247 lines
33 KiB
XML
247 lines
33 KiB
XML
<document id="3E2C3E2F3A3B8EBC2303A6B835BFF03B" ID-DOI="10.5281/zenodo.5276563" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630012980284" checkinUser="carolina" docAuthor="Assing, V." docDate="2009" docId="6D0E87E4EA07617644F8F127FE5BFA7B" docLanguage="en" docName="LinzerbiolBeitr.2009.41.2.1191-1246.pdf" docOrigin="Linzer biologische Beiträge 41 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba (Tropogastrosipalia) heliophila Assing 2009, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1199" masterDocId="9137FF9CEA01617E444AF536FFEAFFD1" masterDocTitle="A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and a catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1246" masterPageNumber="1191" pageNumber="1197" updateTime="1699136907799" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="82410D00DFC6EC36C4FA9EB468A7730F">
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<mods:title id="16318B55FD2D2196A6F6D70EF1626F36">A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and a catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B72811DDF6A858D29EB1ECFCB7287E54" type="personal">
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<mods:role id="27CD19204C71314501F5CBF1D02ACF28">
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<mods:roleTerm id="846B7C655F245700D33D97EE9E746961">Author</mods:roleTerm>
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<mods:namePart id="59920B1FB07EBDF73B2482C56A9844A2">Assing, V.</mods:namePart>
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<mods:typeOfResource id="1C55B3CFF4225BA45F96D37D0B19BBD3">text</mods:typeOfResource>
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<mods:title id="6B9D02E48F424855D78C8F8D037FF9AE">Linzer biologische Beiträge</mods:title>
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</mods:titleInfo>
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<mods:part id="493D9A666D47D4CA1E47308E7A4FF0F1">
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<mods:date id="F55F5F9F8B10797CA8523EF6D6D613BA">2009</mods:date>
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<mods:detail id="E4E58E3E1079FE2CA2445F39745BB050" type="pubDate">
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<mods:number id="E9DCDA3021B2B804C6E67A9B135D3B1F">2009-12-18</mods:number>
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</mods:detail>
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<mods:detail id="2E95D36DA7B4D937878B51D2F15CE795" type="volume">
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<mods:number id="487FC8559D60E7714FD2260602B129C9">41</mods:number>
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<mods:detail id="E6D65ACA6D625C2C5F6F6E9542360331" type="issue">
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<mods:number id="586A82A49636023A9FF73C3307AB50C0">2</mods:number>
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<mods:start id="EE79076DC0447EAF966A0A1608CF5EA0">1191</mods:start>
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<mods:end id="338C647F78193542FA5BBBA9EE38F1D2">1246</mods:end>
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<mods:classification id="78794CB8A3F6F1A03AB8668D96549C18">journal article</mods:classification>
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<mods:identifier id="A1211E298BF4C676035DE5CA003FB5F5" type="DOI">10.5281/zenodo.5276563</mods:identifier>
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<mods:identifier id="829052667DD0E7C4CCAC08BAD8B9A771" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="B79981300AA0B6D1BBADB2A0C44AE9F2" type="Zenodo-Dep">5276563</mods:identifier>
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<treatment id="6D0E87E4EA07617644F8F127FE5BFA7B" LSID="urn:lsid:plazi:treatment:6D0E87E4EA07617644F8F127FE5BFA7B" httpUri="http://treatment.plazi.org/id/6D0E87E4EA07617644F8F127FE5BFA7B" lastPageId="8" lastPageNumber="1199" pageId="6" pageNumber="1197">
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<subSubSection id="ADBD6579EA07617844F8F127FCC8FBF6" box="[178,802,1041,1063]" pageId="6" pageNumber="1197" type="nomenclature">
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<paragraph id="E51836F2EA07617844F8F127FCC8FBF6" blockId="6.[178,802,1041,1063]" box="[178,802,1041,1063]" pageId="6" pageNumber="1197">
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<heading id="BE50819EEA07617844F8F127FCC8FBF6" box="[178,802,1041,1063]" fontSize="8" level="2" pageId="6" pageNumber="1197" reason="2">
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<emphasis id="D7D3EAE0EA07617844F8F127FD41FBF6" bold="true" box="[178,683,1041,1063]" pageId="6" pageNumber="1197">
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<taxonomicName id="22A74D71EA07617844F8F127FDB2FBF6" ID-CoL="6KCW8" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[178,600,1041,1063]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1197" phylum="Arthropoda" rank="species" species="heliophila" status="sp. nov." subGenus="Tropogastrosipalia">
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<emphasis id="D7D3EAE0EA07617844F8F127FEE7FBF6" bold="true" box="[178,269,1041,1063]" italics="true" pageId="6" pageNumber="1197">Geostiba</emphasis>
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(
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<emphasis id="D7D3EAE0EA0761784556F127FE0BFBF6" bold="true" box="[284,481,1041,1063]" italics="true" pageId="6" pageNumber="1197">Tropogastrosipalia</emphasis>
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)
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<emphasis id="D7D3EAE0EA07617845A5F127FDB2FBF6" bold="true" box="[495,600,1041,1063]" italics="true" pageId="6" pageNumber="1197">heliophila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CCE0579BEA0761784615F127FD41FBF6" box="[607,683,1041,1063]" pageId="6" pageNumber="1197" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="7D9C2A77EA07617846F2F127FCF1FBF6" box="[696,795,1041,1063]" captionStart-0="Figs 9-14" captionStart-1="Fig" captionStartId-0="7.[178,218,1397,1417]" captionStartId-1="9.[178,211,764,784]" captionTargetBox-0="[178,1085,219,1376]" captionTargetBox-1="[178,1085,219,753]" captionTargetId-0="figure-0@7.[178,1085,219,1393]" captionTargetId-1="figure-274@9.[178,1085,219,753]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." captionText-1="Fig. 15: Type locality of G. heliophila nov.sp. (photo: P. Wunderle)." figureDoi-0="http://doi.org/10.5281/zenodo.5276569" figureDoi-1="http://doi.org/10.5281/zenodo.5276571" httpUri-0="https://zenodo.org/record/5276569/files/figure.png" httpUri-1="https://zenodo.org/record/5276571/files/figure.png" pageId="6" pageNumber="1197">Figs 9-15</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ADBD6579EA07617644F8F10BFE5BFA7B" lastPageId="8" lastPageNumber="1199" pageId="6" pageNumber="1197" type="materials_examined">
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<paragraph id="E51836F2EA07617844F8F10BFDF0FB44" blockId="6.[178,1085,1084,1173]" pageId="6" pageNumber="1197">
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<materialsCitation id="55CF3CAFEA07617844F8F10BFEC7FBAE" collectingDate="2009-04-09" collectorName="V. Assing" determinerName="V. Assing" elevation="1090" latitude="41.70139" location="Kastamonu" longLatPrecision="19" longitude="33.47139" pageId="6" pageNumber="1197" specimenCount="2" stateProvince="Kastamonu" typeStatus="holotype">
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<typeStatus id="3A1C8850EA07617844F8F10BFEEDFB80" box="[178,263,1085,1105]" pageId="6" pageNumber="1197" type="holotype">Holotype</typeStatus>
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Ƌ: "TR [36] -
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<collectingRegion id="2763F810EA07617845D4F10BFDE2FB80" box="[414,520,1085,1105]" country="Turkey" name="Kastamonu" pageId="6" pageNumber="1197">Kastamonu</collectingRegion>
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,
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<quantity id="225F9B17EA076178465EF10BFDBBFB80" box="[532,593,1085,1105]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="1197" unit="km" value="40.0">40 km</quantity>
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NW
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<collectingRegion id="2763F810EA07617846C2F10BFD18FB80" box="[648,754,1085,1105]" country="Turkey" name="Kastamonu" pageId="6" pageNumber="1197">Kastamonu</collectingRegion>
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,
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<geoCoordinate id="80935035EA07617846B4F10BFC81FB80" box="[766,875,1085,1105]" degrees="41" direction="north" minutes="42" orientation="latitude" pageId="6" pageNumber="1197" precision="15" seconds="05" value="41.70139">41°42'05"N</geoCoordinate>
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,
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<geoCoordinate id="80935035EA0761784732F10BFC08FB80" box="[888,994,1085,1105]" degrees="33" direction="east" minutes="28" orientation="longitude" pageId="6" pageNumber="1197" precision="15" seconds="17" value="33.47139">33°28'17"E</geoCoordinate>
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,
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<quantity id="225F9B17EA07617847A5F10BFBD3FB80" box="[1007,1081,1085,1105]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="6" pageNumber="1197" unit="m" value="1090.0">
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<elevation id="6E8AD1C1EA07617847A5F10BFBD3FB80" box="[1007,1081,1085,1105]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="6" pageNumber="1197" unit="m" value="1090.0">1090 m</elevation>
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</quantity>
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, calcareous slope,
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<date id="91191032EA0761784512F162FE5DFBB9" box="[344,439,1108,1128]" pageId="6" pageNumber="1197" value="2009-04-09">
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<collectingDate id="815DE9DAEA0761784512F162FE5DFBB9" box="[344,439,1108,1128]" pageId="6" pageNumber="1197" value="2009-04-09">9.IV.2009</collectingDate>
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</date>
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,
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<collectorName id="48525324EA0761784588F162FDCAFBB9" box="[450,544,1108,1128]" pageId="6" pageNumber="1197">V. Assing</collectorName>
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/
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<typeStatus id="3A1C8850EA076178467DF162FD7CFBB9" box="[567,662,1108,1128]" pageId="6" pageNumber="1197" type="holotype">Holotypus</typeStatus>
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Ƌ
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<taxonomicName id="22A74D71EA07617846F0F162FC9AFBB9" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[698,880,1108,1128]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1197" phylum="Arthropoda" rank="species" species="heliophila" status="sp. nov.">
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<emphasis id="D7D3EAE0EA07617846F0F162FC9AFBB9" box="[698,880,1108,1128]" italics="true" pageId="6" pageNumber="1197">Geostiba heliophila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CCE0579BEA0761784732F162FC47FBB9" box="[888,941,1108,1128]" pageId="6" pageNumber="1197" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<determinerName id="10C4DA36EA0761784794F162FBD6FBB9" box="[990,1084,1108,1128]" pageId="6" pageNumber="1197">V. Assing</determinerName>
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2009" (cAss)
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</materialsCitation>
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||
.
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<materialsCitation id="55CF3CAFEA076178457DF15DFCAAFBAE" box="[311,832,1131,1155]" collectingDate="2009-04-09" collectionCode="OOLL" collectorName="V. Assing" determinerName="V. Assing" elevation="1090" latitude="41.70139" location="Kastamonu" longLatPrecision="19" longitude="33.47139" pageId="6" pageNumber="1197" specimenCount="10" specimenCount-female="10" stateProvince="Kastamonu" typeStatus="paratype">
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<typeStatus id="3A1C8850EA076178457DF15DFE79FBAE" box="[311,403,1131,1151]" pageId="6" pageNumber="1197" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="F3A1FD7BEA07617845D6F15DFE21FB52" box="[412,459,1131,1155]" pageId="6" pageNumber="1197" type="female">10♀♀</specimenCount>
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||
: same data as holotype (cAss,
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<collectionCode id="83B6AE37EA07617846B3F15DFCD0FBAE" box="[761,826,1131,1151]" pageId="6" pageNumber="1197">OÖLL</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="55CF3CAFEA0761784701F15DFDFCFB44" collectingDate="2009-04-09" collectorName="P. Wunderle" determinerName="V. Assing" elevation="1090" latitude="41.70139" location="Kastamonu" longLatPrecision="19" longitude="33.47139" pageId="6" pageNumber="1197" specimenCount="9" specimenCount-female="9" stateProvince="Kastamonu" typeStatus="paratype">
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||
2ƋƋ,
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<specimenCount id="F3A1FD7BEA07617847C3F15DFC46FB52" box="[905,940,1131,1155]" pageId="6" pageNumber="1197" type="female">9♀♀</specimenCount>
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: same data as holotype, but leg.
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<collectorName id="48525324EA076178451CF1B7FE23FB44" box="[342,457,1153,1173]" pageId="6" pageNumber="1197">P. Wunderle</collectorName>
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(cWun)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="E51836F2EA07617844F8F193FD3EFAC6" blockId="6.[178,1085,1189,1303]" pageId="6" pageNumber="1197">D e s c r i p t i o n: Body length 2.4-3.0 mm. Coloration variable: head and abdomen blackish, occasionally with abdominal segments III-IV and VIII-X more or less distinctly paler brown; pronotum and elytra reddish to dark brown; legs yellowish; antennae brown to dark brown, with the basal 2-3 antennomeres paler.</paragraph>
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<paragraph id="E51836F2EA07617844F8F01EFD98FAAA" blockId="6.[178,1085,1320,1403]" pageId="6" pageNumber="1197">
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Head approximately as long as wide (
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<figureCitation id="7D9C2A77EA076178461FF01EFD2AFAEF" box="[597,704,1320,1342]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="6" pageNumber="1197">Figs 9-10</figureCitation>
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); punctation extremely fine, barely noticeable; surface with shallow microreticulation. Eyes approximately half as long as postocular region in dorsal view, or smaller.
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</paragraph>
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<paragraph id="E51836F2EA07617844F8F0BAFBC7FA11" blockId="6.[178,1084,1420,1472]" pageId="6" pageNumber="1197">
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Pronotum with pronounced sexual dimorphism (
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<figureCitation id="7D9C2A77EA07617846E6F0BAFCFFFA73" box="[684,789,1420,1442]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="6" pageNumber="1197">Figs 9-10</figureCitation>
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); punctation extremely fine, barely noticeable; microreticulation similar to that of head or slightly more pronounced.
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</paragraph>
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<paragraph id="E51836F2EA07617844F8F0E7FE1EF9F4" blockId="6.[178,1085,1489,1573]" pageId="6" pageNumber="1197">
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Elytra with moderately pronounced sexual dimorphism, 0.42-0.50 times as long as pronotum (
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<figureCitation id="7D9C2A77EA076178454BF0C6FE82F9D7" box="[257,360,1520,1542]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="6" pageNumber="1197">Figs 9-10</figureCitation>
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); microsculpture very shallow, less pronounced than that of head and pronotum. Hind wings reduced.
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</paragraph>
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<caption id="B1D8667AEA06617944F8F043FC33FA67" ID-DOI="http://doi.org/10.5281/zenodo.5276569" ID-Zenodo-Dep="5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="7" pageNumber="1198" startId="7.[178,218,1397,1417]" targetBox="[178,1085,219,1376]" targetPageId="7">
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||
<paragraph id="E51836F2EA06617944F8F043FC33FA67" blockId="7.[178,1084,1397,1462]" pageId="7" pageNumber="1198">
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||
<emphasis id="D7D3EAE0EA06617944F8F043FEE1FA58" bold="true" box="[178,267,1397,1417]" pageId="7" pageNumber="1198">Figs 9-14</emphasis>
|
||
:
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||
<taxonomicName id="22A74D71EA0661794553F043FE24FA58" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[281,462,1397,1417]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1198" phylum="Arthropoda" rank="species" species="heliophila" status="sp. nov.">
|
||
<emphasis id="D7D3EAE0EA0661794553F043FE24FA58" box="[281,462,1397,1417]" italics="true" pageId="7" pageNumber="1198">Geostiba heliophila</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="CCE0579BEA066179459CF043FDF3FA58" box="[470,537,1397,1417]" pageId="7" pageNumber="1198" rank="species">nov.sp.</taxonomicNameLabel>
|
||
(9, 11-13: holotype): (
|
||
<emphasis id="D7D3EAE0EA06617946A5F043FD10FA58" bold="true" box="[751,762,1397,1417]" pageId="7" pageNumber="1198">9</emphasis>
|
||
) male forebody; (
|
||
<emphasis id="D7D3EAE0EA06617947EFF043FC56FA58" bold="true" box="[933,956,1397,1417]" pageId="7" pageNumber="1198">10</emphasis>
|
||
) female forebody; (
|
||
<emphasis id="D7D3EAE0EA06617944B9F0BAFEE0FA71" bold="true" box="[243,266,1420,1440]" pageId="7" pageNumber="1198">11</emphasis>
|
||
) male abdomen; (
|
||
<emphasis id="D7D3EAE0EA06617945FBF0BAFE22FA71" bold="true" box="[433,456,1420,1440]" pageId="7" pageNumber="1198">12</emphasis>
|
||
) male tergites VI-VIII in lateral view; (
|
||
<emphasis id="D7D3EAE0EA066179477FF0BAFCA6FA71" bold="true" box="[821,844,1420,1440]" pageId="7" pageNumber="1198">13</emphasis>
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||
) median lobe of aedeagus in lateral view; (
|
||
<emphasis id="D7D3EAE0EA066179450DF094FEB4FA67" bold="true" box="[327,350,1442,1462]" pageId="7" pageNumber="1198">14</emphasis>
|
||
) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E51836F2EA09617644F8F5EAFDBFFE9F" blockId="8.[178,1085,220,334]" pageId="8" pageNumber="1199">
|
||
Abdomen (
|
||
<figureCitation id="7D9C2A77EA096176456DF5EAFE99FF23" box="[295,371,220,242]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 11</figureCitation>
|
||
) approximately as wide as elytra; punctation very fine and sparse; microreticulation distinct, but rather shallow; posterior margin of tergite VII without palisade fringe; tergites IV, V, and VII with sexual dimorphism; posterior margin of tergite VIII weakly convex in both sexes.
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA09617644F8F468FF3BFD9D" blockId="8.[178,1085,350,588]" pageId="8" pageNumber="1199">
|
||
Ƌ (with fully developed secondary sexual characters): pronotum elongated, of oblong ovoid shape, produced posteriorly, approximately 1.15 times as long as wide, lateral margins not sinuate posteriorly, posterior margin weakly and narrowly concave in the middle; elytra with short and rather weakly elevated sutural carinae in anterior half, postero-laterally with oblique impressions, punctation distinctly granulose (
|
||
<figureCitation id="7D9C2A77EA09617647A3F4ECFBDAFE21" box="[1001,1072,474,496]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 9</figureCitation>
|
||
); abdominal tergites III and IV with median tubercle (
|
||
<figureCitation id="7D9C2A77EA096176468EF4CFFCE4FDDE" box="[708,782,505,527]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 11</figureCitation>
|
||
); process of tergite VII rather short and not very slender in lateral view (
|
||
<figureCitation id="7D9C2A77EA0961764625F721FD56FDFC" box="[623,700,535,557]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 12</figureCitation>
|
||
); median lobe of aedeagus as in
|
||
<figureCitation id="7D9C2A77EA096176405EF721FF24FD9D" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 13</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA09617644F8F757FCD4FD1E" blockId="8.[178,1085,605,719]" pageId="8" pageNumber="1199">
|
||
<specimenCount id="F3A1FD7BEA09617644F8F757FF2AFDA9" box="[178,192,609,632]" pageId="8" pageNumber="1199" type="female">♀</specimenCount>
|
||
: pronotum of shortly ovoid shape, weakly transverse or, at most, approximately as long as wide, posterior margin truncate or weakly convex; elytra with, at most, weakly granulose punctation (
|
||
<figureCitation id="7D9C2A77EA09617645D0F7ACFE02FD61" box="[410,488,666,688]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 10</figureCitation>
|
||
); tergites III, IV, and VII unmodified; spermatheca with proximal portion of capsule conspicuously transparent (
|
||
<figureCitation id="7D9C2A77EA09617646ACF78FFCDBFD1E" box="[742,817,697,719]" captionStart="Figs 9-14" captionStartId="7.[178,218,1397,1417]" captionTargetBox="[178,1085,219,1376]" captionTargetId="figure-0@7.[178,1085,219,1393]" captionTargetPageId="7" captionText="Figs 9-14: Geostiba heliophila nov.sp. (9, 11-13: holotype): (9) male forebody; (10) female forebody; (11) male abdomen; (12) male tergites VI-VIII in lateral view; (13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 9-11: 0.5 mm; 12: 0.2 mm; 13-14: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276569" httpUri="https://zenodo.org/record/5276569/files/figure.png" pageId="8" pageNumber="1199">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA09617644F8F7D6FDDCFCC5" blockId="8.[178,1085,736,788]" pageId="8" pageNumber="1199">
|
||
E t y m o l o g y: The name (adjective) refers to the fact that, surprisingly, the
|
||
<typeStatus id="3A1C8850EA096176405BF7D6FBD7FD24" box="[1041,1085,736,757]" pageId="8" pageNumber="1199">type</typeStatus>
|
||
material was collected on a SW-slope.
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA09617644F8F613FF15FB38" blockId="8.[178,1085,805,1257]" pageId="8" pageNumber="1199">
|
||
C o m p a r a t i v e n o t e s: UsingthekeyinASSING (2005a),
|
||
<taxonomicName id="22A74D71EA096176473AF613FC1DFCEA" authorityName="Assing" authorityYear="2009" box="[880,1015,805,827]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="heliophila">
|
||
<emphasis id="D7D3EAE0EA096176473AF613FC1DFCEA" box="[880,1015,805,827]" italics="true" pageId="8" pageNumber="1199">G. heliophila</emphasis>
|
||
</taxonomicName>
|
||
would key out at couplet 68, together with
|
||
<taxonomicName id="22A74D71EA096176466AF672FC6DFC8B" authority="ASSING 2000" authorityName="ASSING" authorityYear="2000" box="[544,903,836,858]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="brachati">
|
||
<emphasis id="D7D3EAE0EA096176466AF672FD7EFC8B" box="[544,660,836,858]" italics="true" pageId="8" pageNumber="1199">G. brachati</emphasis>
|
||
ASSING 2000 (
|
||
<collectingRegion id="2763F810EA0961764767F672FC6AFC8B" box="[813,896,836,858]" country="Turkey" name="Antalya" pageId="8" pageNumber="1199">Antalya</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="22A74D71EA09617647F2F672FE66FCA8" authority="ASSING 2001" authorityName="ASSING" authorityYear="2001" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="bitlisensis">
|
||
<emphasis id="D7D3EAE0EA09617647F2F672FBD6FC8B" box="[952,1084,836,858]" italics="true" pageId="8" pageNumber="1199">G. bitlisensis</emphasis>
|
||
ASSING 2001 (
|
||
<collectingRegion id="2763F810EA0961764503F655FE6FFCA8" box="[329,389,867,889]" country="Turkey" name="Bitlis" pageId="8" pageNumber="1199">Bitlis</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
. It is distinguished from both species by the more slender habitus and smaller average size, the slightly more slender antennae, the shape of the male pronotum (more oblong, posterior margin more distinctly concave in the middle), the shape of the cristal process of the aedeagus, and by the proximally completely transparent spermatheca. In addition, it is separated from
|
||
<taxonomicName id="22A74D71EA096176477CF6EBFC59FC22" authorityName="ASSING" authorityYear="2000" box="[822,947,989,1011]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="brachati">
|
||
<emphasis id="D7D3EAE0EA096176477CF6EBFC59FC22" box="[822,947,989,1011]" italics="true" pageId="8" pageNumber="1199">G. brachati</emphasis>
|
||
</taxonomicName>
|
||
by the less pronounced microsculpture of the forebody and by the shorter, stouter, and less erect spine-like process of the male abdominal tergite VII. From
|
||
<taxonomicName id="22A74D71EA0961764700F12DFBFBFBE0" authorityName="PACE" authorityYear="1983" box="[842,1041,1051,1073]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="kastamonuensis">
|
||
<emphasis id="D7D3EAE0EA0961764700F12DFBFBFBE0" box="[842,1041,1051,1073]" italics="true" pageId="8" pageNumber="1199">G. kastamonuensis</emphasis>
|
||
</taxonomicName>
|
||
, its geographically closest consubgener, it is separated by smaller average size and more slender habitus, the more slender and more oblong male pronotum, the presence of tubercles on the male abdominal tergites III and IV, as well as by the shape of the process of the male abdominal tergite VIII and of the cristal process of the aedeagus. For illustrations of
|
||
<taxonomicName id="22A74D71EA0961764518F183FE20FB1A" authorityName="ASSING" authorityYear="2000" box="[338,458,1205,1227]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="brachati">
|
||
<emphasis id="D7D3EAE0EA0961764518F183FE20FB1A" box="[338,458,1205,1227]" italics="true" pageId="8" pageNumber="1199">G. brachati</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="22A74D71EA0961764591F183FD88FB1A" authorityName="ASSING" authorityYear="2001" box="[475,610,1205,1227]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="bitlisensis">
|
||
<emphasis id="D7D3EAE0EA0961764591F183FD88FB1A" box="[475,610,1205,1227]" italics="true" pageId="8" pageNumber="1199">G. bitlisensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="22A74D71EA09617646EBF183FC8EFB1A" authorityName="PACE" authorityYear="1983" box="[673,868,1205,1227]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="species" species="kastamonuensis">
|
||
<emphasis id="D7D3EAE0EA09617646EBF183FC8EFB1A" box="[673,868,1205,1227]" italics="true" pageId="8" pageNumber="1199">G. kastamonuensis</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="81364B03EA09617647D2F183FBD2FB1A" author="ASSING V" box="[920,1080,1205,1227]" pageId="8" pageNumber="1199" pagination="1033 - 1042" refId="ref43250" refString="ASSING V. (2000 b): The Turkish species of Geostiba s. str. THOMSON 1858 (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 32 (2) 1033 - 1042." type="journal article" year="2000">ASSING (2000b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="81364B03EA09617644F8F1E5FF1EFB38" author="ASSING V" box="[178,244,1235,1257]" pageId="8" pageNumber="1199" pagination="137 - 185" refId="ref43326" refString="ASSING V. (2001 a): A revision of the Turkish species of Geostiba THOMSON 1858 and Tropimenelytron PACE 1983 (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 33 (1): 137 - 185." type="journal article" year="2001">2001a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E51836F2EA09617644F8F1CDFE5BFA7B" blockId="8.[178,1085,1274,1450]" pageId="8" pageNumber="1199">
|
||
D i s t r i b u t i o n a n d b i o n o m i c s: As can be inferred from the restricted distributions of other Turkish representatives of the subgenus
|
||
<taxonomicName id="22A74D71EA0961764703F02FFBE6FAFE" box="[841,1036,1305,1327]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1199" phylum="Arthropoda" rank="subGenus" subGenus="Tropogastrosipalia">
|
||
<emphasis id="D7D3EAE0EA0961764703F02FFBE6FAFE" box="[841,1036,1305,1327]" italics="true" pageId="8" pageNumber="1199">Tropogastrosipalia</emphasis>
|
||
</taxonomicName>
|
||
, the species is probably endemic to the Karyatağı Dağı and possibly also adjacent mountain ranges to the northwest of
|
||
<collectingRegion id="2763F810EA096176458BF061FDDCFABD" box="[449,566,1367,1388]" country="Turkey" name="Kastamonu" pageId="8" pageNumber="1199">Kastamonu</collectingRegion>
|
||
,
|
||
<collectingRegion id="2763F810EA096176460BF061FCFCFABD" box="[577,790,1366,1388]" country="Turkey" name="Kastamonu" pageId="8" pageNumber="1199">Kastamonu province</collectingRegion>
|
||
, northern
|
||
<collectingCountry id="9DB07662EA0961764734F060FC32FABD" box="[894,984,1366,1388]" name="Turkey" pageId="8" pageNumber="1199">Anatolia</collectingCountry>
|
||
. The
|
||
<typeStatus id="3A1C8850EA096176405BF061FBD7FABD" box="[1041,1085,1367,1388]" pageId="8" pageNumber="1199">type</typeStatus>
|
||
specimens were collected by turning stones on a grassy calcareous SW-slope at an altitude of
|
||
<quantity id="225F9B17EA09617644B5F0A2FEA1FA78" box="[255,331,1428,1450]" metricMagnitude="3" metricUnit="m" metricValue="1.09" pageId="8" pageNumber="1199" unit="m" value="1090.0">1090 m</quantity>
|
||
(
|
||
<figureCitation id="7D9C2A77EA0961764513F0A2FE4FFA7B" box="[345,421,1428,1450]" captionStart="Fig" captionStartId="9.[178,211,764,784]" captionTargetBox="[178,1085,219,753]" captionTargetId="figure-274@9.[178,1085,219,753]" captionTargetPageId="9" captionText="Fig. 15: Type locality of G. heliophila nov.sp. (photo: P. Wunderle)." figureDoi="http://doi.org/10.5281/zenodo.5276571" httpUri="https://zenodo.org/record/5276571/files/figure.png" pageId="8" pageNumber="1199">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |