
.
<emphasis id="1FD15144FFC0AF50FED6FD95FE38FDE0" bold="true" box="[331,489,581,606]" pageId="2" pageNumber="187">
<typeStatus id="F21E33F4FFC0AF50FED6FD95FE38FDE0" box="[331,489,581,606]" pageId="2" pageNumber="187" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, ♂ &quot;
<collectingCountry id="55B2CDC6FFC0AF50FD82FD95FDB7FDE0" box="[543,614,581,606]" name="Iran" pageId="2" pageNumber="187">
<emphasis id="1FD15144FFC0AF50FD82FD95FDB7FDE0" bold="true" box="[543,614,581,606]" pageId="2" pageNumber="187">IRAN</emphasis>
</collectingCountry>
, Guilan province, Rudsar, Orkom, N 36˚ 45.739',
<geoCoordinate id="4891EB91FFC0AF50FB05FD96FAE3FDE0" box="[1176,1330,581,606]" direction="east" orientation="longitude" pageId="2" pageNumber="187" precision="1" value="0.8383883">E 0 50 18.198</geoCoordinate>
',
<quantity id="EA5D20B3FFC0AF50FADDFD96FA46FDE3" box="[1344,1431,582,606]" metricMagnitude="3" metricUnit="m" metricValue="1.201" pageId="2" pageNumber="187" unit="m" value="1201.0">1201 m</quantity>
,
<date id="591BAB96FFC0AF50FF0AFDBAFEDDFD3C" box="[151,268,618,642]" pageId="2" pageNumber="187" value="2010-05-31">31.V.2010</date>
, M. Khayrandish&quot; (
<collectionCode id="4BB41593FFC0AF50FE75FDBAFDEEFD3C" box="[488,575,618,642]" pageId="2" pageNumber="187">TMUC</collectionCode>
).
<emphasis id="1FD15144FFC0AF50FDCCFDB9FD34FD3C" bold="true" box="[593,741,617,642]" pageId="2" pageNumber="187">
<typeStatus id="F21E33F4FFC0AF50FDCCFDB9FD34FD3C" box="[593,741,617,642]" pageId="2" pageNumber="187" type="paratype">PARATYPE</typeStatus>
</emphasis>
: ♂, same data as
<typeStatus id="F21E33F4FFC0AF50FC31FDB9FBDFFD3C" box="[940,1038,617,642]" pageId="2" pageNumber="187" type="holotype">holotype</typeStatus>
(
<collectionCode id="4BB41593FFC0AF50FB80FDBAFB57FD3C" box="[1053,1158,618,642]" pageId="2" pageNumber="187">ICHMM</collectionCode>
).
</paragraph>
<paragraph id="2D1A8D56FFC0AF50FF5AFD5DFC1AFCAC" blockId="2.[151,1437,581,786]" pageId="2" pageNumber="187">
<emphasis id="1FD15144FFC0AF50FF5AFD5DFE87FD18" bold="true" box="[199,342,653,678]" pageId="2" pageNumber="187">Recognition</emphasis>
. Mid-sized species with black, pale setose thorax and partly yellow abdomen (
<figureCitation id="B59E91D3FFC0AF50FAA2FD5DFA5BFD18" box="[1343,1418,653,678]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="2" pageNumber="187">Fig. 1</figureCitation>
); postpronotal lobe, scutum and scutellum shining; scape of antenna elongate, nearly
<date id="591BAB96FFC0AF50FB0AFD62FB6BFD74" box="[1175,1210,690,714]" pageId="2" pageNumber="187">3X</date>
longer than wide; postpedicel about
<date id="591BAB96FFC0AF50FEF7FD05FE5DFD50" box="[362,396,725,750]" pageId="2" pageNumber="187">6X</date>
longer than wide (
<figureCitation id="B59E91D3FFC0AF50FDF0FD05FD65FD50" box="[621,692,725,750]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,263,1786,1809]" captionTargetBox="[340,1240,208,1765]" captionTargetId="figure@3.[340,1247,193,1765]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Empis (Lissempis) guilanensis sp. nov. 5. antenna; 6. male genitalia (lateral view) (scale: 0.5 mm)." httpUri="https://zenodo.org/record/225535/files/figure.png" pageId="2" pageNumber="187">Fig. 6</figureCitation>
); legs largely yellow, covered with unmodified setae, hind tibia thickened towards apex (
<figureCitation id="B59E91D3FFC0AF50FE32FD29FE22FCAC" box="[431,499,761,786]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="2" pageNumber="187">Fig. 4</figureCitation>
), hind basitarsus slightly swollen (
<figureCitation id="B59E91D3FFC0AF50FCE5FD29FC6CFCAC" box="[888,957,761,786]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="2" pageNumber="187">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="79DADDDEFFC0AF50FF0AF8A3FF30F816" httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="2" pageNumber="187" targetBox="[340,1246,839,1884]" targetPageId="2">
<paragraph id="2D1A8D56FFC0AF50FF0AF8A3FF30F816" blockId="2.[151,1437,1907,1960]" pageId="2" pageNumber="187">
<emphasis id="1FD15144FFC0AF50FF0AF8A3FEE2F836" bold="true" box="[151,307,1907,1930]" pageId="2" pageNumber="187">FIGURES 1–4</emphasis>
.
<taxonomicName id="EAA5F6D5FFC0AF50FEA2F8A4FDA2F836" box="[319,627,1907,1929]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="guilanensis" status="sp. nov." subGenus="Lissempis">
<emphasis id="1FD15144FFC0AF50FEA2F8A4FEAEF837" box="[319,383,1908,1929]" italics="true" pageId="2" pageNumber="187">Empis</emphasis>
(
<emphasis id="1FD15144FFC0AF50FE10F8A4FE20F837" box="[397,497,1908,1929]" italics="true" pageId="2" pageNumber="187">Lissempis</emphasis>
)
<emphasis id="1FD15144FFC0AF50FE62F8A3FDA2F836" box="[511,627,1907,1928]" italics="true" pageId="2" pageNumber="187">guilanensis</emphasis>
</taxonomicName>
<emphasis id="1FD15144FFC0AF50FDE5F8A4FD0AF836" bold="true" box="[632,731,1907,1929]" pageId="2" pageNumber="187">
<taxonomicNameLabel id="04E2EC3FFFC0AF50FDE5F8A4FD18F837" box="[632,713,1908,1929]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
1
</emphasis>
. male habitus (lateral view);
<emphasis id="1FD15144FFC0AF50FB9EF8A3FBDEF836" bold="true" box="[1027,1039,1907,1928]" pageId="2" pageNumber="187">2</emphasis>
. fore leg;
<emphasis id="1FD15144FFC0AF50FBEBF8A3FB52F836" bold="true" box="[1142,1155,1907,1928]" pageId="2" pageNumber="187">3</emphasis>
. mid leg;
<emphasis id="1FD15144FFC0AF50FB74F8A3FB24F836" bold="true" box="[1257,1269,1907,1928]" pageId="2" pageNumber="187">4</emphasis>
. hind leg (scale: 1 mm).
</paragraph>
</caption>
<caption id="79DADDDEFFC1AF51FF0AF92AFAF9F8AF" box="[151,1320,1786,1809]" httpUri="https://zenodo.org/record/225535/files/figure.png" pageId="3" pageNumber="188" targetBox="[340,1240,208,1765]" targetPageId="3">
<paragraph id="2D1A8D56FFC1AF51FF0AF92AFAF9F8AF" blockId="3.[151,1320,1786,1809]" box="[151,1320,1786,1809]" pageId="3" pageNumber="188">
<emphasis id="1FD15144FFC1AF51FF0AF92AFEE2F8AE" bold="true" box="[151,307,1786,1809]" pageId="3" pageNumber="188">FIGURES 5–6</emphasis>
.
<taxonomicName id="EAA5F6D5FFC1AF51FEDDF92CFDA5F8AE" box="[320,628,1787,1809]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="guilanensis" status="sp. nov." subGenus="Lissempis">
<emphasis id="1FD15144FFC1AF51FEDDF92CFE51F8AF" box="[320,384,1788,1809]" italics="true" pageId="3" pageNumber="188">Empis</emphasis>
(
<emphasis id="1FD15144FFC1AF51FE13F92CFE23F8AF" box="[398,498,1788,1809]" italics="true" pageId="3" pageNumber="188">Lissempis</emphasis>
)
<emphasis id="1FD15144FFC1AF51FD9DF92BFDA5F8AE" box="[512,628,1787,1808]" italics="true" pageId="3" pageNumber="188">guilanensis</emphasis>
</taxonomicName>
<emphasis id="1FD15144FFC1AF51FDE7F92BFD0CF8AE" bold="true" box="[634,733,1787,1808]" pageId="3" pageNumber="188">
<taxonomicNameLabel id="04E2EC3FFFC1AF51FDE7F92BFD1BF8AE" box="[634,714,1787,1808]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
5
</emphasis>
. antenna;
<emphasis id="1FD15144FFC1AF51FCDAF92BFC82F8AE" bold="true" box="[839,851,1787,1808]" pageId="3" pageNumber="188">6</emphasis>
. male genitalia (lateral view) (scale: 0.5 mm).
</paragraph>
</caption>
<subSubSection id="65BFDEDDFFC1AF57FF5AF8E8FE73FEDB" lastPageId="5" lastPageNumber="190" pageId="3" pageNumber="188" type="description">
<paragraph id="2D1A8D56FFC1AF56FF5AF8E8FCEAFEA2" blockId="3.[151,1437,1848,2017]" lastBlockId="4.[151,1437,151,573]" lastPageId="4" lastPageNumber="189" pageId="3" pageNumber="188">
<emphasis id="1FD15144FFC1AF51FF5AF8E8FE80F8EF" bold="true" box="[199,337,1848,1873]" pageId="3" pageNumber="188">Description</emphasis>
.
<emphasis id="1FD15144FFC1AF51FEC2F8E8FE4DF8EF" bold="true" box="[351,412,1848,1873]" pageId="3" pageNumber="188">Male</emphasis>
(
<figureCitation id="B59E91D3FFC1AF51FE36F8E8FDC0F8EF" box="[427,529,1848,1873]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 6" captionStart-2="FIGURE 7" captionStartId-0="2.[151,263,1907,1930]" captionStartId-1="3.[151,263,1786,1809]" captionStartId-2="4.[151,249,1325,1347]" captionTargetBox-0="[340,1246,839,1884]" captionTargetBox-1="[340,1240,208,1765]" captionTargetBox-2="[344,1237,630,1300]" captionTargetId-0="figure@2.[340,1247,838,1886]" captionTargetId-1="figure@3.[340,1247,193,1765]" captionTargetId-2="figure@4.[340,1247,625,1303]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." captionText-1="FIGURES 5 – 6. Empis (Lissempis) guilanensis sp. nov. 5. antenna; 6. male genitalia (lateral view) (scale: 0.5 mm)." captionText-2="FIGURE 7. Empis (Lissempis) guilanensis sp. nov., male genitalia, lateral view. Abbreviations: cerc—cercus; epandepandrium; hypd— hypandrium; phal—phallus (scale: 0.5 mm)." httpUri-0="https://zenodo.org/record/225534/files/figure.png" httpUri-1="https://zenodo.org/record/225535/files/figure.png" httpUri-2="https://zenodo.org/record/225536/files/figure.png" pageId="3" pageNumber="188">Figs 1–7</figureCitation>
). Body length
<quantity id="EA5D20B3FFC1AF51FD2BF8E8FCF9F8EE" box="[694,808,1848,1873]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="3" pageNumber="188" unit="mm" value="5.15" valueMax="5.3" valueMin="5.0">5–5.3 mm</quantity>
(in
<typeStatus id="F21E33F4FFC1AF51FCC8F8E8FC66F8EF" box="[853,951,1848,1873]" pageId="3" pageNumber="188" type="holotype">holotype</typeStatus>
5.3); wing length
<quantity id="EA5D20B3FFC1AF51FB1BF8E9FADCF8EE" box="[1158,1293,1849,1873]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="3" pageNumber="188" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 mm</quantity>
(in
<typeStatus id="F21E33F4FFC1AF51FAA7F8E8FA4DF8EF" box="[1338,1436,1848,1873]" pageId="3" pageNumber="188" type="holotype">holotype</typeStatus>
4.2). Head largely black in ground-colour; vertex, ocellar tubercle, frons and face subshining, faintly greyish pollinose, occiput light brown. Eyes dichoptic. Frons narrow, above antennae narrower than anterior ocellus; with several thin yellow setae. Face very narrow, below antennae nearly as broad as frons. Ocellar tubercle with 2 long, thin brownish yellow and several minute setae. Occiput covered with numerous thin, pale setae. Antenna dark brown; scape with scattered short dark setae, nearly
<date id="591BAB96FFC6AF56FD7AFF48FCD8FF0E" box="[743,777,152,176]" pageId="4" pageNumber="189">3X</date>
longer than wide, about 2.5X longer than pedicel; pedicel with short dark setae; postpedicel very long, 5.8–6.0X longer than wide near base, with straight margins; stylus short (
<figureCitation id="B59E91D3FFC6AF56FF42FF0FFEF5FF46" box="[223,292,223,248]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,263,1786,1809]" captionTargetBox="[340,1240,208,1765]" captionTargetId="figure@3.[340,1247,193,1765]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Empis (Lissempis) guilanensis sp. nov. 5. antenna; 6. male genitalia (lateral view) (scale: 0.5 mm)." httpUri="https://zenodo.org/record/225535/files/figure.png" pageId="4" pageNumber="189">Fig. 5</figureCitation>
). Proboscis yellow, 2 times longer than head height; labrum yellow, about 1.5 times longer than head height; palpus yellowish brown, with scattered yellow setae.
</paragraph>
<paragraph id="2D1A8D56FFC6AF56FF5AFEF7FE6BFD82" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="189">Thorax black in ground-colour, only postalar callus yellow, with pale to yellow setae; postpronotal lobe, scutum (except notopleuron) and scutellum shining; remaining prothoracic sclerites and entire mesopleuron faintly pollinose. Prosternum and proepisternum bare. Antepronotum and postpronotal lobe with several short, thin setae. Mesonotum with 2 notopleurals (anterior seta distinctly longer, as long as apical scutellars), 2 short postalar, 6 scutellars (apical and median pairs long, lateral pair short); additionally, notopleuron with several hair-like setae; acrostichals uniserial, short, rather scattered, also present on prescutellar depression; dorsocentrals uniserial, nearly as long as acrostichals, prescutellar pair long (nearly as long as lateral scutellar bristles). Laterotergite with numerous long pale setae.</paragraph>
<caption id="79DADDDEFFC6AF56FF0AFAFDFCF5FADC" httpUri="https://zenodo.org/record/225536/files/figure.png" pageId="4" pageNumber="189" targetBox="[344,1237,630,1300]" targetPageId="4">
<paragraph id="2D1A8D56FFC6AF56FF0AFAFDFCF5FADC" blockId="4.[151,1436,1325,1378]" pageId="4" pageNumber="189">
<emphasis id="1FD15144FFC6AF56FF0AFAFDFEC3FAFC" bold="true" box="[151,274,1325,1347]" pageId="4" pageNumber="189">FIGURE 7</emphasis>
.
<taxonomicName id="EAA5F6D5FFC6AF56FEB9FAFEFDB2FAFC" box="[292,611,1325,1347]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="guilanensis" status="sp. nov." subGenus="Lissempis">
<emphasis id="1FD15144FFC6AF56FEB9FAFEFEB2FAFD" box="[292,355,1326,1347]" italics="true" pageId="4" pageNumber="189">Empis</emphasis>
(
<emphasis id="1FD15144FFC6AF56FEE5FAFEFE0DFAFD" box="[376,476,1326,1347]" italics="true" pageId="4" pageNumber="189">Lissempis</emphasis>
)
<emphasis id="1FD15144FFC6AF56FE6DFAFDFDB2FAFC" box="[496,611,1325,1346]" italics="true" pageId="4" pageNumber="189">guilanensis</emphasis>
</taxonomicName>
<emphasis id="1FD15144FFC6AF56FDF2FAFDFD14FAFC" bold="true" box="[623,709,1325,1346]" pageId="4" pageNumber="189">
<taxonomicNameLabel id="04E2EC3FFFC6AF56FDF2FAFDFD14FAFC" box="[623,709,1325,1346]" pageId="4" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male genitalia, lateral view. Abbreviations: cerc—cercus; epandepandrium; hypd— hypandrium; phal—phallus (scale: 0.5 mm).
</paragraph>
</caption>
<paragraph id="2D1A8D56FFC6AF56FF5AFA5FFAE7F88D" blockId="4.[151,1437,1422,2026]" pageId="4" pageNumber="189">
Legs with coxae, fore and mid femora and tibiae entirely yellow; hind femur brown on about apical 1/4, otherwise yellow (
<figureCitation id="B59E91D3FFC6AF56FEF8FA62FE75FA75" box="[357,420,1458,1483]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig 4</figureCitation>
); hind tibia brownish yellow to yellow near base, otherwise brownish (
<figureCitation id="B59E91D3FFC6AF56FB20FA62FAD3FA75" box="[1213,1282,1458,1483]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 4</figureCitation>
); tarsi brown, except yellow basal 2/3 of fore and mid basitarsus (
<figureCitation id="B59E91D3FFC6AF56FD48FA06FCEDFA51" box="[725,828,1494,1519]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Figs 2–4</figureCitation>
). Coxae and trochanters with ordinary pale setae. Fore femur slender, with rows of pale (closer to base) moderately long setae (
<figureCitation id="B59E91D3FFC6AF56FC41FA2AFBF2F9AD" box="[988,1059,1530,1555]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 2</figureCitation>
). Fore tibia lacking strong setae, denser pubescent closer to apex, with 2 short, black, spine-like subapical setae ventrally (
<figureCitation id="B59E91D3FFC6AF56FB24F9CEFAD3F989" box="[1209,1282,1566,1591]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 2</figureCitation>
). Fore tarsus unmodified; clothed in short, ordinary setae (
<figureCitation id="B59E91D3FFC6AF56FD0FF992FD06F9E5" box="[658,727,1602,1627]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 2</figureCitation>
). Mid femur slightly narrower than fore femur; with short, thin ventral pale and brown setae and 1 subapical back seta (
<figureCitation id="B59E91D3FFC6AF56FC89F9B6FC88F9C1" box="[788,857,1638,1663]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 3</figureCitation>
). Mid tibia with 2 black, spine-like subapical setae anteriorly, otherwise clothed in short ordinary setae (
<figureCitation id="B59E91D3FFC6AF56FD7AF95AFCFAF91D" box="[743,811,1674,1699]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 3</figureCitation>
). Mid tarsomeres unmodified.
<collectingCountry id="55B2CDC6FFC6AF56FB1BF95AFB6FF91D" box="[1158,1214,1674,1699]" name="India" pageId="4" pageNumber="189">Hind</collectingCountry>
femur narrowed on basal part; with dense silvery white pubescent ventrally, with 1 strong, black anteroventral subapical seta and similar subapical seta anteriorly (
<figureCitation id="B59E91D3FFC6AF56FD8BF902FD8CF955" box="[534,605,1746,1771]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 3</figureCitation>
).
<collectingCountry id="55B2CDC6FFC6AF56FDE8F902FD7CF955" box="[629,685,1746,1771]" name="India" pageId="4" pageNumber="189">Hind</collectingCountry>
tibia evenly thickened towards apex; pubescent ventrally, with 1 black dorsal bristle near base and some scattered, short setae closer to apex dorsally (
<figureCitation id="B59E91D3FFC6AF56FB14F926FB03F8B1" box="[1161,1234,1782,1807]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 4</figureCitation>
).
<collectingCountry id="55B2CDC6FFC6AF56FB76F926FAF2F8B1" box="[1259,1315,1782,1807]" name="India" pageId="4" pageNumber="189">Hind</collectingCountry>
basitarsus slightly uniformly thickened but narrower than hind tibia at apex; covered with unmodified setae (
<figureCitation id="B59E91D3FFC6AF56FB7EF8CAFAF9F88D" box="[1251,1320,1818,1843]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="2D1A8D56FFC6AF56FF5AF8EEFAFCF81F" blockId="4.[151,1437,1422,2026]" pageId="4" pageNumber="189">
Wing hyaline, anal lobe weakly developed, anal angle obtuse; 1 rather short, thin costal seta; stigma distinct, brownish yellow. Veins R5 and M1 slightly divergent near wing margin; radial fork rather broad; vein CuA+CuP incomplete; discal median cell elongate apically. Calypter yellow, with pale setae. Halter yellow (
<figureCitation id="B59E91D3FFC6AF56FB46F859FACDF81F" box="[1243,1308,1929,1954]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="2D1A8D56FFC6AF57FF5AF87DFDECFE81" blockId="4.[151,1437,1422,2026]" lastBlockId="5.[151,1437,151,680]" lastPageId="5" lastPageNumber="190" pageId="4" pageNumber="189">
Abdomen largely brownish to yellowish brown, tergites 1–4 yellowish laterally and narrowly along posterior margin, sternites 1–4 entirely pale yellow (
<figureCitation id="B59E91D3FFC6AF56FD1EF801FD16F857" box="[643,711,2001,2026]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,263,1907,1930]" captionTargetBox="[340,1246,839,1884]" captionTargetId="figure@2.[340,1247,838,1886]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Empis (Lissempis) guilanensis sp. nov. 1. male habitus (lateral view); 2. fore leg; 3. mid leg; 4. hind leg (scale: 1 mm)." httpUri="https://zenodo.org/record/225534/files/figure.png" pageId="4" pageNumber="189">Fig. 1</figureCitation>
); covered with scattered, short, pale setae; pregenital segments unmodified. Terminalia (
<figureCitation id="B59E91D3FFC7AF57FE24FF47FDD9FF0E" box="[441,520,151,176]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,263,1786,1809]" captionTargetBox="[340,1240,208,1765]" captionTargetId="figure@3.[340,1247,193,1765]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Empis (Lissempis) guilanensis sp. nov. 5. antenna; 6. male genitalia (lateral view) (scale: 0.5 mm)." httpUri="https://zenodo.org/record/225535/files/figure.png" pageId="5" pageNumber="190">Figs 6</figureCitation>
,
<figureCitation id="B59E91D3FFC7AF57FD87FF48FDF8FF0E" box="[538,553,152,176]" captionStart="FIGURE 7" captionStartId="4.[151,249,1325,1347]" captionTargetBox="[344,1237,630,1300]" captionTargetId="figure@4.[340,1247,625,1303]" captionTargetPageId="4" captionText="FIGURE 7. Empis (Lissempis) guilanensis sp. nov., male genitalia, lateral view. Abbreviations: cerc—cercus; epandepandrium; hypd— hypandrium; phal—phallus (scale: 0.5 mm)." httpUri="https://zenodo.org/record/225536/files/figure.png" pageId="5" pageNumber="190">7</figureCitation>
) moderately large. Cercus brown, covered with short black setae; viewed laterally subglobular on basal part, with two apical projections, upper projection short and digitiform, lower projection large and bifurcate apically. Epandrial lamella dark brown; with long, pale setae along lower margin; with deep cleft apically. Hypandrium yellowish brown, bare, concave apically. Phallus yellow, rather thin, evenly bowed, with two loops on apical part.
</paragraph>
<paragraph id="2D1A8D56FFC7AF57FF5AFE9CFE73FEDB" blockId="5.[151,1437,151,680]" box="[199,418,332,357]" pageId="5" pageNumber="190">
<emphasis id="1FD15144FFC7AF57FF5AFE9CFECFFEDB" bold="true" box="[199,286,332,357]" pageId="5" pageNumber="190">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="65BFDEDDFFC7AF57FF5AFEBFFED5FE13" pageId="5" pageNumber="190" type="etymology">
<paragraph id="2D1A8D56FFC7AF57FF5AFEBFFED5FE13" blockId="5.[151,1437,151,680]" pageId="5" pageNumber="190">
<emphasis id="1FD15144FFC7AF57FF5AFEBFFE97FE36" bold="true" box="[199,326,367,392]" pageId="5" pageNumber="190">Etymology</emphasis>
. The name of the new species refers to Guilan Province of
<collectingCountry id="55B2CDC6FFC7AF57FB8AFEA0FB95FE39" box="[1047,1092,368,391]" name="Iran" pageId="5" pageNumber="190">Iran</collectingCountry>
where the
<typeStatus id="F21E33F4FFC7AF57FB50FEA0FB2CFE36" box="[1229,1277,368,392]" pageId="5" pageNumber="190">type</typeStatus>
material was collected.
</paragraph>
</subSubSection>
<subSubSection id="65BFDEDDFFC7AF57FF5AFE67FDD3FE6E" box="[199,514,439,464]" pageId="5" pageNumber="190" type="distribution">
<paragraph id="2D1A8D56FFC7AF57FF5AFE67FDD3FE6E" blockId="5.[151,1437,151,680]" box="[199,514,439,464]" pageId="5" pageNumber="190">
<emphasis id="1FD15144FFC7AF57FF5AFE67FE88FE6E" bold="true" box="[199,345,439,464]" pageId="5" pageNumber="190">Distribution</emphasis>
.
<collectingCountry id="55B2CDC6FFC7AF57FEFAFE68FE45FE71" box="[359,404,440,463]" name="Iran" pageId="5" pageNumber="190">Iran</collectingCountry>
(Guilan).
</paragraph>
</subSubSection>
<subSubSection id="65BFDEDDFFC7AF57FF5AFE0CFBEFFD16" pageId="5" pageNumber="190" type="discussion">
<paragraph id="2D1A8D56FFC7AF57FF5AFE0CFBEFFD16" blockId="5.[151,1437,151,680]" pageId="5" pageNumber="190">
<emphasis id="1FD15144FFC7AF57FF5AFE0CFEE5FE4B" bold="true" box="[199,308,476,501]" pageId="5" pageNumber="190">Remarks</emphasis>
.
<taxonomicName id="EAA5F6D5FFC7AF57FED5FE0DFDCAFE4A" box="[328,539,476,500]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="guilanensis" status="sp. nov.">
<emphasis id="1FD15144FFC7AF57FED5FE0DFDCAFE4A" box="[328,539,476,500]" italics="true" pageId="5" pageNumber="190">Empis guilanensis</emphasis>
</taxonomicName>
<emphasis id="1FD15144FFC7AF57FDB7FE0CFD5BFE4B" bold="true" box="[554,650,476,501]" pageId="5" pageNumber="190">
<taxonomicNameLabel id="04E2EC3FFFC7AF57FDB7FE0CFD5BFE4B" box="[554,650,476,501]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely related to
<taxonomicName id="EAA5F6D5FFC7AF57FC09FE0DFA4DFE4A" authority="Shamshev et Kustov, 2013" authorityName="Shamshev et Kustov" authorityYear="2013" box="[916,1436,476,501]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azishtauensis">
<emphasis id="1FD15144FFC7AF57FC09FE0DFC74FE4A" box="[916,933,477,500]" italics="true" pageId="5" pageNumber="190">E</emphasis>
.
<emphasis id="1FD15144FFC7AF57FC27FE0CFB9EFE4A" box="[954,1103,476,500]" italics="true" pageId="5" pageNumber="190">azishtauensis</emphasis>
<bibRefCitation id="4934F0A7FFC7AF57FBC0FE0CFA4DFE4A" author="Shamshev" box="[1117,1436,476,501]" pageId="5" pageNumber="190" refString="Shamshev, I. &amp; Kustov, S. (2013) Two new species of the Empis subgenus Lissempis (Diptera: Empididae) from the Caucasus. Zootaxa, 3637 (1), 74 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3637.1.8" type="journal article" year="2013">Shamshev et Kustov, 2013</bibRefCitation>
</taxonomicName>
described recently from the North Caucasus (
<collectingCountry id="55B2CDC6FFC7AF57FD31FE2FFD2DFDA6" box="[684,764,511,536]" name="Russia" pageId="5" pageNumber="190">Russia</collectingCountry>
: Republic of Adygea, Krasnodar Territory) (
<bibRefCitation id="4934F0A7FFC7AF57FA9BFE2FFEFCFD82" author="Shamshev" pageId="5" pageNumber="190" refString="Shamshev, I. &amp; Kustov, S. (2013) Two new species of the Empis subgenus Lissempis (Diptera: Empididae) from the Caucasus. Zootaxa, 3637 (1), 74 - 78. http: // dx. doi. org / 10.11646 / zootaxa. 3637.1.8" type="journal article" year="2013">Shamshev &amp; Kustov 2013</bibRefCitation>
). However, the new species differs from
<taxonomicName id="EAA5F6D5FFC7AF57FD60FDF5FC63FD82" box="[765,946,548,572]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azishtauensis">
<emphasis id="1FD15144FFC7AF57FD60FDF5FCDFFD82" box="[765,782,549,572]" italics="true" pageId="5" pageNumber="190">E</emphasis>
.
<emphasis id="1FD15144FFC7AF57FC80FDF4FC63FD82" box="[797,946,548,572]" italics="true" pageId="5" pageNumber="190">azishtauensis</emphasis>
</taxonomicName>
by colour of fore and mid tibiae which are entirely yellow (with brown apex in
<taxonomicName id="EAA5F6D5FFC7AF57FDD8FD99FD2DFDDE" box="[581,764,584,608]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azishtauensis">
<emphasis id="1FD15144FFC7AF57FDD8FD99FD87FDDE" box="[581,598,585,608]" italics="true" pageId="5" pageNumber="190">E</emphasis>
.
<emphasis id="1FD15144FFC7AF57FDFAFD98FD2DFDDE" box="[615,764,584,608]" italics="true" pageId="5" pageNumber="190">azishtauensis</emphasis>
</taxonomicName>
); by the structure of the hind basitarsus which is slightly uniformly thickened (strongly swollen on basal half in
<taxonomicName id="EAA5F6D5FFC7AF57FD61FDBDFC60FD3A" box="[764,945,620,644]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azishtauensis">
<emphasis id="1FD15144FFC7AF57FD61FDBDFCDCFD3A" box="[764,781,621,644]" italics="true" pageId="5" pageNumber="190">E</emphasis>
.
<emphasis id="1FD15144FFC7AF57FC81FDBCFC60FD3A" box="[796,945,620,644]" italics="true" pageId="5" pageNumber="190">azishtauensis</emphasis>
</taxonomicName>
); by ordinary setation of abdominal tergites (with spinule-like setae in
<taxonomicName id="EAA5F6D5FFC7AF57FE5DFD41FDA5FD16" box="[448,628,656,680]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azishtauensis">
<emphasis id="1FD15144FFC7AF57FE5DFD41FE00FD16" box="[448,465,657,680]" italics="true" pageId="5" pageNumber="190">E</emphasis>
.
<emphasis id="1FD15144FFC7AF57FE42FD40FDA5FD16" box="[479,628,656,680]" italics="true" pageId="5" pageNumber="190">azishtauensis</emphasis>
</taxonomicName>
) and by structure of the male terminalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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