156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="E9CCBB06CA76C9F91CCD2918EFC10DFE" ID-DOI="10.11646/zootaxa.3931.4.3" ID-GBIF-Dataset="d954dbb2-b586-4b02-984c-0bb8f3709ab0" ID-ISSN="1175-5326" ID-Zenodo-Dep="234953" ID-ZooBank="918B6F76-B5CA-4FC6-9F32-F499DDE7C420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461011379296" checkinUser="plazi" docAuthor="Deschodt, Christian M., Davis, Adrian L. V. & Scholtz, Clarke H." docDate="2015" docId="7A32765450549F4EF5E307ABB081FD30" docLanguage="en" docName="zt03931p527.pdf" docOrigin="Zootaxa 3931 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scarabaeus (Scarabaeolus) niemandi Deschodt and Davis" docType="treatment" docVersion="7" lastPageNumber="510" masterDocId="860B0E2C50509F4BF5740364B06DFFB4" masterDocTitle="Status changes, new synonymies, key and descriptions of seven new species in the subgenus Scarabaeus (Scarabaeolus) Balthasar 1965 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="527" masterPageNumber="505" pageNumber="509" updateTime="1698567509984" updateUser="plazi">
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<mods:titleInfo id="093C0F75F1277FD183133148F6474911">
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<mods:title id="E30C389DEBA835DA88861A2E08A0CCC2">Status changes, new synonymies, key and descriptions of seven new species in the subgenus Scarabaeus (Scarabaeolus) Balthasar 1965 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="8CB9E02DFE0655B94395DF013234C4D0" type="personal">
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<mods:roleTerm id="D7EA5576A1C151F0C1CE64EE7E60A88C">Author</mods:roleTerm>
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<mods:namePart id="7D3897344F222ACEAFB3D865EDA2D30C">Deschodt, Christian M.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="EC4DC6459AFC0EBF72C9F4753B2B19A2">Author</mods:roleTerm>
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<mods:namePart id="B4AA865DCA011613712D17F47C1E1AD0">Davis, Adrian L. V.</mods:namePart>
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</mods:name>
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<mods:name id="736E1080C7648F13FC2173B81ACBB21E" type="personal">
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<mods:roleTerm id="829AD6F8EF612F8C019C9EC3826FB60D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58FD6E16B7B88C3F0FE8F9A0D71FFDD2">Scholtz, Clarke H.</mods:namePart>
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<mods:typeOfResource id="4B7C30DE77E52AC85418D3628D8502D5">text</mods:typeOfResource>
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<mods:title id="A051C6A749352B54BA210767E21DA777">Zootaxa</mods:title>
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<mods:part id="72C74667036FA624472D2EBDDA436836">
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<mods:date id="67FAE8F0181DAFDC2292741DC255ADCE">2015</mods:date>
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<mods:detail id="54EB8A3617BDA626A861014A6037C4E5" type="volume">
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<mods:number id="1EFB4801BD9FDDEC11A2B7FD0B591914">3931</mods:number>
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<mods:detail id="573827FA97D55D3524126FB7323CBC70" type="issue">
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<mods:number id="B2189ACB23D9059DCAF8E44649A5CA01">4</mods:number>
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<mods:start id="DC267224800A9F6BA111DB554B33BF45">505</mods:start>
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<mods:classification id="7F4340E73D04EC5BA4E5DFBF6B5B1F1E">journal article</mods:classification>
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<mods:identifier id="F5D79629D27277856144A43DCB348736" type="DOI">10.11646/zootaxa.3931.4.3</mods:identifier>
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<mods:identifier id="E1D8D9B34D4EA92146A0C4DA309037F6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="7A32765450549F4EF5E307ABB081FD30" ID-DOI="http://doi.org/10.5281/zenodo.6108502" ID-GBIF-Taxon="119600185" ID-Zenodo-Dep="6108502" LSID="urn:lsid:plazi:treatment:7A32765450549F4EF5E307ABB081FD30" httpUri="http://treatment.plazi.org/id/7A32765450549F4EF5E307ABB081FD30" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509">
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<subSubSection id="BA8194C950549F4FF5E307ABB140FABF" pageId="4" pageNumber="509" type="nomenclature">
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<paragraph id="F224C74250549F4FF5E307ABB334FB5E" blockId="4.[151,857,1231,1291]" box="[151,857,1231,1258]" pageId="4" pageNumber="509">
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<heading id="A96C702E50549F4FF5E307ABB334FB5E" bold="true" box="[151,857,1231,1258]" fontSize="11" level="1" pageId="4" pageNumber="509" reason="1">
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<taxonomicName id="359BBCC150549F4FF5E307ABB334FB5E" ID-CoL="79W69" authority="Deschodt and Davis" authorityName="Deschodt and Davis" box="[151,857,1231,1258]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="niemandi" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF5E307ABB334FB5E" bold="true" box="[151,857,1231,1258]" pageId="4" pageNumber="509">
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<emphasis id="C0EF1B5050549F4FF5E307ABB237FB5D" bold="true" box="[151,602,1231,1257]" italics="true" pageId="4" pageNumber="509">Scarabaeus (Scarabaeolus) niemandi</emphasis>
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Deschodt and Davis
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="F224C74250549F4FF5E30796B140FABF" blockId="4.[151,857,1231,1291]" box="[151,301,1266,1291]" pageId="4" pageNumber="509">Figure 3, 6, 8</paragraph>
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</subSubSection>
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<subSubSection id="BA8194C950549F4FF5E3065EB412FA56" pageId="4" pageNumber="509" type="diagnosis">
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<paragraph id="F224C74250549F4FF5E3065EB412FA56" blockId="4.[151,1437,1338,2011]" pageId="4" pageNumber="509">
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<emphasis id="C0EF1B5050549F4FF5E3065EB17CFAE7" bold="true" box="[151,273,1338,1363]" pageId="4" pageNumber="509">Diagnosis.</emphasis>
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<taxonomicName id="359BBCC150549F4FF469065FB2A1FAE7" box="[285,716,1339,1363]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="niemandi" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF469065FB2A1FAE7" box="[285,716,1339,1363]" italics="true" pageId="4" pageNumber="509">Scarabaeus (Scarabaeolus) niemandi</emphasis>
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</taxonomicName>
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is characterized by short black setae on the head and often punctate interstriae compared to brown setae on the head with interstriae that are smooth along their entire length in
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<taxonomicName id="359BBCC150549F4FF5E306E7B126FA2F" box="[151,331,1411,1435]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="lucidulus" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF5E306E7B126FA2F" box="[151,331,1411,1435]" italics="true" pageId="4" pageNumber="509">S. (S.) lucidulus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="359BBCC150549F4FF42E06E7B27DFA2F" box="[346,528,1411,1435]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="niemandi" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF42E06E7B27DFA2F" box="[346,528,1411,1435]" italics="true" pageId="4" pageNumber="509">S. (S.) niemandi</emphasis>
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</taxonomicName>
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can be separated from
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<taxonomicName id="359BBCC150549F4FF66806E7B5F1FA2F" authority="Waterhouse, 1890" authorityName="Waterhouse" authorityYear="1890" box="[796,1436,1410,1435]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="anderseni" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF66806E7B4A5FA2F" box="[796,1224,1411,1435]" italics="true" pageId="4" pageNumber="509">Scarabaeus (Scarabaeolus) anderseni</emphasis>
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<bibRefCitation id="960ABAB350549F4FF1A506E6B5F1FA2F" author="Waterhouse" box="[1233,1436,1410,1435]" pageId="4" pageNumber="509" refString="Waterhouse, C. O. (1890) New Scarabaeidae in the British Museum. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 29, 365 - 373." type="journal article" year="1890">Waterhouse, 1890</bibRefCitation>
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</taxonomicName>
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by its larger size and black setae on the head compared to yellow setae on the head of
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<taxonomicName id="359BBCC150549F4FF12806C3B578FA0A" box="[1116,1301,1446,1470]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="anderseni" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF12806C3B578FA0A" box="[1116,1301,1446,1470]" italics="true" pageId="4" pageNumber="509">S. (S.) anderseni</emphasis>
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</taxonomicName>
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. Punctation on the elytra of
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<taxonomicName id="359BBCC150549F4FF43C06AFB196FA57" box="[328,507,1483,1507]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="niemandi" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF43C06AFB196FA57" box="[328,507,1483,1507]" italics="true" pageId="4" pageNumber="509">S. (S.) niemandi</emphasis>
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</taxonomicName>
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does not often open into the striae as in
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<taxonomicName id="359BBCC150549F4FF6CB06AFB415FA57" box="[959,1144,1483,1507]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="anderseni" subGenus="Scarabaeolus">
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<emphasis id="C0EF1B5050549F4FF6CB06AFB415FA57" box="[959,1144,1483,1507]" italics="true" pageId="4" pageNumber="509">S. (S.) anderseni</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="BA8194C950549F4EF5B3068AB1B9FE19" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509" type="description">
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<paragraph id="F224C74250549F4FF5B3068AB084F99F" blockId="4.[151,1437,1338,2011]" pageId="4" pageNumber="509">
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<emphasis id="C0EF1B5050549F4FF5B3068AB134F9B3" bold="true" box="[199,345,1518,1543]" pageId="4" pageNumber="509">Description.</emphasis>
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<typeStatus id="2D2079E050549F4FF417068AB1BDF9B3" box="[355,464,1518,1543]" pageId="4" pageNumber="509" type="holotype">Holotype</typeStatus>
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: Body length
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<quantity id="35636AA750549F4FF702068BB2B2F9B2" box="[630,735,1519,1543]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="4" pageNumber="509" unit="mm" value="11.7">11.7 mm</quantity>
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, metallic green. Pubescence dark brown to black. Antennae yellow.
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</paragraph>
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<paragraph id="F224C74250549F4FF5B30552B31BF9C6" blockId="4.[151,1437,1338,2011]" pageId="4" pageNumber="509">Head: Frons punctate, each puncture with associated short, stocky black setae. Smooth, puncture free midlongitudinal area. Clypeus punctate with short stocky black setae.</paragraph>
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<paragraph id="F224C74250549F4FF5B3051BB324F96B" blockId="4.[151,1437,1338,2011]" pageId="4" pageNumber="509">Pronotum: Metallic green, punctate, space between punctures smooth, punctures separated by less than one puncture diameter, punctures with short stocky black associated setae. Longitudinal smooth area not extending to anterior edge of pronotum. Lateral margins curved, crenulate.</paragraph>
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<paragraph id="F224C74250549F4FF5B3058EB166F8FE" blockId="4.[151,1437,1338,2011]" pageId="4" pageNumber="509">Elytra: Metallic green, lateral sides more or less parallel. Striae shallow with regular small punctures separated by less than two puncture diameters; punctures extending into margins of interstriae. Interstriae shiny with irregular punctures.</paragraph>
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<paragraph id="F224C74250549F4FF5B30432B316F8DB" blockId="4.[151,1437,1338,2011]" box="[199,891,1878,1903]" pageId="4" pageNumber="509">Pygidium: Emarginate and densely punctate, punctures small.</paragraph>
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<paragraph id="F224C74250549F4FF5B3041FB2F1F86F" blockId="4.[151,1437,1338,2011]" pageId="4" pageNumber="509">Metasternum: Long thin fovea between the mesocoxae, distally wider, with a shorter fovea more distally. Metasternal elevation ending with rounded angle and gradually sloping toward mesosternum. Area behind mesocoxae densely punctate, punctures small.</paragraph>
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<paragraph id="F224C74250559F4EF5B303F3B48FFF04" blockId="5.[151,1436,151,644]" box="[199,1250,151,176]" pageId="5" pageNumber="510">Abdominal sternites: Each sternite with row of punctures and associated dark setae anteriorly.</paragraph>
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<paragraph id="F224C74250559F4EF5B303D9B38EFE3C" blockId="5.[151,1436,151,644]" pageId="5" pageNumber="510">Legs: Anterior femora shallowly punctate ventrally with black setae. Outside margin of anterior tibiae bearing small saw-like teeth proximally and four outward pointing teeth distally, each separated by six to eight saw-like denticles. Inside margin crenulate. Meso- and metafemora with median row of punctures and long fine dark brown setae. Inside margin of mesotibiae smooth with row of black setae, outside margin with two brushes of dark brown setae in distal third. Vestigial second mesotibial spur present. Metatibiae with black setae. All tarsomeres of metatarsi with black setae, last tarsomere longest with claws half its length.</paragraph>
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<paragraph id="F224C74250559F4EF5B302F0B1B9FE19" blockId="5.[151,1436,151,644]" box="[199,468,404,429]" pageId="5" pageNumber="510">Aedeagus: see Figure 6.</paragraph>
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</subSubSection>
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<subSubSection id="BA8194C950559F4EF5B302D3B082FE41" pageId="5" pageNumber="510" type="etymology">
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<paragraph id="F224C74250559F4EF5B302D3B082FE41" blockId="5.[151,1436,151,644]" pageId="5" pageNumber="510">
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<emphasis id="C0EF1B5050559F4EF5B302D3B123FE64" bold="true" box="[199,334,439,464]" pageId="5" pageNumber="510">Etymology.</emphasis>
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This species is named in honour of Lukas Niemand who collected most of the
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<typeStatus id="2D2079E050559F4EF1A002DCB569FE64" box="[1236,1284,440,464]" pageId="5" pageNumber="510">type</typeStatus>
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series of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="BA8194C950559F4EF5B3029BB081FD30" pageId="5" pageNumber="510" type="materials_examined">
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||
<paragraph id="F224C74250559F4EF5B3029BB081FD30" blockId="5.[151,1436,151,644]" pageId="5" pageNumber="510">
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<emphasis id="C0EF1B5050559F4EF5B3029BB1D6FDAC" bold="true" box="[199,443,511,536]" pageId="5" pageNumber="510">Material examined.</emphasis>
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<typeStatus id="2D2079E050559F4EF4B8029BB255FDAC" box="[460,568,511,536]" pageId="5" pageNumber="510" type="holotype">Holotype</typeStatus>
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: ♂, Soutpansberg, Limpopo,
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<geoCoordinate id="97AFA18550559F4EF6D8029BB44EFDAC" box="[940,1059,511,536]" direction="south" orientation="latitude" pageId="5" pageNumber="510" precision="1" value="-22.87561">S22.87561</geoCoordinate>
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<geoCoordinate id="97AFA18550559F4EF157029BB4DEFDAC" box="[1059,1203,511,536]" direction="east" orientation="longitude" pageId="5" pageNumber="510" precision="1" value="29.73128">ºE29.73128º</geoCoordinate>
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,
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<date id="8625E18250559F4EF1BC029BB526FDAC" box="[1224,1355,511,536]" pageId="5" pageNumber="510" value="2009-02-03" valueMax="2009-02-04" valueMin="2009-02-03">3–4.ii.2009</date>
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, L. J. Niemand,
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<collectionCode id="948A5F8750559F4EF4650140B10FFD88" box="[273,354,548,572]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="510">TMSA</collectionCode>
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||
.
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<typeStatus id="2D2079E050559F4EF4070141B189FD89" box="[371,484,549,573]" pageId="5" pageNumber="510" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="E49D0CCB50559F4EF4800140B2EAFD89" box="[500,647,548,573]" pageId="5" pageNumber="510" type="generic">5 specimens</specimenCount>
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||
, unsexed, same data as
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<typeStatus id="2D2079E050559F4EF6D90140B47EFD89" box="[941,1043,548,573]" pageId="5" pageNumber="510" type="holotype">holotype</typeStatus>
|
||
. 2
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<collectionCode id="948A5F8750559F4EF1490140B4EAFD88" box="[1085,1159,548,572]" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="5" pageNumber="510">UPSA</collectionCode>
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||
, 1
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<collectionCode id="948A5F8750559F4EF1C70140B569FD88" box="[1203,1284,548,572]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="510">TMSA</collectionCode>
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, 2
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<collectionCode id="948A5F8750559F4EF0440140B513FD88" LSID="urn:lsid:biocol.org:col:34248" box="[1328,1406,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="5" pageNumber="510">SANC</collectionCode>
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||
.
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<specimenCount id="E49D0CCB50559F4EF0FA0140B169FDD4" pageId="5" pageNumber="510" type="generic">1 specimen</specimenCount>
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, unsexed, S.Afr.; Limpopo Prov., Mabote farm, 24[º].07[’] S 28[º]. 39[’] E,
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<date id="8625E18250559F4EF12C012CB4BAFDD4" box="[1112,1239,584,608]" pageId="5" pageNumber="510" value="2009-11-14">14.11.2009</date>
|
||
, leg.Ruth Müller,
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<collectionCode id="948A5F8750559F4EF5E30108B084FD30" box="[151,233,620,644]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="5" pageNumber="510">TMSA</collectionCode>
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.
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</paragraph>
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</subSubSection>
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</treatment>
|
||
</document> |