treatments-xml/data/55/0B/87/550B87C3C16FFFB4FF1F76F9FC460008.xml
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<document id="EEFED96221F0B43D8BE5EC5EBC5D20F4" ID-DOI="10.5281/zenodo.4651080" ID-ISSN="1638-9395" ID-Zenodo-Dep="4651080" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617195131926" checkinUser="felipe" docAuthor="Bianucci, Giovanni, Lambert, Olivier &amp; Post, Klaas" docDate="2007" docId="550B87C3C16FFFB4FF1F76F9FC460008" docLanguage="en" docName="Geodiversitas.29.4.561-618.pdf" docOrigin="Geodiversitas 29 (4)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Khoikhoicetus agulhasis Bianucci &amp; Lambert &amp; Post 2007, n. sp." docType="treatment" docUuid="34B13052-445A-442F-A1F0-5474DD717091" docUuidSource="ZooBank" docVersion="5" lastPageNumber="578" masterDocId="A932FFBBC17EFFA5FFC47277FFBE056E" masterDocTitle="A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa" masterLastPageNumber="618" masterPageNumber="561" pageNumber="578" updateTime="1698897450673" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="BD284093AC847B4416378A062FA01C0C">A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa</mods:title>
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<mods:name id="9C3C48A82BFE73A60DDC6FA47B5ABB22" type="personal">
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<mods:namePart id="56519F83C577BA9DE9E7E8326A1187A8">Bianucci, Giovanni</mods:namePart>
<mods:affiliation id="0F7866B3F5FEF1A8AE7F309CA0D58E8D">Dipartimento di Scienze della Terra, Università di Pisa, via S. Maria, 53, I- 56126 Pisa (Italy) bianucci @ dst. unipi. it</mods:affiliation>
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<mods:namePart id="BD3F3EB28661E3F6D269C6513E769D32">Lambert, Olivier</mods:namePart>
<mods:affiliation id="713EDAA12EDCC0780BC90471A60CB00E">Institut royal des Sciences naturelles de Belgique, Département de Paléontologie, rue Vautier, 29, B- 1000 Brussels (Belgium) olivier. lambert @ naturalsciences. be</mods:affiliation>
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<mods:namePart id="9D7C28514B6E7C068B3C4B3E42C4D254">Post, Klaas</mods:namePart>
<mods:affiliation id="7D1F3A196BE33628712EF04D0E887484">Natuurhistorisch Museum Rotterdam, P. O. Box 23452, NL- 3001 Rotterdam (The Netherlands) klaas @ fiskano. nl</mods:affiliation>
<mods:nameIdentifier id="33C025DE759E87563160842119886E70" type="email">klaas@fiskano.nl</mods:nameIdentifier>
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<mods:title id="DE8CFAC9B63701EB75D18A8D35C6C4B2">Geodiversitas</mods:title>
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<mods:date id="216A5351986D835546D0EE18690A9CAE">2007</mods:date>
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<subSubSection id="95B8655EC16FFFB4FF1F76F9FDA301C7" box="[219,541,1166,1193]" pageId="17" pageNumber="578" type="nomenclature">
<paragraph id="DD1D36D5C16FFFB4FF1F76F9FDA301C7" blockId="17.[219,541,1166,1193]" box="[219,541,1166,1193]" pageId="17" pageNumber="578">
<heading id="865581B9C16FFFB4FF1F76F9FDA301C7" box="[219,541,1166,1193]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="578" reason="2">
<taxonomicName id="1AA24D56C16FFFB4FF1F76F9FE6201C7" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[219,476,1166,1193]" class="Mammalia" family="Ziphiidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="species" species="agulhasis" status="sp. nov.">
<emphasis id="EFD6EAC7C16FFFB4FF1F76F9FE6201C7" bold="true" box="[219,476,1166,1193]" italics="true" pageId="17" pageNumber="578">Khoikhoicetus agulhasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC16FFFB4FE2676F8FDA301C7" box="[482,541,1167,1193]" pageId="17" pageNumber="578" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC16FFFB4FFB576B1FEBD007C" pageId="17" pageNumber="578" type="materials_examined">
<paragraph id="DD1D36D5C16FFFB4FFB576B1FEBD007C" blockId="17.[113,647,1222,1299]" pageId="17" pageNumber="578">
<materialsCitation id="6DCA3C88C16FFFB4FFB576B1FEBD007C" collectionCode="SAM" pageId="17" pageNumber="578" specimenCode="PQ 2678" specimenCount="1" typeStatus="holotype">
<typeStatus id="02198877C16FFFB4FFB576B1FF6501B2" box="[113,219,1222,1245]" pageId="17" pageNumber="578" type="holotype">HOLOTYPE</typeStatus>
. —
<collectionCode id="BBB3AE10C16FFFB4FECC76B1FE8501B0" box="[264,315,1222,1246]" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="17" pageNumber="578">SAM</collectionCode>
<specimenCode id="8D049EAEC16FFFB4FE8776B1FE1C01B0" box="[323,418,1222,1246]" collectionCode="SAM" country="South Africa" httpUri="http://grbio.org/cool/mhew-9czz" name="South African Museum" pageId="17" pageNumber="578">PQ 2678</specimenCode>
, partial skull including most of the rostrum, the anterior part of the cranium, and the vertex.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC16FFFB4FFB5775EFD8100FE" pageId="17" pageNumber="578" type="etymology">
<paragraph id="DD1D36D5C16FFFB4FFB5775EFD8100FE" blockId="17.[113,648,1321,1424]" pageId="17" pageNumber="578">
ETYMOLOGY. — The
<typeStatus id="02198877C16FFFB4FE84775EFE28002F" box="[320,406,1321,1345]" pageId="17" pageNumber="578" type="holotype">holotype</typeStatus>
was found offshore from
<collectingRegion id="1F66F837C16FFFB4FFB57734FF1F0035" box="[113,161,1347,1371]" country="South Africa" name="Western Cape" pageId="17" pageNumber="578">Cape</collectingRegion>
Agulhas, a locality on the south coast of
<collectingCountry id="A5B57645C16FFFB4FDC97734FD3A0035" box="[525,644,1347,1371]" name="South Africa" pageId="17" pageNumber="578">South Africa</collectingCountry>
. The oceanic current running along the south-east coast of
<collectingCountry id="A5B57645C16FFFB4FF4F770FFEB800FE" box="[139,262,1400,1424]" name="South Africa" pageId="17" pageNumber="578">South Africa</collectingCountry>
is also named Agulhas Current.
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC16FFFB4FFB577D0FDD100B7" pageId="17" pageNumber="578" type="materials_examined">
<paragraph id="DD1D36D5C16FFFB4FFB577D0FDD100B7" blockId="17.[113,646,1446,1497]" pageId="17" pageNumber="578">
<collectionCode id="BBB3AE10C16FFFB4FFB577D0FF4500D3" box="[113,251,1447,1470]" pageId="17" pageNumber="578">TYPE LOCALITY</collectionCode>
. — No exact locality.Trawled west of
<collectingRegion id="1F66F837C16FFFB4FD9177D1FD3800D0" box="[597,646,1446,1470]" country="South Africa" name="Western Cape" pageId="17" pageNumber="578">Cape</collectingRegion>
Agulhas, south coast of
<collectingCountry id="A5B57645C16FFFB4FE9D77B6FE6900B7" box="[345,471,1473,1497]" name="South Africa" pageId="17" pageNumber="578">South Africa</collectingCountry>
, Indian Ocean.
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC16FFFB4FFB57798FBB90452" pageId="17" pageNumber="578" type="diagnosis">
<paragraph id="DD1D36D5C16FFFB4FFB57798FBB90452" blockId="17.[113,647,1519,1699]" lastBlockId="17.[677,1210,215,316]" pageId="17" pageNumber="578">
DIAGNOSIS. —
<taxonomicName id="1AA24D56C16FFFB4FECA7798FE4F0369" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[270,497,1519,1543]" class="Mammalia" family="Ziphiidae" genus="Khoikhoicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="species" species="agulhasis" status="n. gen., n. sp.">
<emphasis id="EFD6EAC7C16FFFB4FECA7798FE4F0369" box="[270,497,1519,1543]" italics="true" pageId="17" pageNumber="578">Khoikhoicetus agulhasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC16FFFB4FE3D7798FD390368" box="[505,647,1519,1543]" pageId="17" pageNumber="578" rank="species">n. gen., n. sp.</taxonomicNameLabel>
differs from all the other
<taxonomicName id="1AA24D56C16FFFB4FEAB747EFDA3034F" authorityName="Gray" authorityYear="1866" box="[367,541,1545,1569]" class="Mammalia" family="Ziphiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="subFamily" subFamily="Hyperoodontinae">Hyperoodontinae</taxonomicName>
in lacking the posterolaterally directed premaxillary crest. It differs from
<taxonomicName id="1AA24D56C16FFFB4FF6C744BFEAB033A" authorityName="Moore" authorityYear="1968" box="[168,277,1596,1620]" class="Mammalia" family="Hyperoodontidae" genus="Indopacetus" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C16FFFB4FF6C744BFEAB033A" box="[168,277,1596,1620]" italics="true" pageId="17" pageNumber="578">Indopacetus</emphasis>
</taxonomicName>
in: smaller size; narrow rostrum base; narrower premaxillary crests and nasals longer than the frontals on the vertex. It differs from
<taxonomicName id="1AA24D56C16FFFB4FDCD7407FD3803E6" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[521,646,1648,1672]" class="Mammalia" family="Ziphiidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C16FFFB4FDCD7407FD3803E6" box="[521,646,1648,1672]" italics="true" pageId="17" pageNumber="578">Africanacetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC16FFFB4FFB574FCFF0803CD" box="[113,182,1675,1699]" pageId="17" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
,
<taxonomicName id="1AA24D56C16FFFB4FF7A74FDFE9203CC" authorityName="Lacepede" authorityYear="1804" box="[190,300,1674,1698]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C16FFFB4FF7A74FDFE9203CC" box="[190,300,1674,1698]" italics="true" pageId="17" pageNumber="578">Hyperoodon</emphasis>
</taxonomicName>
,
<taxonomicName id="1AA24D56C16FFFB4FEF274FDFE3A03CC" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[310,388,1674,1698]" class="Mammalia" family="Ziphiidae" genus="Ihlengesi" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C16FFFB4FEF274FDFE3A03CC" box="[310,388,1674,1698]" italics="true" pageId="17" pageNumber="578">Ihlengesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC16FFFB4FE4D74FCFE7203CD" box="[393,460,1675,1699]" pageId="17" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
, and
<taxonomicName id="1AA24D56C16FFFB4FE3874FDFDD603CC" authorityName="Gervais" authorityYear="1850" box="[508,616,1674,1698]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C16FFFB4FE3874FDFDD603CC" box="[508,616,1674,1698]" italics="true" pageId="17" pageNumber="578">Mesoplodon</emphasis>
</taxonomicName>
in: shorter intrusion of the nasal in the premaxillary crest; premaxillary crest not overhanging the premaxillary sac fossa. The space between the premaxillary crests is wider than in
<taxonomicName id="1AA24D56C16FFFB4FD357353FCDF0452" authorityName="Lacepede" authorityYear="1804" box="[753,865,292,316]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C16FFFB4FD357353FCDF0452" box="[753,865,292,316]" italics="true" pageId="17" pageNumber="578">Hyperoodon</emphasis>
</taxonomicName>
and
<taxonomicName id="1AA24D56C16FFFB4FC577353FBBF0452" authorityName="Gervais" authorityYear="1850" box="[915,1025,292,316]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus">
<emphasis id="EFD6EAC7C16FFFB4FC577353FBBF0452" box="[915,1025,292,316]" italics="true" pageId="17" pageNumber="578">Mesoplodon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="95B8655EC16FFFB4FD617320FC460008" pageId="17" pageNumber="578" type="description">
<paragraph id="DD1D36D5C16FFFB4FD617320FB8C041C" blockId="17.[677,1210,343,1382]" box="[677,1074,343,370]" pageId="17" pageNumber="578">
DESCRIPTION (
<figureCitation id="45992A50C16FFFB4FC81732FFC25041C" box="[837,923,344,370]" captionStart="FIG" captionStartId="18.[113,124,1657,1674]" captionTargetBox="[134,1185,225,1632]" captionTargetId="figure-22@18.[56,1268,217,1636]" captionTargetPageId="18" captionText="FIG. 13. — Skull of Khoikhoicetus agulhasis n. gen., n. sp. (SAM PQ 2678, holotype): A, dorsal view; B, lateral view; C, ventral view. Scale bar: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.4651105" httpUri="https://zenodo.org/record/4651105/files/figure.png" pageId="17" pageNumber="578">FIGS 13</figureCitation>
; 14;
<tableCitation id="9020036EC16FFFB4FC147320FB94041C" box="[976,1066,343,370]" captionStart="TABLE" captionStartId="52.[113,122,1081,1098]" captionText="TABLE 4. — Measurements (in mm) of the skulls of Hyperoodontinae from the Neogene of South Africa,Khoikhoicetus agulhasis n. gen., n. sp., Ihlengesi saldanhae n. gen., n. sp., Africanacetus ceratopsis n. gen., n. sp., and Mesoplodon slangkopi n. sp. Abbreviations: e, estimate; +, nearly complete; -, no data." pageId="17" pageNumber="578">TABLE 4</tableCitation>
)
</paragraph>
<paragraph id="DD1D36D5C16FFFB4FD617301FB2E07C3" blockId="17.[677,1210,343,1382]" pageId="17" pageNumber="578">
On the only known skull, the rostrum was probably relatively short, not much longer than the preserved length. It is somewhat wider than high along its anterior half and as wide as high posteriorly. The mesorostral groove is filled by the vomer, with a median suture on the posterior half. The ventral margin of the alveolar groove, lacking marks of alveoli, extends to the antorbital notch; contrary to
<taxonomicName id="1AA24D56C16FFFB4FBCE7023FB0C0700" authorityName="Haast" authorityYear="1876" box="[1034,1202,596,622]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="species" species="grayi">
<emphasis id="EFD6EAC7C16FFFB4FBCE7023FB0C0700" box="[1034,1202,596,622]" italics="true" pageId="17" pageNumber="578">Mesoplodon grayi</emphasis>
</taxonomicName>
,
<taxonomicName id="1AA24D56C16FFFB4FD617003FCB307E0" baseAuthorityName="Gray" baseAuthorityYear="1871" box="[677,781,628,654]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="species" species="hectori">
<emphasis id="EFD6EAC7C16FFFB4FD617003FCB307E0" box="[677,781,628,654]" italics="true" pageId="17" pageNumber="578">M. hectori</emphasis>
</taxonomicName>
,
<taxonomicName id="1AA24D56C16FFFB4FCDF7003FC3007E0" baseAuthorityName="Gray" baseAuthorityYear="1865" box="[795,910,628,654]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="species" species="layardii">
<emphasis id="EFD6EAC7C16FFFB4FCDF7003FC3007E0" box="[795,910,628,654]" italics="true" pageId="17" pageNumber="578">M. layardii</emphasis>
</taxonomicName>
, and
<taxonomicName id="1AA24D56C16FFFB4FC0D7003FBF207E0" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[969,1100,628,654]" class="Mammalia" family="Hyperoodontidae" genus="Mesoplodon" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="species" species="slangkopi" status="sp. nov.">
<emphasis id="EFD6EAC7C16FFFB4FC0D7003FBF207E0" box="[969,1100,628,654]" italics="true" pageId="17" pageNumber="578">M. slangkopi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC16FFFB4FB977003FB3207E0" box="[1107,1164,628,654]" pageId="17" pageNumber="578" rank="species">n. sp.</taxonomicNameLabel>
(see below) this feature is not visible in dorsal view.
</paragraph>
<paragraph id="DD1D36D5C16FFFB4FD7F70C4FB1B06E2" blockId="17.[677,1210,343,1382]" pageId="17" pageNumber="578">
The rostrum base lacks a prominental notch; the antorbital notch is wide and probably shallow, at the level of the premaxillary foramen and slightly anterior to the small maxillary foramen lying along the premaxilla-maxilla suture. Posterior to the antorbital notch, a low maxillary crest thickens the preorbital process, with a lateral slope similar to
<taxonomicName id="1AA24D56C16FFFB4FC177106FBE906E5" authority="Bianucci &amp; Lambert &amp; Post, 2007" authorityName="Bianucci &amp; Lambert &amp; Post" authorityYear="2007" box="[979,1111,881,907]" class="Mammalia" family="Ziphiidae" genus="Africanacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="578" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="EFD6EAC7C16FFFB4FC177106FBE906E5" box="[979,1111,881,907]" italics="true" pageId="17" pageNumber="578">Africanacetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E557BCC16FFFB4FB997105FB1B06E2" box="[1117,1189,882,908]" pageId="17" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
<paragraph id="DD1D36D5C16FFFB4FD7F71E6FC2B0127" blockId="17.[677,1210,343,1382]" pageId="17" pageNumber="578">The weakly asymmetric premaxillary sac fossae are anteromedially tilted. The ascending process is moderately constricted in anterior view. The foramina for the terminal nerve in the mesethmoid are strongly asymmetric; the left foramen is much smaller than the right.</paragraph>
<paragraph id="DD1D36D5C16FFFB4FD7F7638FC460008" blockId="17.[677,1210,343,1382]" pageId="17" pageNumber="578">The premaxillary crest is thin and laterally elongated, with a vertical anterior surface. The posterior projection of the premaxilla along the nasal reaches the frontal. The anterolateral corner of the roughly pentagon-shaped nasals forms only a small part of the premaxillary crest; the dorsal surface is anteromedially depressed. The naso-frontal suture is anteriorly pointed; the left nasal extends longer posteriorly than the right nasal.</paragraph>
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</treatment>
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