289 lines
38 KiB
XML
289 lines
38 KiB
XML
<document id="43635A297CBDCC76E9A1FAD509E528C4" ID-DOI="10.5281/zenodo.5276563" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630012980284" checkinUser="carolina" docAuthor="Assing, V." docDate="2009" docId="6D0E87E4EA08617244F8F60FFD30FEDF" docLanguage="en" docName="LinzerbiolBeitr.2009.41.2.1191-1246.pdf" docOrigin="Linzer biologische Beiträge 41 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba (Tropogastrosipalia) hasanica Assing 2009, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1203" masterDocId="9137FF9CEA01617E444AF536FFEAFFD1" masterDocTitle="A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and a catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1246" masterPageNumber="1191" pageNumber="1200" updateTime="1699136907799" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="ED229CAC0BF0B5EF209060CD7D9C25FE">A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and a catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B201AD2CC722A6EBACEA781837224811" type="personal">
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<mods:roleTerm id="A74E5DA16A877EDE01C08BA5DA7B7ED7">Author</mods:roleTerm>
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<mods:namePart id="35A099C4DA462C833DE90822BF77AB80">Assing, V.</mods:namePart>
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<mods:typeOfResource id="5891926355D9B7966C1497B352645947">text</mods:typeOfResource>
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<mods:title id="0321C155ECAEB48D58F1C4286CA0142B">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="E4DA51FB500CA6B5AE9CB55DD56482E3">
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<mods:date id="C9D7F034225BE66FA3EA926E166F2190">2009</mods:date>
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<mods:detail id="2382009DB92C6D02BB5166325F5DD998" type="pubDate">
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<mods:number id="E00AE92FC6F06633AA7B1EF1C5082DDA">2009-12-18</mods:number>
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<mods:detail id="C0E99A5B5658B32691B8B496640CFD7F" type="volume">
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<mods:number id="0C5C519BE8D976DA27AD62EB2F98F32A">41</mods:number>
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<mods:detail id="E538BB3017F1BCD9137677C5EBC1E838" type="issue">
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<mods:number id="BC09CF20B20D0AE60A8D8F39BFD607FE">2</mods:number>
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<mods:start id="88A7B59DB23F292DB77F76AA9E47F855">1191</mods:start>
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<mods:end id="523C75B1D7065056A748D06B13BE3084">1246</mods:end>
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<mods:classification id="EA193F3B525B626A850680C8A92337DA">journal article</mods:classification>
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<mods:identifier id="DCA97C335BE1BB84699CDF748FDC1057" type="DOI">10.5281/zenodo.5276563</mods:identifier>
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<mods:identifier id="89B11F976E64109138FE05F022C33617" type="ISSN">0253-116X</mods:identifier>
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<treatment id="6D0E87E4EA08617244F8F60FFD30FEDF" LSID="urn:lsid:plazi:treatment:6D0E87E4EA08617244F8F60FFD30FEDF" httpUri="http://treatment.plazi.org/id/6D0E87E4EA08617244F8F60FFD30FEDF" lastPageId="12" lastPageNumber="1203" pageId="9" pageNumber="1200">
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<subSubSection id="ADBD6579EA08617744F8F60FFCCEFC9E" box="[178,804,825,847]" pageId="9" pageNumber="1200" type="nomenclature">
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<paragraph id="E51836F2EA08617744F8F60FFCCEFC9E" blockId="9.[178,804,825,847]" box="[178,804,825,847]" pageId="9" pageNumber="1200">
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<heading id="BE50819EEA08617744F8F60FFCCEFC9E" box="[178,804,825,847]" fontSize="8" level="2" pageId="9" pageNumber="1200" reason="2">
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<emphasis id="D7D3EAE0EA08617744F8F60FFD4AFC9E" bold="true" box="[178,672,825,847]" pageId="9" pageNumber="1200">
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<taxonomicName id="22A74D71EA08617744F8F60FFDA7FC9E" ID-CoL="3FSV9" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[178,589,825,847]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1200" phylum="Arthropoda" rank="species" species="hasanica" status="sp. nov." subGenus="Tropogastrosipalia">
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<emphasis id="D7D3EAE0EA08617744F8F60FFEE7FC9E" bold="true" box="[178,269,825,847]" italics="true" pageId="9" pageNumber="1200">Geostiba</emphasis>
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(
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<emphasis id="D7D3EAE0EA0861774556F60FFE0BFC9E" bold="true" box="[284,481,825,847]" italics="true" pageId="9" pageNumber="1200">Tropogastrosipalia</emphasis>
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)
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<emphasis id="D7D3EAE0EA08617745A5F60FFDA7FC9E" bold="true" box="[495,589,825,847]" italics="true" pageId="9" pageNumber="1200">hasanica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CCE0579BEA0861774619F60FFD4AFC9E" box="[595,672,825,847]" pageId="9" pageNumber="1200" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="7D9C2A77EA08617746E7F60FFCF6FC9E" box="[685,796,825,847]" captionStart-0="Figs 16-22" captionStart-1="Fig" captionStartId-0="10.[178,218,1318,1338]" captionStartId-1="11.[178,211,1527,1547]" captionTargetBox-0="[184,1085,219,1296]" captionTargetBox-1="[178,1085,947,1515]" captionTargetId-0="figure-143@10.[178,1085,219,1306]" captionTargetId-1="figure-362@11.[178,1085,947,1515]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." captionText-1="Fig. 23: Type locality of G. hasanica nov.sp. (photo: P. Wunderle)." figureDoi-0="http://doi.org/10.5281/zenodo.5276573" figureDoi-1="http://doi.org/10.5281/zenodo.5276575" httpUri-0="https://zenodo.org/record/5276573/files/figure.png" httpUri-1="https://zenodo.org/record/5276575/files/figure.png" pageId="9" pageNumber="1200">Figs 16-23</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ADBD6579EA08617244F8F653FD30FEDF" lastPageId="12" lastPageNumber="1203" pageId="9" pageNumber="1200" type="materials_examined">
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<paragraph id="E51836F2EA08617744F8F653FDE7FC6C" blockId="9.[178,1085,868,957]" pageId="9" pageNumber="1200">
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<materialsCitation id="55CF3CAFEA08617744F8F653FEDAFC76" collectingDate="2009-04-10" collectorName="V. Assing" determinerName="V. Assing" elevation="1370" latitude="41.760834" location="Inebolu" longLatPrecision="19" longitude="34.04333" pageId="9" pageNumber="1200" specimenCount="2" stateProvince="Kastamonu" typeStatus="holotype">
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<typeStatus id="3A1C8850EA08617744F8F653FEEDFCA8" box="[178,263,869,889]" pageId="9" pageNumber="1200" type="holotype">Holotype</typeStatus>
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Ƌ: "TR [38] -
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<collectingRegion id="2763F810EA08617745F8F653FDF6FCA8" box="[434,540,869,889]" country="Turkey" name="Kastamonu" pageId="9" pageNumber="1200">Kastamonu</collectingRegion>
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,
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<quantity id="225F9B17EA0861774666F653FD86FCA8" box="[556,620,869,889]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="1200" unit="km" value="30.0">30 km</quantity>
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SE
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<location id="E0786029EA08617746EAF653FD02FCA8" LSID="urn:lsid:plazi:treatment:6D0E87E4EA08617244F8F60FFD30FEDF:E0786029EA08617746EAF653FD02FCA8" box="[672,744,869,889]" latitude="41.760834" longLatPrecision="19" longitude="34.04333" name="Inebolu" pageId="9" pageNumber="1200" stateProvince="Kastamonu">Inebolu</location>
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,
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<geoCoordinate id="80935035EA08617746B2F653FC8EFCA8" box="[760,868,869,889]" degrees="41" direction="north" minutes="45" orientation="latitude" pageId="9" pageNumber="1200" precision="15" seconds="39" value="41.760834">41°45'39"N</geoCoordinate>
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,
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<geoCoordinate id="80935035EA086177473FF653FC34FCA8" box="[885,990,869,889]" degrees="34" direction="east" minutes="02" orientation="longitude" pageId="9" pageNumber="1200" precision="15" seconds="36" value="34.04333">34°02'36"E</geoCoordinate>
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<quantity id="225F9B17EA08617747A1F653FBD3FCA8" box="[1003,1081,869,889]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="9" pageNumber="1200" unit="m" value="1370.0">
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<elevation id="6E8AD1C1EA08617747A1F653FBD3FCA8" box="[1003,1081,869,889]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="9" pageNumber="1200" unit="m" value="1370.0">1370 m</elevation>
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</quantity>
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, calcareous slope,
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<date id="91191032EA0861774512F64AFE28FC41" box="[344,450,892,912]" pageId="9" pageNumber="1200" value="2009-04-10">
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<collectingDate id="815DE9DAEA0861774512F64AFE28FC41" box="[344,450,892,912]" pageId="9" pageNumber="1200" value="2009-04-10">10.IV.2009</collectingDate>
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</date>
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,
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<collectorName id="48525324EA0861774587F64AFDC1FC41" box="[461,555,892,912]" pageId="9" pageNumber="1200">V. Assing</collectorName>
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/
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<typeStatus id="3A1C8850EA0861774608F64AFD4BFC41" box="[578,673,892,912]" pageId="9" pageNumber="1200" type="holotype">Holotypus</typeStatus>
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Ƌ
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<taxonomicName id="22A74D71EA086177468EF64AFC9AFC41" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[708,880,892,912]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1200" phylum="Arthropoda" rank="species" species="hasanica" status="sp. nov.">
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<emphasis id="D7D3EAE0EA086177468EF64AFC9AFC41" box="[708,880,892,912]" italics="true" pageId="9" pageNumber="1200">Geostiba hasanica</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="CCE0579BEA0861774733F64AFC47FC41" box="[889,941,892,912]" pageId="9" pageNumber="1200" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<determinerName id="10C4DA36EA0861774795F64AFBD6FC41" box="[991,1084,892,912]" pageId="9" pageNumber="1200">V. Assing</determinerName>
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2009" (cAss)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="55CF3CAFEA0861774576F6A5FC4BFC76" box="[316,929,913,940]" collectingDate="2009-04-10" collectionCode="OOLL" collectorName="V. Assing" determinerName="V. Assing" elevation="1370" latitude="41.760834" location="Inebolu" longLatPrecision="19" longitude="34.04333" pageId="9" pageNumber="1200" specimenCount="17" specimenCount-female="17" stateProvince="Kastamonu" typeStatus="paratype">
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<typeStatus id="3A1C8850EA0861774576F6A5FE72FC76" box="[316,408,915,935]" pageId="9" pageNumber="1200" type="paratype">Paratypes</typeStatus>
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: 14ƋƋ,
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<specimenCount id="F3A1FD7BEA08617745A5F6A5FDF4FC7D" box="[495,542,915,940]" pageId="9" pageNumber="1200" type="female">17♀♀</specimenCount>
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||
: same data as holotype (cAss,
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<collectionCode id="83B6AE37EA0861774710F6A5FC71FC76" box="[858,923,915,935]" pageId="9" pageNumber="1200">OÖLL</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="55CF3CAFEA08617747E4F6A5FDE2FC6C" collectingDate="2009-04-10" collectionCode="OOLL" collectorName="Wunderle" determinerName="V. Assing" elevation="1370" latitude="41.760834" location="Inebolu" longLatPrecision="19" longitude="34.04333" pageId="9" pageNumber="1200" specimenCount="19" specimenCount-female="19" stateProvince="Kastamonu" typeStatus="paratype">
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10ƋƋ,
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<specimenCount id="F3A1FD7BEA08617747B0F6A5FBC2FC7D" box="[1018,1064,915,940]" pageId="9" pageNumber="1200" type="female">19♀♀</specimenCount>
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||
: same data, but leg.
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<collectorName id="48525324EA086177452BF69FFE51FC6C" box="[353,443,937,957]" pageId="9" pageNumber="1200">Wunderle</collectorName>
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(cWun)
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</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="E51836F2EA08617744F8F6F8FF01FB8F" blockId="9.[178,1085,973,1118]" pageId="9" pageNumber="1200">
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D e s c r i p t i o n: Body length
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<quantity id="225F9B17EA086177466BF6FBFD76FC32" box="[545,668,973,995]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.1" metricValueMin="2.3" pageId="9" pageNumber="1200" unit="mm" value="2.7" valueMax="3.1" valueMin="2.3">2.3-3.1 mm</quantity>
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. Habitus as in
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<figureCitation id="7D9C2A77EA0861774776F6FBFC60FC32" box="[828,906,973,995]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="9" pageNumber="1200">Fig. 16</figureCitation>
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. Coloration variable: head reddish brown to blackish; pronotum and elytra reddish to dark brown; abdomen blackish, occasionally with abdominal segments III-IV and VIII-X more or less distinctly paler brown; legs yellowish; antennae brown, with the basal 2-3 antennomeres paler.
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</paragraph>
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<paragraph id="E51836F2EA08617744F8F159FC5DFB13" blockId="9.[178,1084,1135,1218]" pageId="9" pageNumber="1200">
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Head 1.00-1.07 times as long as wide (
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||
<figureCitation id="7D9C2A77EA086177461FF159FD20FB54" box="[597,714,1135,1157]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="9" pageNumber="1200">Figs 17-18</figureCitation>
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); punctation extremely fine, barely noticeable; surface with very shallow, sometimes almost obsolete microreticulation. Eyes relatively small, 0.35-0.45 times as long as postocular region in dorsal view.
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</paragraph>
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<paragraph id="E51836F2EA08617744F8F1E5FF02FAF7" blockId="9.[178,1085,1235,1318]" pageId="9" pageNumber="1200">
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Pronotum with rather weakly pronounced sexual dimorphism (
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||
<figureCitation id="7D9C2A77EA086177470CF1E5FC51FB38" box="[838,955,1235,1257]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="9" pageNumber="1200">Figs 17-18</figureCitation>
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); punctation extremely fine, barely noticeable; microreticulation somewhat more distinct than that of head.
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</paragraph>
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<paragraph id="E51836F2EA08617744F8F001FE6FFA5B" blockId="9.[178,1086,1335,1418]" pageId="9" pageNumber="1200">
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||
Elytra with moderately pronounced sexual dimorphism, 0.50-0.55 times as long as pronotum (
|
||
<figureCitation id="7D9C2A77EA08617744B5F060FE9BFABD" box="[255,369,1366,1388]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="9" pageNumber="1200">Figs 17-18</figureCitation>
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); microsculpture very shallow, less pronounced than that of pronotum. Hind wings reduced.
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</paragraph>
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<paragraph id="E51836F2EA08617744F8F0ADFE66F9DC" blockId="9.[178,1085,1435,1549]" pageId="9" pageNumber="1200">
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Abdomen (
|
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<figureCitation id="7D9C2A77EA086177456DF0ADFE9EFA60" box="[295,372,1435,1457]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="9" pageNumber="1200">Fig. 19</figureCitation>
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) approximately as wide as elytra; punctation very fine and sparse; microreticulation distinct; posterior margin of tergite VII without palisade fringe; tergites IV and V without, VII with sexual dimorphism; posterior margin of tergite VIII weakly convex in both sexes.
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</paragraph>
|
||
<caption id="B1D8667AEA0B617444F8F010FDEEFAAF" ID-DOI="http://doi.org/10.5281/zenodo.5276573" ID-Zenodo-Dep="5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="10" pageNumber="1201" startId="10.[178,218,1318,1338]" targetBox="[184,1085,219,1296]" targetPageId="10">
|
||
<paragraph id="E51836F2EA0B617444F8F010FDEEFAAF" blockId="10.[178,1085,1318,1406]" pageId="10" pageNumber="1201">
|
||
<emphasis id="D7D3EAE0EA0B617444F8F010FEF3FAEB" bold="true" box="[178,281,1318,1338]" pageId="10" pageNumber="1201">Figs 16-22</emphasis>
|
||
:
|
||
<taxonomicName id="22A74D71EA0B61744563F010FE32FAEB" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[297,472,1318,1338]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1201" phylum="Arthropoda" rank="species" species="hasanica" status="sp. nov.">
|
||
<emphasis id="D7D3EAE0EA0B61744563F010FE32FAEB" box="[297,472,1318,1338]" italics="true" pageId="10" pageNumber="1201">Geostiba hasanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CCE0579BEA0B617445A8F010FDCEFAEB" box="[482,548,1318,1338]" pageId="10" pageNumber="1201" rank="species">nov.sp.</taxonomicNameLabel>
|
||
(16-17, 19-20: holotype): (
|
||
<emphasis id="D7D3EAE0EA0B61744765F010FCACFAEB" bold="true" box="[815,838,1318,1338]" pageId="10" pageNumber="1201">16</emphasis>
|
||
) male habitus; (
|
||
<emphasis id="D7D3EAE0EA0B617447A2F010FC15FAEB" bold="true" box="[1000,1023,1318,1338]" pageId="10" pageNumber="1201">17</emphasis>
|
||
) male forebody; (
|
||
<emphasis id="D7D3EAE0EA0B61744550F00BFEDBFA80" bold="true" box="[282,305,1341,1361]" pageId="10" pageNumber="1201">18</emphasis>
|
||
) female forebody; (
|
||
<emphasis id="D7D3EAE0EA0B617445BBF00BFDE2FA80" bold="true" box="[497,520,1341,1361]" pageId="10" pageNumber="1201">19</emphasis>
|
||
) male abdomen; (
|
||
<emphasis id="D7D3EAE0EA0B617446FDF00BFD24FA80" bold="true" box="[695,718,1341,1361]" pageId="10" pageNumber="1201">20</emphasis>
|
||
) male tergites VI-VIII in lateral view; (
|
||
<emphasis id="D7D3EAE0EA0B617444F3F065FF3AFAB6" bold="true" box="[185,208,1363,1383]" pageId="10" pageNumber="1201">21</emphasis>
|
||
) median lobe of aedeagus in lateral view; (
|
||
<emphasis id="D7D3EAE0EA0B61744625F065FD6CFAB6" bold="true" box="[623,646,1363,1383]" pageId="10" pageNumber="1201">22</emphasis>
|
||
) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E51836F2EA0B617544F8F0A2FE1EFEFF" blockId="10.[178,1085,1428,1603]" lastBlockId="11.[178,1085,219,302]" lastPageId="11" lastPageNumber="1202" pageId="10" pageNumber="1201">
|
||
Ƌ (with fully developed secondary sexual characters): pronotum 1.00-1.05 times as long as wide, posterior margin convex in the middle, lateral margins smoothly and weakly convex; elytra with short and rather weakly elevated sutural carinae in anterior half, postero-laterally with oblique impressions, punctation weakly to moderately granulose (
|
||
<figureCitation id="7D9C2A77EA0B617444F3F339FEE2F9F4" box="[185,264,1551,1573]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="10" pageNumber="1201">Fig. 17</figureCitation>
|
||
); abdominal tergites III and IV without distinct median tubercle, occasionally tergite III with very weak indication of such a tubercle (
|
||
<figureCitation id="7D9C2A77EA0B617446B6F31BFCA3F992" box="[764,841,1581,1603]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="10" pageNumber="1201">Fig. 18</figureCitation>
|
||
); process of tergite VII moderately long, slender and apically rounded in antero-dorsal view, broad-based and apically acute in lateral view (
|
||
<figureCitation id="7D9C2A77EA0A617545B3F5CCFDA3FEC1" box="[505,585,250,272]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="11" pageNumber="1202">Fig. 20</figureCitation>
|
||
); median lobe of aedeagus with broad daggershaped cristal process (
|
||
<figureCitation id="7D9C2A77EA0A617545D6F42EFE0DFEFF" box="[412,487,280,302]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="11" pageNumber="1202">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA0A617544F8F474FE6BFE40" blockId="11.[178,1085,319,401]" pageId="11" pageNumber="1202">
|
||
<specimenCount id="F3A1FD7BEA0A617544F8F474FF2AFE88" box="[178,192,322,345]" pageId="11" pageNumber="1202" type="female">♀</specimenCount>
|
||
: pronotum weakly transverse, posterior margin broadly convex; punctation of elytra not distinctly granulose (
|
||
<figureCitation id="7D9C2A77EA0A617545F9F46BFDEAFEA2" box="[435,512,349,371]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="11" pageNumber="1202">Fig. 18</figureCitation>
|
||
); tergites III, IV, and VII unmodified; spermatheca not distinctive (
|
||
<figureCitation id="7D9C2A77EA0A61754563F44DFE9FFE40" box="[297,373,379,401]" captionStart="Figs 16-22" captionStartId="10.[178,218,1318,1338]" captionTargetBox="[184,1085,219,1296]" captionTargetId="figure-143@10.[178,1085,219,1306]" captionTargetPageId="10" captionText="Figs 16-22: Geostiba hasanica nov.sp. (16-17, 19-20: holotype): (16) male habitus; (17) male forebody; (18) female forebody; (19) male abdomen; (20) male tergites VI-VIII in lateral view; (21) median lobe of aedeagus in lateral view; (22) spermatheca. Scale bars: 16: 1.0 mm; 17-19: 0.5 mm; 20: 0.2 mm; 21-22: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276573" httpUri="https://zenodo.org/record/5276573/files/figure.png" pageId="11" pageNumber="1202">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA0A617544F8F495FE9BFE07" blockId="11.[178,1085,418,470]" pageId="11" pageNumber="1202">
|
||
E t y m o l o g y: The name (adjective) is derived from the Hasan Dağı, where the
|
||
<typeStatus id="3A1C8850EA0A6175405BF495FBD7FE69" box="[1041,1085,419,440]" pageId="11" pageNumber="1202">type</typeStatus>
|
||
locality is situated.
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA0A617544F8F4D1FD54FC77" blockId="11.[178,1085,487,934]" pageId="11" pageNumber="1202">
|
||
C o m p a r a t i v e n o t e s: Using the key in
|
||
<bibRefCitation id="81364B03EA0A61754688F4D1FC83FE2C" author="ASSING V" box="[706,873,487,509]" pageId="11" pageNumber="1202" pagination="903 - 1006" refId="ref43550" refString="ASSING V. (2005 a): A revision of the species of Geostiba THOMSON and Tropimenelytron PACE of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 37 (2): 903 - 1006." type="journal article" year="2005">ASSING (2005a)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="22A74D71EA0A6175473CF4D1FC19FE2C" authorityName="Assing" authorityYear="2009" box="[886,1011,487,509]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="hasanica">
|
||
<emphasis id="D7D3EAE0EA0A6175473CF4D1FC19FE2C" box="[886,1011,487,509]" italics="true" pageId="11" pageNumber="1202">G. hasanica</emphasis>
|
||
</taxonomicName>
|
||
would key out at couplets 83-86, together with two species from central southern
|
||
<collectingCountry id="9DB07662EA0A617547F6F733FBF9FDCA" box="[956,1043,517,539]" name="Turkey" pageId="11" pageNumber="1202">Anatolia</collectingCountry>
|
||
(
|
||
<taxonomicName id="22A74D71EA0A6175406EF733FE62FDE8" authority="PACE 1983" authorityName="PACE" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="D7D3EAE0EA0A6175406EF733FEE6FDE8" italics="true" pageId="11" pageNumber="1202">G. simulans</emphasis>
|
||
PACE 1983
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="22A74D71EA0A617545DCF715FD41FDE8" authority="ASSING 2004" authorityName="ASSING" authorityYear="2004" box="[406,683,547,569]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="marasica">
|
||
<emphasis id="D7D3EAE0EA0A617545DCF715FDFCFDE8" box="[406,534,547,569]" italics="true" pageId="11" pageNumber="1202">G. marasica</emphasis>
|
||
ASSING 2004
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="22A74D71EA0A617546ADF715FBE8FDE8" authority="PACE 1983" authorityName="PACE" authorityYear="1983" box="[743,1026,547,569]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="arganthonia">
|
||
<emphasis id="D7D3EAE0EA0A617546ADF715FC6DFDE8" box="[743,903,547,569]" italics="true" pageId="11" pageNumber="1202">G. arganthonia</emphasis>
|
||
PACE 1983
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="2763F810EA0A617544F8F774FEEDFD89" box="[178,263,578,600]" country="Turkey" name="Istanbul" pageId="11" pageNumber="1202">Istanbul</collectingRegion>
|
||
. It is distinguished from all these species by the shape of the cristal process of the aedeagus, from
|
||
<taxonomicName id="22A74D71EA0A61754537F756FDF1FDA7" authorityName="PACE" authorityYear="1983" box="[381,539,608,630]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="arganthonia">
|
||
<emphasis id="D7D3EAE0EA0A61754537F756FDF1FDA7" box="[381,539,608,630]" italics="true" pageId="11" pageNumber="1202">G. arganthonia</emphasis>
|
||
</taxonomicName>
|
||
additionally by the sexual dimorphism of the pronotum (in
|
||
<taxonomicName id="22A74D71EA0A6175454CF749FE4EFD44" authorityName="PACE" authorityYear="1983" box="[262,420,639,661]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="arganthonia">
|
||
<emphasis id="D7D3EAE0EA0A6175454CF749FE4EFD44" box="[262,420,639,661]" italics="true" pageId="11" pageNumber="1202">G. arganthonia</emphasis>
|
||
</taxonomicName>
|
||
absent), as well as by the longer and more erect process of the male abdominal tergite VII, from
|
||
<taxonomicName id="22A74D71EA0A61754646F7ABFD6FFD62" authorityName="PACE" authorityYear="1983" box="[524,645,669,691]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="D7D3EAE0EA0A61754646F7ABFD6FFD62" box="[524,645,669,691]" italics="true" pageId="11" pageNumber="1202">G. simulans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="22A74D71EA0A617546F2F7ABFCDFFD62" authorityName="ASSING" authorityYear="2004" box="[696,821,669,691]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="marasica">
|
||
<emphasis id="D7D3EAE0EA0A617546F2F7ABFCDFFD62" box="[696,821,669,691]" italics="true" pageId="11" pageNumber="1202">G. marasica</emphasis>
|
||
</taxonomicName>
|
||
also by the modifications of the male elytra. For illustrations of
|
||
<taxonomicName id="22A74D71EA0A61754670F78DFD3CFD00" authorityName="PACE" authorityYear="1983" box="[570,726,699,721]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="arganthonia">
|
||
<emphasis id="D7D3EAE0EA0A61754670F78DFD3CFD00" box="[570,726,699,721]" italics="true" pageId="11" pageNumber="1202">G. arganthonia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="22A74D71EA0A617546AEF78DFCB4FD00" authorityName="PACE" authorityYear="1983" box="[740,862,699,721]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="D7D3EAE0EA0A617546AEF78DFCB4FD00" box="[740,862,699,721]" italics="true" pageId="11" pageNumber="1202">G. simulans</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="22A74D71EA0A617547D2F78DFBFCFD00" authorityName="ASSING" authorityYear="2004" box="[920,1046,699,721]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="marasica">
|
||
<emphasis id="D7D3EAE0EA0A617547D2F78DFBFCFD00" box="[920,1046,699,721]" italics="true" pageId="11" pageNumber="1202">G. marasica</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="81364B03EA0A617544F8F7EDFEABFD21" author="PACE R" box="[178,321,730,752]" pageId="11" pageNumber="1202" pagination="3 - 46" refId="ref43801" refString="PACE R. (1983 b): Specie del genere Geostiba THOMSON raccolte dal Dr. C. Besuchet e collaboratori in Marocco, nella Penisola Iberica e Balcanica, e nel Medio Oriente (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 90: 3 - 46." type="journal article" year="1983">PACE (1983b)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="81364B03EA0A61754530F7ECFDC8FD21" author="ASSING V" box="[378,546,730,752]" pageId="11" pageNumber="1202" pagination="615 - 638" refId="ref43470" refString="ASSING V. (2004 a): A revision of the Turkish species of Geostiba THOMSON. V. New species and additional records (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 36 (2): 615 - 638." type="journal article" year="2004">ASSING (2004a)</bibRefCitation>
|
||
. From the geographically closest consubgeners,
|
||
<taxonomicName id="22A74D71EA0A6175406EF7ECFEB8FCDF" authorityName="PACE" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="kastamonuensis">
|
||
<emphasis id="D7D3EAE0EA0A6175406EF7ECFEB8FCDF" italics="true" pageId="11" pageNumber="1202">G. kastamonuensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="22A74D71EA0A617545CDF7CEFDE5FCDF" authorityName="Assing" authorityYear="2009" box="[391,527,760,782]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="heliophila">
|
||
<emphasis id="D7D3EAE0EA0A617545CDF7CEFDE5FCDF" box="[391,527,760,782]" italics="true" pageId="11" pageNumber="1202">G. heliophila</emphasis>
|
||
</taxonomicName>
|
||
, the new species is readily distinguished by the shallower microreticulation of the head, the weakly pronounced sexual dimorphism of the male pronotum (posterior margin of the male pronotum convex in the middle), and by the shape of the cristal process of the aedeagus, from
|
||
<taxonomicName id="22A74D71EA0A61754696F665FC8DFCB8" authorityName="Assing" authorityYear="2009" box="[732,871,851,873]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="heliophila">
|
||
<emphasis id="D7D3EAE0EA0A61754696F665FC8DFCB8" box="[732,871,851,873]" italics="true" pageId="11" pageNumber="1202">G. heliophila</emphasis>
|
||
</taxonomicName>
|
||
additionally by the absence of distinct tubercles on the male abdominal tergites III and IV, and by the proximally more distinctly sclerotised spermatheca.
|
||
</paragraph>
|
||
<caption id="B1D8667AEA0A617544F8F0C1FCF3F9DA" ID-DOI="http://doi.org/10.5281/zenodo.5276575" ID-Zenodo-Dep="5276575" box="[178,793,1527,1547]" httpUri="https://zenodo.org/record/5276575/files/figure.png" pageId="11" pageNumber="1202" startId="11.[178,211,1527,1547]" targetBox="[178,1085,947,1515]" targetPageId="11">
|
||
<paragraph id="E51836F2EA0A617544F8F0C1FCF3F9DA" blockId="11.[178,793,1527,1547]" box="[178,793,1527,1547]" pageId="11" pageNumber="1202">
|
||
<emphasis id="D7D3EAE0EA0A617544F8F0C1FF1EF9DA" bold="true" box="[178,244,1527,1547]" pageId="11" pageNumber="1202">Fig. 23</emphasis>
|
||
: Type locality of
|
||
<taxonomicName id="22A74D71EA0A617545DCF0C1FDEEF9DA" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[406,516,1527,1547]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1202" phylum="Arthropoda" rank="species" species="hasanica" status="sp. nov.">
|
||
<emphasis id="D7D3EAE0EA0A617545DCF0C1FDEEF9DA" box="[406,516,1527,1547]" italics="true" pageId="11" pageNumber="1202">G. hasanica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CCE0579BEA0A61754640F0C1FDA6F9DA" box="[522,588,1527,1547]" pageId="11" pageNumber="1202" rank="species">nov.sp.</taxonomicNameLabel>
|
||
(photo: P. Wunderle).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E51836F2EA0A617544F8F314FBD6F984" blockId="11.[178,1085,1570,1621]" pageId="11" pageNumber="1202">
|
||
D i s t r i b u t i o n a n d b i o n o m i c s: Thespeciesisprobablyendemictothe Hasan Dağı to the northeast of
|
||
<collectingRegion id="2763F810EA0A617545B8F376FD82F984" box="[498,616,1600,1621]" country="Turkey" name="Kastamonu" pageId="11" pageNumber="1202">Kastamonu</collectingRegion>
|
||
,
|
||
<collectingRegion id="2763F810EA0A61754639F376FCA2F984" box="[627,840,1599,1621]" country="Turkey" name="Kastamonu" pageId="11" pageNumber="1202">Kastamonu province</collectingRegion>
|
||
, northern
|
||
<collectingCountry id="9DB07662EA0A617547FAF309FBE0F984" box="[944,1034,1599,1621]" name="Turkey" pageId="11" pageNumber="1202">Anatolia</collectingCountry>
|
||
. The
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA0D617244F8F5EDFD30FEDF" blockId="12.[178,1084,219,270]" pageId="12" pageNumber="1203">
|
||
type specimens were collected by turning stones, partly near snow, on a calcareous slope near a forest margin at an altitude of
|
||
<quantity id="225F9B17EA0D61724662F5CEFD9EFEDF" box="[552,628,248,270]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="12" pageNumber="1203" unit="m" value="1370.0">1370 m</quantity>
|
||
(
|
||
<figureCitation id="7D9C2A77EA0D617246C8F5CEFD25FEDF" box="[642,719,248,270]" captionStart="Fig" captionStartId="11.[178,211,1527,1547]" captionTargetBox="[178,1085,947,1515]" captionTargetId="figure-362@11.[178,1085,947,1515]" captionTargetPageId="11" captionText="Fig. 23: Type locality of G. hasanica nov.sp. (photo: P. Wunderle)." figureDoi="http://doi.org/10.5281/zenodo.5276575" httpUri="https://zenodo.org/record/5276575/files/figure.png" pageId="12" pageNumber="1203">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |