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<document id="C1C078403C08E5D11E868C5542742CFC" ID-CLB-Dataset="285334" ID-DOI="10.11646/zootaxa.5399.4.5" ID-GBIF-Dataset="86601a1d-fb31-46f3-9f04-8c990a0e9e7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="10517106" ID-ZooBank="FA43F23F-419F-46C7-B019-D29B71796245" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1705392275901" checkinUser="plazi" docAuthor="Reategui, Natália Soares, Ferreira, Rodrigo Lopes, Rafael, José Albertino &amp; Silva-Neto, Alberto Moreira Da" docDate="2024" docId="7A62A918FFA5FF835F9ACD8FFD37FAC4" docLanguage="en" docName="zootaxa.5399.4.5.pdf" docOrigin="Zootaxa 5399 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5399.4.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptiloneuropsis Roesler 1940" docType="treatment" docVersion="2" lastPageNumber="382" masterDocId="865BD160FFA4FF825F0DCE20FF87FFC9" masterDocTitle="Synopsis of the Ptiloneuropsis Roesler, 1940 (Psocodea: ‘ Psocoptera’: Ptiloneuridae), with a new cave-dwelling species from Brazil" masterLastPageNumber="396" masterPageNumber="381" pageNumber="382" updateTime="1705393137950" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4C958F59E0B959038DD6473A5FBAE3F7">Synopsis of the Ptiloneuropsis Roesler, 1940 (Psocodea: ‘ Psocoptera’: Ptiloneuridae), with a new cave-dwelling species from Brazil</mods:title>
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<mods:namePart id="6824F3F4AD60774AC13D5A6096099269">Rafael, José Albertino</mods:namePart>
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<taxonomicName id="35CB638DFFA5FF835F9ACD8FFE75FC00" ID-CoL="8MTVR" authority="Roesler, 1940" authorityName="Roesler" authorityYear="1940" box="[151,498,943,970]" class="Insecta" family="Ptiloneuridae" genus="Ptiloneuropsis" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="382" phylum="Arthropoda" rank="genus">
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<emphasis id="C0BFC41CFFA5FF835F9ACD8FFEC2FC03" bold="true" box="[151,325,943,970]" italics="true" pageId="1" pageNumber="382">Ptiloneuropsis</emphasis>
Roesler, 1940
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(
<figureCitation id="6AF0048BFFA5FF835F92CDF3FE97FC24" box="[159,272,979,1005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-12="FIGURE 13" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="2.[151,250,1856,1880]" captionStartId-1="3.[151,250,1245,1269]" captionStartId-10="11.[151,250,1980,2004]" captionStartId-11="12.[151,250,424,448]" captionStartId-12="12.[151,250,1354,1378]" captionStartId-2="4.[151,250,1782,1806]" captionStartId-3="5.[151,250,1539,1563]" captionStartId-4="6.[151,250,1442,1466]" captionStartId-5="7.[151,250,1355,1379]" captionStartId-6="8.[151,250,1758,1782]" captionStartId-7="9.[151,250,1943,1967]" captionStartId-8="10.[151,250,1269,1293]" captionStartId-9="11.[151,250,1082,1106]" captionTargetBox-0="[157,1430,185,1824]" captionTargetBox-1="[159,1430,181,1216]" captionTargetBox-10="[156,1430,1172,1952]" captionTargetBox-11="[156,1431,187,395]" captionTargetBox-12="[151,1436,508,1330]" captionTargetBox-2="[151,1436,741,1758]" captionTargetBox-3="[153,1434,525,1501]" captionTargetBox-4="[156,1432,386,1414]" captionTargetBox-5="[156,1430,388,1326]" captionTargetBox-6="[158,1431,280,1728]" captionTargetBox-7="[200,1388,187,1913]" captionTargetBox-8="[156,1430,185,1241]" captionTargetBox-9="[157,1432,187,1058]" captionTargetId-0="figure-25@2.[151,1436,179,1830]" captionTargetId-1="figure-19@3.[151,1436,181,1221]" captionTargetId-10="figure-62@11.[151,1435,1167,1957]" captionTargetId-11="figure-25@12.[151,1435,181,400]" captionTargetId-12="figure-49@12.[151,1436,508,1330]" captionTargetId-2="figure-256@4.[151,1436,741,1758]" captionTargetId-3="figure-134@5.[151,1436,525,1515]" captionTargetId-4="figure-128@6.[151,1435,381,1418]" captionTargetId-5="figure-110@7.[151,1436,381,1332]" captionTargetId-6="figure-67@8.[151,1436,273,1735]" captionTargetId-7="figure-19@9.[194,1394,181,1919]" captionTargetId-8="figure-25@10.[151,1435,181,1245]" captionTargetId-9="figure-19@11.[151,1436,181,1058]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="11" captionTargetPageId-11="12" captionTargetPageId-12="12" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURE 1. Ptiloneuropsis immaculata Roesler (Male neotype).A. Front view of head. B. Left forewing. C. Left hindwing. D. Hypandrium. E. Phallosome. F. Clunium, right paraproct and epiproct. Scale in mm." captionText-1="FIGURE 2. Ptiloneuropsis immaculata Roesler (Male neotype).A. labrum. B. Labium. C. left maxillary palpus and left lacinial tip. D. Right mandible in dorsal view. E. Left mandible in dorsal view. Scales in mm." captionText-10="FIGURE 11. Variation of the forewing M vein in paratypes of Ptiloneuropsis krenaki sp. nov. A–E. Forewings. Scales in mm." captionText-11="FIGURE 12. Variation of the hindwing M vein in paratypes of Ptiloneuropsis krenaki sp. nov. A–B. Hindwings. Scales in mm." captionText-12="FIGURE 13. Distribution of the Ptiloneuropsis species." captionText-2="FIGURE 3. Ptiloneuropsis immaculata Roesler (Male neotype). A. Forelegs. B. Hindlegs. Scales in mm." captionText-3="FIGURE 4. Ptiloneuropsis krenaki sp. nov. Male holotype. A. Side view. B. Zoom of lateral view of head. C. Front view of head. Scales in mm." captionText-4="FIGURE 5. Ptiloneuropsis krenaki sp. nov. Male holotype. A. Right forewing. B. Left forewing. C. Right hindwing. D. Left hindwing. E. Right foreleg. F. Right midleg. G. Right hindleg. Scales in mm." captionText-5="FIGURE 6. Ptiloneuropsis krenaki sp. nov. Male holotype. A. labrum. B. Labium. C. left maxillary palpus. D. Right mandible in dorsal view. E. Left mandible in dorsal view. F. Right lacinial tip. G. Left lacinial tip. Scales in mm." captionText-6="FIGURE 7. Ptiloneuropsis krenaki sp. nov. Male holotype. A. End of abdomen in side view. B. Hypandrium. C. Phallosome. D. Clunium, right paraproct and epiproct. Scales in mm." captionText-7="FIGURE 8. Ptiloneuropsis krenaki sp. nov. Female paratype. A. Side view. B. Zoom of lateral view of head. C. Front view of head. D. Subgenital plate. E. Ninth sternum and left gonapophyses. F. Clunium, right paraproct and epiproct. Scales in mm." captionText-8="FIGURE 9. Ptiloneuropsis krenaki sp. nov. Female paratype. A. Right forewing. B. Left forewing. C. Right hindwing. D. Left hindwing. E. Right foreleg. F. Right midleg. G. Right hindleg. Scales in mm." captionText-9="FIGURE 10. Ptiloneuropsis krenaki sp. nov. Female paratype.A. labrum. B. Labium. C. left maxillary palpus. D. Left mandible in dorsal view. E. Right mandible in dorsal view. F. Right lacinial tip. G. Left lacinial tip. Scales in mm." figureDoi-0="http://doi.org/10.5281/zenodo.10517108" figureDoi-1="http://doi.org/10.5281/zenodo.10517110" figureDoi-10="http://doi.org/10.5281/zenodo.10517130" figureDoi-11="http://doi.org/10.5281/zenodo.10517134" figureDoi-12="http://doi.org/10.5281/zenodo.10517136" figureDoi-2="http://doi.org/10.5281/zenodo.10517112" figureDoi-3="http://doi.org/10.5281/zenodo.10517114" figureDoi-4="http://doi.org/10.5281/zenodo.10517116" figureDoi-5="http://doi.org/10.5281/zenodo.10517118" figureDoi-6="http://doi.org/10.5281/zenodo.10517120" figureDoi-7="http://doi.org/10.5281/zenodo.10517124" figureDoi-8="http://doi.org/10.5281/zenodo.10517126" figureDoi-9="http://doi.org/10.5281/zenodo.10517128" httpUri-0="https://zenodo.org/record/10517108/files/figure.png" httpUri-1="https://zenodo.org/record/10517110/files/figure.png" httpUri-10="https://zenodo.org/record/10517130/files/figure.png" httpUri-11="https://zenodo.org/record/10517134/files/figure.png" httpUri-12="https://zenodo.org/record/10517136/files/figure.png" httpUri-2="https://zenodo.org/record/10517112/files/figure.png" httpUri-3="https://zenodo.org/record/10517114/files/figure.png" httpUri-4="https://zenodo.org/record/10517116/files/figure.png" httpUri-5="https://zenodo.org/record/10517118/files/figure.png" httpUri-6="https://zenodo.org/record/10517120/files/figure.png" httpUri-7="https://zenodo.org/record/10517124/files/figure.png" httpUri-8="https://zenodo.org/record/10517126/files/figure.png" httpUri-9="https://zenodo.org/record/10517128/files/figure.png" pageId="1" pageNumber="382">Figs 1–13</figureCitation>
)
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<emphasis id="C0BFC41CFFA5FF835F9ACA3BFEBEFBFC" box="[151,313,1051,1077]" italics="true" pageId="1" pageNumber="382">Ptiloneuropsis</emphasis>
Roesler, 1940: 236
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;
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<bibRefCitation id="965A65FFFFA5FF835D12CA3BFD78FBFC" author="Smithers, C. N." box="[543,767,1051,1077]" pageId="1" pageNumber="382" pagination="1 - 349" refId="ref8208" refString="Smithers, C. N. (1972) The classification and phylogeny of the Psocoptera. Australian Museum Memoirs, 14, 1 - 349. https: // doi. org / 10.3853 / j. 0067 - 1967.14.1972.424" type="journal article" year="1972">Smithers, 1972: 112</bibRefCitation>
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(classification and phylogeny);
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(catalogue);
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<bibRefCitation id="965A65FFFFA5FF835E58CA1FFDFFFB90" author="Garcia Aldrete, A. N." box="[341,632,1087,1113]" pageId="1" pageNumber="382" pagination="145 - 148" refId="ref7392" refString="Garcia Aldrete, A. N. (2004) Rediscovery of Ptiloneuropsis immaculata Roesler 1940 (Psocoptera: Ptiloneuridae), augmented description and neotype designation. Studies on Neotropical Fauna and Environment, 39 (2), 145 - 148. https: // doi. org / 10.1080 / 01650520412331333792" type="journal article" year="2004">García Aldrete, 2004: 146</bibRefCitation>
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(
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designation);
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<bibRefCitation id="965A65FFFFA5FF835C8CCA1FFB36FB93" author="Silva-Neto, A. M. &amp; Garcia Aldrete, A. N. &amp; Rafael, J. A." box="[897,1201,1087,1114]" pageId="1" pageNumber="382" pagination="567 - 574" refId="ref7988" refString="Silva-Neto, A. M., Garcia Aldrete, A. N. &amp; Rafael, J. A. (2018) After seventy-seven years Ptiloneuropsis Roesler (Psocodea: Psocomorpha: Ptiloneuridae) ceases to be a monotypic genus: a new species from Bahia, Brazil. Zootaxa, 4388 (4), 567 - 574. https: // doi. org / 10.11646 / zootaxa. 4388.4.8" type="journal article" year="2018">
Silva-Neto
<emphasis id="C0BFC41CFFA5FF835B0CCA60FBB5FB90" box="[1025,1074,1087,1113]" italics="true" pageId="1" pageNumber="382">et al</emphasis>
. 2018: 569
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(taxonomy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="BAD14B85FFA5FF835FCACA43FD37FAC4" pageId="1" pageNumber="382" type="diagnosis">
<paragraph id="F274180EFFA5FF835FCACA43FD37FAC4" blockId="1.[151,1437,1051,1294]" pageId="1" pageNumber="382">
<emphasis id="C0BFC41CFFA5FF835FCACA43FE27FBB4" bold="true" box="[199,416,1123,1149]" pageId="1" pageNumber="382">Update diagnosis:</emphasis>
Forewings M vein with four (see fig.
<quantity id="3533B5EBFFA5FF835C6ECA43FC1AFBB4" box="[867,925,1123,1149]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="1" pageNumber="382" unit="in" value="11.0">11 in</quantity>
<bibRefCitation id="965A65FFFFA5FF835CAACA43FB2EFBB4" author="Silva-Neto, A. M. &amp; Garcia Aldrete, A. N. &amp; Rafael, J. A." box="[935,1193,1123,1150]" pageId="1" pageNumber="382" pagination="567 - 574" refId="ref7988" refString="Silva-Neto, A. M., Garcia Aldrete, A. N. &amp; Rafael, J. A. (2018) After seventy-seven years Ptiloneuropsis Roesler (Psocodea: Psocomorpha: Ptiloneuridae) ceases to be a monotypic genus: a new species from Bahia, Brazil. Zootaxa, 4388 (4), 567 - 574. https: // doi. org / 10.11646 / zootaxa. 4388.4.8" type="journal article" year="2018">
Silva-Neto
<emphasis id="C0BFC41CFFA5FF835B27CA44FBD8FBB4" box="[1066,1119,1123,1149]" italics="true" pageId="1" pageNumber="382">et al</emphasis>
. 2018
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) to seven (
<figureCitation id="6AF0048BFFA5FF835A21CA43FA13FBB4" box="[1324,1428,1123,1149]" captionStart="FIGURE 11" captionStartId="11.[151,250,1980,2004]" captionTargetBox="[156,1430,1172,1952]" captionTargetId="figure-62@11.[151,1435,1167,1957]" captionTargetPageId="11" captionText="FIGURE 11. Variation of the forewing M vein in paratypes of Ptiloneuropsis krenaki sp. nov. A–E. Forewings. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517130" httpUri="https://zenodo.org/record/10517130/files/figure.png" pageId="1" pageNumber="382">Fig. 11D</figureCitation>
) primary branches, forewing areola postica triangular, joined to M vein by a cross vein arising from its apex (
<figureCitation id="6AF0048BFFA5FF835A67CAA7FF3DFB0C" captionStart="FIGURE 1" captionStartId="2.[151,250,1856,1880]" captionTargetBox="[157,1430,185,1824]" captionTargetId="figure-25@2.[151,1436,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Ptiloneuropsis immaculata Roesler (Male neotype).A. Front view of head. B. Left forewing. C. Left hindwing. D. Hypandrium. E. Phallosome. F. Clunium, right paraproct and epiproct. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.10517108" httpUri="https://zenodo.org/record/10517108/files/figure.png" pageId="1" pageNumber="382">Figs 1B</figureCitation>
;
<figureCitation id="6AF0048BFFA5FF835FC8CA8BFE8DFB0F" box="[197,266,1195,1222]" captionStart="FIGURE 5" captionStartId="6.[151,250,1442,1466]" captionTargetBox="[156,1432,386,1414]" captionTargetId="figure-128@6.[151,1435,381,1418]" captionTargetPageId="6" captionText="FIGURE 5. Ptiloneuropsis krenaki sp. nov. Male holotype. A. Right forewing. B. Left forewing. C. Right hindwing. D. Left hindwing. E. Right foreleg. F. Right midleg. G. Right hindleg. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517116" httpUri="https://zenodo.org/record/10517116/files/figure.png" pageId="1" pageNumber="382">5A–B</figureCitation>
,
<figureCitation id="6AF0048BFFA5FF835E18CA8BFEEEFB0C" box="[277,361,1195,1221]" captionStart="FIGURE 11" captionStartId="11.[151,250,1980,2004]" captionTargetBox="[156,1430,1172,1952]" captionTargetId="figure-62@11.[151,1435,1167,1957]" captionTargetPageId="11" captionText="FIGURE 11. Variation of the forewing M vein in paratypes of Ptiloneuropsis krenaki sp. nov. A–E. Forewings. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517130" httpUri="https://zenodo.org/record/10517130/files/figure.png" pageId="1" pageNumber="382">11A–G</figureCitation>
,
<figureCitation id="6AF0048BFFA5FF835E7ECA8BFE4FFB0C" box="[371,456,1195,1221]" captionStart="FIGURE 12" captionStartId="12.[151,250,424,448]" captionTargetBox="[156,1431,187,395]" captionTargetId="figure-25@12.[151,1435,181,400]" captionTargetPageId="12" captionText="FIGURE 12. Variation of the hindwing M vein in paratypes of Ptiloneuropsis krenaki sp. nov. A–B. Hindwings. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517134" httpUri="https://zenodo.org/record/10517134/files/figure.png" pageId="1" pageNumber="382">12A–C</figureCitation>
). Hindwings M with three to five primary branches (
<figureCitation id="6AF0048BFFA5FF835B06CA8BFBE3FB0C" box="[1035,1124,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1856,1880]" captionTargetBox="[157,1430,185,1824]" captionTargetId="figure-25@2.[151,1436,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Ptiloneuropsis immaculata Roesler (Male neotype).A. Front view of head. B. Left forewing. C. Left hindwing. D. Hypandrium. E. Phallosome. F. Clunium, right paraproct and epiproct. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.10517108" httpUri="https://zenodo.org/record/10517108/files/figure.png" pageId="1" pageNumber="382">Figs 1C</figureCitation>
,
<figureCitation id="6AF0048BFFA5FF835B7DCA8BFB32FB0F" box="[1136,1205,1195,1222]" captionStart="FIGURE 5" captionStartId="6.[151,250,1442,1466]" captionTargetBox="[156,1432,386,1414]" captionTargetId="figure-128@6.[151,1435,381,1418]" captionTargetPageId="6" captionText="FIGURE 5. Ptiloneuropsis krenaki sp. nov. Male holotype. A. Right forewing. B. Left forewing. C. Right hindwing. D. Left hindwing. E. Right foreleg. F. Right midleg. G. Right hindleg. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517116" httpUri="https://zenodo.org/record/10517116/files/figure.png" pageId="1" pageNumber="382">5C–D</figureCitation>
,
<figureCitation id="6AF0048BFFA5FF835BB2CA8BFB59FB0C" box="[1215,1246,1195,1221]" captionStart="FIGURE 12" captionStartId="12.[151,250,424,448]" captionTargetBox="[156,1431,187,395]" captionTargetId="figure-25@12.[151,1435,181,400]" captionTargetPageId="12" captionText="FIGURE 12. Variation of the hindwing M vein in paratypes of Ptiloneuropsis krenaki sp. nov. A–B. Hindwings. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517134" httpUri="https://zenodo.org/record/10517134/files/figure.png" pageId="1" pageNumber="382">12</figureCitation>
). Hypandrium of one sclerite, with a protuberance in its anterior median region, in which the anterior ends of the side struts fit (
<figureCitation id="6AF0048BFFA5FF835A67CAEFFF3CFAC4" captionStart="FIGURE 1" captionStartId="2.[151,250,1856,1880]" captionTargetBox="[157,1430,185,1824]" captionTargetId="figure-25@2.[151,1436,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Ptiloneuropsis immaculata Roesler (Male neotype).A. Front view of head. B. Left forewing. C. Left hindwing. D. Hypandrium. E. Phallosome. F. Clunium, right paraproct and epiproct. Scale in mm." figureDoi="http://doi.org/10.5281/zenodo.10517108" httpUri="https://zenodo.org/record/10517108/files/figure.png" pageId="1" pageNumber="382">Figs 1D</figureCitation>
,
<figureCitation id="6AF0048BFFA5FF835FCACAD4FF6FFAC7" box="[199,232,1268,1294]" captionStart="FIGURE 7" captionStartId="8.[151,250,1758,1782]" captionTargetBox="[158,1431,280,1728]" captionTargetId="figure-67@8.[151,1436,273,1735]" captionTargetPageId="8" captionText="FIGURE 7. Ptiloneuropsis krenaki sp. nov. Male holotype. A. End of abdomen in side view. B. Hypandrium. C. Phallosome. D. Clunium, right paraproct and epiproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.10517120" httpUri="https://zenodo.org/record/10517120/files/figure.png" pageId="1" pageNumber="382">7B</figureCitation>
and see fig.
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<bibRefCitation id="965A65FFFFA5FF835EA5CAD3FD23FAC4" author="Silva-Neto, A. M. &amp; Garcia Aldrete, A. N. &amp; Rafael, J. A." box="[424,676,1267,1293]" pageId="1" pageNumber="382" pagination="567 - 574" refId="ref7988" refString="Silva-Neto, A. M., Garcia Aldrete, A. N. &amp; Rafael, J. A. (2018) After seventy-seven years Ptiloneuropsis Roesler (Psocodea: Psocomorpha: Ptiloneuridae) ceases to be a monotypic genus: a new species from Bahia, Brazil. Zootaxa, 4388 (4), 567 - 574. https: // doi. org / 10.11646 / zootaxa. 4388.4.8" type="journal article" year="2018">
Silva-Neto
<emphasis id="C0BFC41CFFA5FF835D25CAD4FDE6FAC4" box="[552,609,1267,1293]" italics="true" pageId="1" pageNumber="382">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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