1219 lines
136 KiB
XML
1219 lines
136 KiB
XML
<document id="8BABD1B747C9DAAEEB20D65A530C93C2" ID-CLB-Dataset="298084" ID-DOI="10.3897/zookeys.1204.117261" ID-GBIF-Dataset="4ee75651-5bbb-424e-bca6-97c5391ca8c5" ID-ZooBank="0704A2BB-6173-44BD-A1F7-CB3EDCFDC5DC" ID-publisher-id="117261" URI-arpha="29A78280-EE32-57DD-97A4-22DB0AEC910C" URI-zoobank="http://zoobank.org/0704A2BB-6173-44BD-A1F7-CB3EDCFDC5DC" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1717518118260" checkinUser="pensoft" docAuthor="Che Engku Abdullah, Che Engku Siti Mariam, Idris, Izwandy, Fahmi, Afiq Durrani Mohd, Flaxman, Beth & Hutchings, Pat" docDate="2024" docId="37BB3E0577F650AD9CA20FA6E8FDE9F4" docLanguage="en" docName="ZooKeys 1204: 65-103" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/117261/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Marphysa merchangensis Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings 2024, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="103" masterDocId="29A78280EE3257DD97A422DB0AEC910C" masterDocTitle="Four new species of Marphysa (Annelida, Eunicida, Eunicidae) from the east coast of Peninsular Malaysia" masterLastPageNumber="103" masterPageNumber="65" pageNumber="65" updateTime="1717537492882" updateUser="ExternalLinkService">
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<mods:title id="234059209F42251CAB5E713D3B7F8A37">Four new species of Marphysa (Annelida, Eunicida, Eunicidae) from the east coast of Peninsular Malaysia</mods:title>
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<mods:name id="EE07C7D03D050E3475CA7C5878920E2B" type="personal">
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<mods:namePart id="24B80378709D9271452EDA0A4DB68575">Che Engku Abdullah, Che Engku Siti Mariam</mods:namePart>
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<mods:nameIdentifier id="EDA14C5D5248E926E84C75B6DA72E77E" type="ORCID">https://orcid.org/0009-0002-8371-1141</mods:nameIdentifier>
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<mods:affiliation id="E48E4213C8CD442F8BF684CDB5C4D521">Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:roleTerm id="DF0D2D1772EBC8C6E7286A7A67A77F1E">Author</mods:roleTerm>
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<mods:namePart id="3D994B212414A99A41B37C78D3D8BEB4">Idris, Izwandy</mods:namePart>
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<mods:nameIdentifier id="086A5C34D0523A135009B1DA857E7DD9" type="ORCID">0000-0003-1516-8175</mods:nameIdentifier>
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<mods:affiliation id="7245A0D04C909514C8BF1CFC8806675B">Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:roleTerm id="6016C853EAAD61AF4DBFC8AC9B3AD3D1">Author</mods:roleTerm>
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<mods:namePart id="DE04749B107125FBE6E5560F1B91B124">Fahmi, Afiq Durrani Mohd</mods:namePart>
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<mods:nameIdentifier id="573B5BA7C0984FFDF1B6770564F184A1" type="ORCID">0000-0002-5131-0098</mods:nameIdentifier>
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<mods:affiliation id="D1B77DF555D82B2EE20DEF2415437D53">Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:roleTerm id="9AC0C74BBF58BFBF25B837CE3F252EAF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B20B56A9274FA7465214CF719F3F756E">Flaxman, Beth</mods:namePart>
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<mods:nameIdentifier id="9CA4E06842DE8626D01C8927407101FF" type="ORCID">0000-0002-0329-9525</mods:nameIdentifier>
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<mods:affiliation id="C9E0D9D6D76209830B7D8EBE891879AD">South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia & Mangrove Research Unit, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:namePart id="FC3518EF3E42092A688DFB2B1961EF3D">Hutchings, Pat</mods:namePart>
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<mods:nameIdentifier id="336753D8AE484B0FF047277DB9B27CA2" type="ORCID">0000-0001-7521-3930</mods:nameIdentifier>
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<mods:affiliation id="226230EF2DAD29EDB9B7132C3C62D578">Mangrove Research Unit, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia & Faculty of Science and Marine Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:title id="1CAEC83F801D5A1AF91270DBAFB60E87">ZooKeys</mods:title>
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<mods:date id="1D4F84A62A336C27C436C20B9E1A1288">2024</mods:date>
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<mods:number id="1786B51F51D11EAE1EC21D01755C9928">2024-06-04</mods:number>
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<mods:detail id="A4142694AF718E0FA7987FA198D5705A" type="volume">
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<mods:number id="4D4DE130D12D7BF40B5BBED7CC88F506">1204</mods:number>
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<mods:start id="02D27348B6A16FC0622DF903BF5BE2F5">65</mods:start>
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<mods:end id="AEE8786F0E5F7BDC23815AE8F4AA3D74">103</mods:end>
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<mods:classification id="4F3FA29C9ABB9418A6B2BC618555B742">journal article</mods:classification>
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<mods:identifier id="F6D37145534C803C87A3CAB1608A34FD" type="CLB-Dataset">298084</mods:identifier>
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<mods:identifier id="E970DBF3D561F18B24B32D564531B2C8" type="DOI">10.3897/zookeys.1204.117261</mods:identifier>
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<mods:identifier id="D5A42A4D7F009BF5897BD220E1A3F8BD" type="ZooBank">0704A2BB-6173-44BD-A1F7-CB3EDCFDC5DC</mods:identifier>
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<treatment id="37BB3E0577F650AD9CA20FA6E8FDE9F4" ID-DOI="http://doi.org/10.5281/zenodo.11479292" ID-GBIF-Taxon="232137203" ID-Zenodo-Dep="11479292" ID-arpha="37BB3E05-77F6-50AD-9CA2-0FA6E8FDE9F4" ID-zoobank="https://zoobank.org/AD77E9BF-8D3D-458F-8AEA-BAC2E7AAFCB7" LSID="urn:lsid:plazi:treatment:37BB3E0577F650AD9CA20FA6E8FDE9F4" LSID-ZBK="urn:lsid:zoobank.org:act:AD77E9BF-8D3D-458F-8AEA-BAC2E7AAFCB7" httpUri="http://treatment.plazi.org/id/37BB3E0577F650AD9CA20FA6E8FDE9F4">
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<subSubSection id="81E2D3F737EF37655A48F842C61DEB21" type="nomenclature">
|
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<paragraph id="444FCB3F1839767562FDABC7D12A1DED">
|
||
<taxonomicName id="1C7365248963D80F4F66D33129D09C6F" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="639EDB2A1A650297E8462E6D19E74983" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="65FF9789A45757ABDF375AE4B5A73B94" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2F0BFF608F714ED8435D7C1C02CE3995" type="description">
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||
<paragraph id="10EE5CE2ED4440F57CCFFA358752E524">
|
||
<figureCitation id="A811E5A0B653EF57A062CB540D25D27C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval)." figureDoi="10.3897/zookeys.1204.117261.figure1" httpUri="https://binary.pensoft.net/fig/1062327">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="DD24029998EB7F0DE236AE8F1B157B8A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H)." figureDoi="10.3897/zookeys.1204.117261.figure2" httpUri="https://binary.pensoft.net/fig/1062328">2</figureCitation>
|
||
,
|
||
<figureCitation id="E1F116C9DD3CF79DE07DFC8E8AA60308" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8</figureCitation>
|
||
,
|
||
<figureCitation id="A2BE06A96A95E479E283424C8055D67F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9</figureCitation>
|
||
,
|
||
<figureCitation id="99C72382A561F8D56A264F2DFEA431FD" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sampling site in Merchang mangrove estuary A habitat of M. merchangensis sp. nov. B – C worm found in decayed root of Exoecaria agallocha (Malay: Bebuta) D live worms." figureDoi="10.3897/zookeys.1204.117261.figure10" httpUri="https://binary.pensoft.net/fig/1062336">10</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E5OAI" type="material">
|
||
<paragraph id="9CE788C956ADB11661E503525B518249">
|
||
<heading id="38546FD6E8F2F7FD8A435264E128FA0B" reason="title">Material examined.</heading>
|
||
</paragraph>
|
||
<paragraph id="E50DC21B491D871F7176D2165AFDB48B">
|
||
<materialsCitation id="14F332D6D016D9F24B55F0938D3D799A" ID-GBIF-Occurrence="4873238304" collectedFrom="mangrove estuary" collectingDate="2021-10" country="Malaysia" latitude="5.0232167" location="Merchang" longLatPrecision="1" longitude="103.2999" specimenCode="UMTAnn 2149" specimenCount="1" stateProvince="Terengganu" typeStatus="holotype">
|
||
<emphasis id="EB64FF52C78B399D217A1DD62A65E7D8" bold="true">
|
||
<emphasis id="27B4B17228E7FFA41AD92EE2A40C2E90" bold="true" italics="true">
|
||
<typeStatus id="AEE02D00F6504DC5493CDA7FC571749A" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="950EA7285A2AC7655FB7B0D5DD7D1DE9">UMTAnn 2149</specimenCode>
|
||
, complete, antero-ventrally dissected, some parapodia mounted for SEM
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="64558F38898688397E19F58D56F79C9F" ID-GBIF-Occurrence="4873238305" collectedFrom="mangrove estuary" collectingDate="2021-10" collectionCode="AM, ZRC" country="Malaysia" latitude="5.0232167" location="Merchang" longLatPrecision="1" longitude="103.2999" specimenCode="AM W. 54044, LACM-AHF 13494, ZRC. ANN. 1604" specimenCount="1" stateProvince="Terengganu" typeStatus="paratype">
|
||
<emphasis id="7E5CD2DE8C828985A399E5C20536DBF7" bold="true">
|
||
<emphasis id="991717A078A2773D1DB679CCE675FA5B" bold="true" italics="true">
|
||
<typeStatus id="55F45FEFA0123C5BB3B58A5BC7375D7C" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="E649C19E9B586DA9E45064DC2F52945B">
|
||
<named-content id="2E4A0E683D885AB16AF9DE2FBCEBCD57" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/australian-museum" xlink_title="Australian Museum, Sydney, Australia">AM</named-content>
|
||
W. 54044
|
||
</specimenCode>
|
||
, complete, some parapodia mounted for SEM.
|
||
<specimenCode id="656829368E655AC88CC4D70D13DAFCA8">LACM-AHF 13494</specimenCode>
|
||
to 13496, complete, some parapodia removed;
|
||
<specimenCode id="6379288DC6256294C7BE71816449C554">ZRC. ANN. 1604</specimenCode>
|
||
to 1606, complete, some parapodia removed; SAM-MB-A 096021, complete, some parapodia removed
|
||
</materialsCitation>
|
||
. All material was collected from the east coast of Peninsular
|
||
<collectingCountry id="9A0DADA73C9C78F7FE7B74C9722ED85B" name="Malaysia">Malaysia</collectingCountry>
|
||
,
|
||
<collectingRegion id="649FE26984CC7DD090786F71F7B79F5B" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
|
||
,
|
||
<location id="FD8A384AEEDDB6B178F57F51E4992A66" LSID="urn:lsid:plazi:treatment:37BB3E0577F650AD9CA20FA6E8FDE9F4:FD8A384AEEDDB6B178F57F51E4992A66" country="Malaysia" latitude="5.0232167" longLatPrecision="1" longitude="103.2999" name="Merchang" stateProvince="Terengganu">Merchang</location>
|
||
<collectedFrom id="0ABB180332AA8E571D4BF6B641155AC3">mangrove estuary</collectedFrom>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVPAI" specific-use="{"type":"Point","coordinates":[103.299900,5.023217]}">
|
||
<geoCoordinate id="7EA5E0C8D71647E89A2234D9E9FEC06C" degrees="05" direction="north" minutes="01.393" orientation="latitude" precision="1" value="5.0232167">05 ° 01.393 ' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="62DD38977261709E121ECD8C9DBE175D" degrees="103" direction="east" minutes="17.994" orientation="longitude" precision="1" value="103.2999">103 ° 17.994 ' E</geoCoordinate>
|
||
</named-content>
|
||
),
|
||
<date id="F3B9512A27CAF632A83CC1EA67F95BEB" value="2021-10">
|
||
<collectingDate id="2FAC9ADC79884C3A4BE7874E536FDF90" value="2021-10">October 2021</collectingDate>
|
||
</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1PAI" type="diagnosis">
|
||
<paragraph id="D99A522D622FB9CC47BE13A373DE10B4">
|
||
<heading id="ED16AD15337F08F294EE7177ACD5DC62" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="A053BED1912A2C925EFC760D6297ED92">Prostomium completely bilobed, five prostomial appendages without articulations; eyes present. Peristomium without peristomial cirri. Maxillary apparatus with four pairs of maxillae, an unpaired on the left side, MI with falcal arch extended at sub-right angle, basal outer edge arched, basal inner edge lacking curvature. MII with triangular teeth and without attachment lamella. MIII slightly curved, with equal-sized triangular teeth, without attachment lamella. MIV with rectangular and curved attachment lamella. Branchiae distributed along entire body. Dorsal cirri without articulations; postchaetal lobe well developed in anterior regions. Ventral cirri with swollen, inflated base. Sub-aciculae black, blunt, and translucent at distal end, pale brown in posterior-most parapodia. Supra-acicular chaetae include limbate, pectinate thin, narrow isodont with short and slender inner teeth, pectinate thick, wide isodont with short or long and slender inner teeth, and pectinate thick, narrow and wide anodont with long and thick inner teeth. Subacicular chaetae include only compound spinigers. Subacicular hook unidentate throughout chaetigers. Pygidium with two pairs of anal cirri, without articulation.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E6PAI" type="description">
|
||
<paragraph id="3E098761313740EA1BDDB34220793561">
|
||
<heading id="18757944F076491AE7CA9DB64D55A379" reason="title">Description</heading>
|
||
</paragraph>
|
||
<paragraph id="690A284239EEA3C2C2AD1458208C0A7F">
|
||
<emphasis id="F4E7E393991DF053957E1F9228A68230" bold="true">
|
||
(based on
|
||
<typeStatus id="4CE86A4994488DEBE41B9212172284C5" type="holotype">holotype</typeStatus>
|
||
, with variation in parentheses for
|
||
<typeStatus id="520B7FBD120C39C9403DEE1538E5751F" type="paratype">paratypes</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Preserved specimen beige (Fig.
|
||
<figureCitation id="199A8F4D5F270B6C5693CD29F83FAF06" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 A</figureCitation>
|
||
), 257 (165–294) chaetigers,
|
||
<quantity id="D2F76A70771A4ED5ABBC39CCB102B1B0" metricMagnitude="-2" metricUnit="m" metricValue="9.4" unit="mm" value="94.0">94 mm</quantity>
|
||
(
|
||
<quantity id="62BF0781A45B5A77A2BAEC0906C7BE9C" metricMagnitude="-2" metricUnit="m" metricValue="9.05" metricValueMax="14.4" metricValueMin="3.7" unit="mm" value="90.5" valueMax="144.0" valueMin="37.0">37–144 mm</quantity>
|
||
) long,
|
||
<abbrev id="ABBRID0ELQAI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
-
|
||
<quantity id="180FEFD33A2654CBED8DB26D854BF646" metricMagnitude="-3" metricUnit="m" metricValue="5.25" unit="mm" value="5.25">5.25 mm</quantity>
|
||
(
|
||
<quantity id="9483477828AEFF247A301015563E9576" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.85" metricValueMin="3.45" unit="mm" value="4.65" valueMax="5.85" valueMin="3.45">3.45–5.85 mm</quantity>
|
||
),
|
||
<abbrev id="ABBRID0EPQAI" xlink_title="width at chaetiger 10">W 10</abbrev>
|
||
-
|
||
<quantity id="2C39A00B9E1CFEA40EB476AD17857CA7" metricMagnitude="-3" metricUnit="m" metricValue="2.85" unit="mm" value="2.85">2.85 mm</quantity>
|
||
(
|
||
<quantity id="18F8F008FCBADE48DBE3CF2714A53B0D" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="3.15" metricValueMin="1.95" unit="mm" value="2.55" valueMax="3.15" valueMin="1.95">1.95–3.15 mm</quantity>
|
||
), excluding parapodia. Anterior region of body with dorsum convex and flat ventrum, without groove (Fig.
|
||
<figureCitation id="A1CAF894426C82C185F6765D639FB807" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 A</figureCitation>
|
||
); body depressed from chaetiger 25, elongated and tapering at distal end. Live specimens pink with red branchiae (Fig.
|
||
<figureCitation id="DDADCA26554A6EF4714D5710ABB0626C" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sampling site in Merchang mangrove estuary A habitat of M. merchangensis sp. nov. B – C worm found in decayed root of Exoecaria agallocha (Malay: Bebuta) D live worms." figureDoi="10.3897/zookeys.1204.117261.figure10" httpUri="https://binary.pensoft.net/fig/1062336">10 D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="E07E80130AB47ACB439CD9D86F323FCA" ID-DOI="10.3897/zookeys.1204.117261.figure8" ID-arpha="5E81F118-174B-5A85-8156-3EAF79CBC5C0" httpUri="https://binary.pensoft.net/fig/1062334" startId="F8">
|
||
<paragraph id="A0F19B5A8AF81A2D3ACDE2B0F8113D75">
|
||
<label id="F3E6DAA3ECBB654CA80F150A1C8FDD2B">Figure 8.</label>
|
||
</paragraph>
|
||
<paragraph id="522B53CD29563636EA8BC76460A85520">
|
||
<taxonomicName id="E4AA8E6BEACB54C4CB21C6F5E0722D4B" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="52BEAFBB2ED0A005AC678DFDE5DA0897" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="63C784710B4E86507A9C6A2F882A4E3F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype UMTAnn 2149 (
|
||
<emphasis id="1B6CE16A1401F62B38CABDDB1376BA69" bold="true">B – H</emphasis>
|
||
), paratype UMTAnn 2148 (
|
||
<emphasis id="901ABAE802C4C14E3919250F7207244F" bold="true">A</emphasis>
|
||
). Light microscopy images and digital drawing
|
||
<emphasis id="7DE41CD69483459D21F03B8196D02C2F" bold="true">A</emphasis>
|
||
anterior end, lateral view
|
||
<emphasis id="FDF85E7697D2F58CDC12756EF035FDFB" bold="true">B</emphasis>
|
||
anterior end, dorsal view. Arrows indicate eyes
|
||
<emphasis id="D5B358162C92F935A39BE45BBACBC37B" bold="true">C</emphasis>
|
||
maxillae, dorsal view
|
||
<emphasis id="4B7E20CBF15D34C74A14F969C6B069CD" bold="true">D</emphasis>
|
||
mandibles, dorsal view
|
||
<emphasis id="EBA00CF4FB5C7603A8F15B62170C87FB" bold="true">E</emphasis>
|
||
parapodium, chaetiger 10
|
||
<emphasis id="22E8E8610C561C5ED243D739C502A9D7" bold="true">F</emphasis>
|
||
parapodium, chaetiger 134
|
||
<emphasis id="8480D6DFB35828D49D81801DE2347B24" bold="true">G</emphasis>
|
||
parapodium, chaetiger 250
|
||
<emphasis id="36DE627735B6FE91DD723130DE9E9A69" bold="true">H</emphasis>
|
||
posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (
|
||
<emphasis id="3D82D29CE040DA3844717068C14AA654" bold="true">A – D, H</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis id="4C7C9B691FF3FC41FF177C94BF684031" bold="true">E – G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E05F2807FB8326A693DEC32581049090">
|
||
Prostomium bilobed, anteriorly rounded with two dorsoventrally flattened lobes with an anterior notch between them (Fig.
|
||
<figureCitation id="3EC3C16E2B3C546AAEBE3A41B2A8656C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 A, B</figureCitation>
|
||
). Prostomial appendages in a semicircle, median antenna isolated by a gap (Fig.
|
||
<figureCitation id="E399BF8DAAE459FA7C73FD7E0DA12CBB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 B</figureCitation>
|
||
). Palps reach middle of second peristomial ring; lateral antennae reaching chaetiger 2; median antenna reaching chaetiger 3. Palpophores and ceratophores ring-shaped, short, and thick; palpostyles and ceratostyles tapering, and slender. Prostomial appendage peduncles absent. A pair of faded brown eyes present at posterior base of prostomium, between palps and lateral antennae (Fig.
|
||
<figureCitation id="574BBF87CF1EFFB3E4D06D1A524C7C5E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 B</figureCitation>
|
||
). Peristomium larger and wider than prostomium; first ring is 2.5 × longer than second ring, separation between rings distinct on all sides.
|
||
</paragraph>
|
||
<paragraph id="AAA1AA596A53EAC51F1E6DAB746BDAD8">
|
||
Maxillae pale brown (Fig.
|
||
<figureCitation id="BF33129578456E1DC1E906D0D1337B69" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 C</figureCitation>
|
||
), and maxillary formula as follows:
|
||
<abbrev id="ABBRID0E6SAI" xlink_title="maxillary formula, roman numerals refer to number of maxilla">MF</abbrev>
|
||
= 1 + 1, 5 (4–5) + 5 (5–6), 7 (6–7) + 0, 4 (4–5) + 8 (5–8), 1 + 1. Maxillary carrier ~ 2.5 × shorter than MI, rectangular anteriorly, triangular posteriorly. MI forceps-like, without attachment lamellae, falcal arch extended at sub-right angle, basal outer edge arched, basal inner edge lacking a curvature. Closing system ~ 3 × shorter than MI. Ligament between MI and MII pale brown. MII without attachment lamella, teeth triangular, distributed on <1 / 2 length of the plate. Ligament between MII and MIII pale brown. MIII single, longer than left MIV slightly curved, with equal-sized triangular teeth, without attachment lamella. Left MIV short (<1 / 2 the size of right MIV) with rectangular attachment lamellae. Right MIV long with curved attachment lamellae, teeth triangular, decreasing in size and teeth curved posteriorly. MV paired. Mandible pale brown, with concentric stripes, longer than MI; cutting plates whitish (Fig.
|
||
<figureCitation id="758646F912C05D439387A0B3F5209521" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A917520AA7DA3748426FCCDDBD234301">
|
||
First few parapodia located ventrolaterally but gradually becoming dorsolateral in subsequent segments. Chaetal lobes rounded in anterior and posterior chaetigers, conical in median chaetigers (Fig.
|
||
<figureCitation id="FD51027B540D851664796EE11979B8EB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 E – G</figureCitation>
|
||
). Prechaetal lobe shorter than chaetal lobe throughout body. Postchaetal lobe digitiform in first three chaetigers then rounded thereafter; longer than chaetal lobe in median chaetigers onwards, become shorter and absent in the posterior-most chaetigers. Dorsal cirri digitiform and slender, longer than ventral cirri anteriorly, slightly longer or similar from mid-body towards posterior-most chaetigers (Fig.
|
||
<figureCitation id="319A39E1B211B4A40A2C1FDD6D2516E1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 E – G</figureCitation>
|
||
). Ventral cirri thumb-shaped with rounded wide tips in first few chaetigers, basally inflated with digitiform tip from chaetiger 15, and gradually becoming conical posteriorly (Fig.
|
||
<figureCitation id="10A46939BD4E61D20BD20F983C2E0700" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 E – G</figureCitation>
|
||
). Branchiae pectinate, from chaetiger 24 (16–27) and continuing to last ~ 10 chaetigers, branchial filaments 4 × longer than dorsal cirri where best developed; number of filaments increasing from three anteriorly to six in mid-body, decreasing to one in last several chaetigers.
|
||
</paragraph>
|
||
<paragraph id="85CD4B197FB0E5D30F7C0CE0259B4700">
|
||
Notoaciculae absent, neuroaciculae black, blunt, and translucent at distal end along most of body, pale brown in posterior-most parapodia; ~ 2 or 3 per parapodium in anterior, one per parapodium in median and posterior chaetigers (Fig.
|
||
<figureCitation id="4DFF95364643C295E43D9439425B9223" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 E – G</figureCitation>
|
||
). Supra-acicular chaetae with limbate capillaries and pectinates.
|
||
<specimenCount id="58901C86B29A1010430FF271B3C3D1AE" count="5" type="generic" typeStatus="types">Five types</specimenCount>
|
||
of pectinate chaetae present (types 1, 4, 5, 6, 8) (see Fig.
|
||
<figureCitation id="0843EE9638145F26DAC1A3596C33EE35" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The schematic drawing of type of pectinate chaetae present in Marphysa from Terengganu A thin, narrow isodont with short and slender inner teeth (type 1) B thin, wide isodont with short and slender inner teeth (type 2) C thin, narrow heterodont with short and slender inner teeth (type 3) D thick, wide isodont with short and slender inner teeth (type 4) E thick, wide isodont with short and slender inner teeth (type 5) F thick, narrow anodont with long and thick inner teeth (type 6) G thick, wide anodont with long and slender inner teeth (type 7) H thick, wide anodont with long and thick inner teeth (type 8). Scale bars: 18 µm (A); 35 µm (B, G); 20 µm (C); 38 µm (D); 32 µm (E); 13 µm (F); 30 µm (H)." figureDoi="10.3897/zookeys.1204.117261.figure2" httpUri="https://binary.pensoft.net/fig/1062328">2</figureCitation>
|
||
): type 1: thin, narrow isodont with 7–12 short and slender inner teeth, outer teeth longer, but with varying lengths, present in anterior and median body region (Fig.
|
||
<figureCitation id="B301E53C31782F18EF81492B0A51C944" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 A, B</figureCitation>
|
||
); type 4: thick, wide isodont with 12–15 short and slender inner teeth, present only in median and posterior region (Fig.
|
||
<figureCitation id="EA5E5B5D75FCFF260CF8B7784C59DFBD" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 C</figureCitation>
|
||
); type 5: thick, wide isodont, with 15–18 long and slender inner teeth, only present in posterior region (Fig.
|
||
<figureCitation id="2DC8F7B0016D14FDD5775183678F451B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 D</figureCitation>
|
||
); type 6: thick, narrow anodont with 11 or 12 long thick teeth, only present in posterior region (Fig.
|
||
<figureCitation id="B1E0B2C44921249CEE6010F34E4D5587" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 E</figureCitation>
|
||
); type 8: thick, wide anodont, with five inner long and thick teeth, only present in the posterior region (Fig.
|
||
<figureCitation id="B48C71E4A52AF86F2616FD0D9962CB6C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 F</figureCitation>
|
||
). Subacicular chaetae with compound spinigers (Fig.
|
||
<figureCitation id="8F9A65C9F7D0E9464D5011E5F1F1F59C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 G</figureCitation>
|
||
). Subacicular hooks pale brown, translucent at distal end, emerge from chaetiger 37 (26–42) and then present on all chaetigers, one per parapodium; subacicular hooks unidentate throughout chaetigers (Fig.
|
||
<figureCitation id="4F8618759A1A1E785C6F73AFF7549ADE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM images of Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (A, B, E, H), paratype AM W. 54044 (C, D, F, G) A, B pectinate chaetae, chaetiger 10 C pectinate chaetae, chaetiger 164 D pectinate chaetae, chaetiger 245 E pectinate chaetae, chaetiger 250 F pectinate chaetae, chaetiger 265 G spiniger chaetae, chaetiger 10 H subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (A – C, E); 10 µm (D, F, H); 50 µm (G)." figureDoi="10.3897/zookeys.1204.117261.figure9" httpUri="https://binary.pensoft.net/fig/1062335">9 H</figureCitation>
|
||
). Pygidium with crenulated margin, with two pairs of tapering pygidial cirri attached to ventral side of pygidium, dorsal pair ~ 4 × longer than ventral (Fig.
|
||
<figureCitation id="9B844B74FC5EC49F91C0EB176E6AE5C9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Marphysa merchangensis sp. nov. Holotype UMTAnn 2149 (B – H), paratype UMTAnn 2148 (A). Light microscopy images and digital drawing A anterior end, lateral view B anterior end, dorsal view. Arrows indicate eyes C maxillae, dorsal view D mandibles, dorsal view E parapodium, chaetiger 10 F parapodium, chaetiger 134 G parapodium, chaetiger 250 H posterior segments and pygidium, ventral view. Arrows show the short pair of pygidial cirri. Abbreviations; MI – MV: maxillae I – V, Ac: aciculae, Dc: dorsal cirrus, Vc: ventral cirrus, PCl: prechaetal lobe, Cl: chaetal lobe, PtCl: postchaetal lobe, Sah: subacicular hook, Bf: branchial filament. Scale bars: 1 mm (A – D, H); 0.1 mm (E – G)." figureDoi="10.3897/zookeys.1204.117261.figure8" httpUri="https://binary.pensoft.net/fig/1062334">8 H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="1B11AF690DFD966B6081C28306878A31" ID-DOI="10.3897/zookeys.1204.117261.figure9" ID-arpha="BA060619-54BA-5ED8-9AF9-B1D2BB0C9F0B" httpUri="https://binary.pensoft.net/fig/1062335" startId="F9">
|
||
<paragraph id="D8C1229641CE08FF4869AFA7E10091A8">
|
||
<label id="2710924759CCF5C1D3243651FAAA32E2">Figure 9.</label>
|
||
</paragraph>
|
||
<paragraph id="96221BC7B69EB5D81B149BFA63A49ADD">
|
||
SEM images of
|
||
<taxonomicName id="407D727C0367463AC36B7B452C49D43D" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="FC2B576A9CBDC9EA65AB4B6A06A10A56" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA27AEB795A1C1F2262F9478EA6844C6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype UMTAnn 2149 (
|
||
<emphasis id="CC5B883356C07126B5F1B61BC4DA6C19" bold="true">A, B, E, H</emphasis>
|
||
), paratype
|
||
<named-content id="A36A12379AFF299EEE935FFF23A6B753" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/australian-museum" xlink_title="Australian Museum, Sydney, Australia">AM</named-content>
|
||
W. 54044 (
|
||
<emphasis id="1B41006522158A87B9331DE9C14857D7" bold="true">C, D, F, G</emphasis>
|
||
)
|
||
<emphasis id="825AB91C4178AD3B2E17B3FEFDB999D5" bold="true">A, B</emphasis>
|
||
pectinate chaetae, chaetiger 10
|
||
<emphasis id="ACAE3A9C90E010FA2EDFDD4F2A889EF3" bold="true">C</emphasis>
|
||
pectinate chaetae, chaetiger 164
|
||
<emphasis id="D74FCACCAB9FCFDCC3BDE7FE48FCB3DA" bold="true">D</emphasis>
|
||
pectinate chaetae, chaetiger 245
|
||
<emphasis id="0F558B04D8765CD020FAB1D14DF26AF3" bold="true">E</emphasis>
|
||
pectinate chaetae, chaetiger 250
|
||
<emphasis id="55DBD3E1F96F915F54698596A6D1A9B8" bold="true">F</emphasis>
|
||
pectinate chaetae, chaetiger 265
|
||
<emphasis id="79FCF177F37EAEA6CCBF3F8267628472" bold="true">G</emphasis>
|
||
spiniger chaetae, chaetiger 10
|
||
<emphasis id="BCAE67CB0BFB841210D9FA333B64EF94" bold="true">H</emphasis>
|
||
subacicular hook, chaetiger 128. Numbers denoted by arrows indicate the type of pectinate chaetae; 1. Thin, narrow isodont; 4, 5. Thick, wide isodont; 6. Thick, narrow anodont; 8. Thick, wide anodont. Scale bars: 5 µm (
|
||
<emphasis id="64B64BCE160D921B85B754FEC52C5953" bold="true">A – C, E</emphasis>
|
||
); 10 µm (
|
||
<emphasis id="CD278248057D111E409265CAD75824B7" bold="true">D, F, H</emphasis>
|
||
); 50 µm (
|
||
<emphasis id="11825FFBF4D805029C98361A85DE0BA8" bold="true">G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUWAI" type="etymology">
|
||
<paragraph id="7960B0EFE6AC7AC70A606D5EB5A5D326">
|
||
<heading id="58E5C5A0B0FD997424D7019135B021FE" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="A2A80B770DA59D5C1DE4FFA8F6709D33">The name denotes the type locality (Merchang estuary) where the specimens were collected.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZWAI" type="type locality">
|
||
<paragraph id="DADAAA2807D08CB0453C7DEF2C957F83">
|
||
<heading id="4E40471ABFB3483DCBFA9F6FDB069A4D" reason="title">Type locality.</heading>
|
||
</paragraph>
|
||
<paragraph id="EF801F622E885D7637C70D98591D8B5D">
|
||
South
|
||
<collectingCountry id="8A20A826DB9F081D1B0740342B779D0A" name="China">China</collectingCountry>
|
||
Sea,
|
||
<collectingCountry id="FA6020F65DE00E6500D95A2C1F9E88CB" name="Malaysia">Malaysia</collectingCountry>
|
||
, east coast of Peninsular,
|
||
<collectingRegion id="096297F8690C2209B5036E34E8465F5F" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
|
||
, Merchang mangrove estuary (see Fig.
|
||
<figureCitation id="DB36925BB45EEBD25186FF2DADCB8F36" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing sampling sites of four new Marphysa species in Terengganu mangrove forest, east coast of Peninsular Malaysia A location of Terengganu on the east of Peninsular Malaysia B symbols indicate each sampling site; Setiu Wetlands (red star), Kuala Ibai (red rhombus), Merchang (red triangle), and Kerteh (red oval)." figureDoi="10.3897/zookeys.1204.117261.figure1" httpUri="https://binary.pensoft.net/fig/1062327">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EDXAI" type="distribution">
|
||
<paragraph id="6DE4F4F0601BEE3184F419B104121727">
|
||
<heading id="B48AC687B65745D8DA1F550736EB9825" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="F2777FD79F3680BA133E8A57648D4317">
|
||
Known only from the type locality and Setiu Wetlands,
|
||
<collectingRegion id="DF7A499B90DBFB2AC7EA8C3B89FC1D09" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
|
||
,
|
||
<collectingCountry id="31765C022C325C642A923815A561420D" name="Malaysia">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EIXAI" type="habitat">
|
||
<paragraph id="D3A22AE903F0F66BE0C42F8EB103B092">
|
||
<heading id="327DA5147B388171FE4CF6C516815CC4" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="66113F690051AFED464EB9FFAF0DE9B0">
|
||
Gravelly and slightly gravelly sand (Table
|
||
<tableCitation id="BA40C1BFA8EF7835B3B4E74FC4384FC8" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Distance of sampling sites from river mouth, type of sediment textures and total organic matter content of sampling sites in Terengganu mangrove forest." httpUri="http://table.plazi.org/id/A7F5E5E96E47328E73CF7254E753C455" tableUuid="A7F5E5E96E47328E73CF7254E753C455">4</tableCitation>
|
||
), burrowing in decayed roots of the mangrove
|
||
<taxonomicName id="2FE53B36D59DAF070572FF437CBD408B" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="agallocha">
|
||
<emphasis id="3ECAFC440CB5641F01F72363B61010F1" italics="true">E. agallocha</emphasis>
|
||
</taxonomicName>
|
||
(Malay: Bebuta) (Fig.
|
||
<figureCitation id="54C33DBA6DAEBF6E7E2FB6B17078313F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Sampling site in Merchang mangrove estuary A habitat of M. merchangensis sp. nov. B – C worm found in decayed root of Exoecaria agallocha (Malay: Bebuta) D live worms." figureDoi="10.3897/zookeys.1204.117261.figure10" httpUri="https://binary.pensoft.net/fig/1062336">10 A – C</figureCitation>
|
||
), burrowing in the sediments within an area populated with
|
||
<taxonomicName id="BC71570358637A065A75856281DF6D8F" class="Magnoliopsida" family="Malvaceae" genus="Talipariti" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="species" species="tiliaceum">
|
||
<emphasis id="4B6CD246CF7ACF202FD22B3E0CA5992B" italics="true">Talipariti tiliaceum</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="218AE0A91CB36EDC4171B3EF49BF757B" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Sampling in Setiu Wetlands (river area) A, C habitat of M. setiuense sp. nov. and M. merchangensis sp. nov. within Talipariti tiliaceum B Marphysa setiuense sp. nov. in situ D live M. setiuense sp. nov." figureDoi="10.3897/zookeys.1204.117261.figure13" httpUri="https://binary.pensoft.net/fig/1062339">13 C</figureCitation>
|
||
) with salinity 26 ‰ during spring low tide.
|
||
</paragraph>
|
||
<caption id="4858D11B0961213F8E98A8E34F72E3CE" ID-DOI="10.3897/zookeys.1204.117261.figure10" ID-arpha="A5026C06-9BC9-543F-9E76-2B0330B1408C" httpUri="https://binary.pensoft.net/fig/1062336" startId="F10">
|
||
<paragraph id="23F53EB91A6C6B6A8B96344787B3073D">
|
||
<label id="FC80F84946EAA83C1048554EE99CAB1D">Figure 10.</label>
|
||
</paragraph>
|
||
<paragraph id="1EBCC6FB0A250E71307964CBE3956A21">
|
||
Sampling site in Merchang mangrove estuary
|
||
<emphasis id="944AE61836D63BE7E403FA2A04113767" bold="true">A</emphasis>
|
||
habitat of
|
||
<taxonomicName id="DEE21F0AE7E1C034DDC938E9EEA2E894" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="96FB53B6CC85E2F58E9C25255DDD4830" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D9A8CD9E19D7B19903904FF688928A9D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="1FACAA9C7E16C512D41B709DBF9D85AA" bold="true">B – C</emphasis>
|
||
worm found in decayed root of
|
||
<emphasis id="14D5B38EFB93128591A75C784FA71D35" italics="true">
|
||
Exoecaria
|
||
<taxonomicName id="42072B8189F55F162344333826D586EE" class="Polychaeta" family="Eunicidae" genus="Eunice" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="agallocha">agallocha</taxonomicName>
|
||
</emphasis>
|
||
(Malay: Bebuta)
|
||
<emphasis id="117D1A35334D8101FAA3AFA431E3F5C3" bold="true">D</emphasis>
|
||
live worms.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZZAI" type="remarks">
|
||
<paragraph id="9703C42C74EAC1B4E3F735A6C190E7FC">
|
||
<heading id="D5AABFA50564C8DE50855EE18509E9A5" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="98CAF5BEB4E7CB60C68513134A3A783F">
|
||
With the presence of only compound spinigers along the whole body and branchiae along most of the body,
|
||
<taxonomicName id="92E6A8DB7609B35FB20F6DE87C44946F" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="A0C4A0A580112511621837E2F89C7610" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9FA35EAAE78229B522D03A85E6308704" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName id="1C7E3A28787F0780F8F7EC18C9F23AC6" authorityName="de Quatrefages" authorityYear="1866" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="genus">
|
||
<emphasis id="89F22458FB60B108D7EFC33AE9DDE625" italics="true">Marphysa</emphasis>
|
||
</taxonomicName>
|
||
Group B (Sanguinea). Other
|
||
<taxonomicName id="F3C79BAA7B519056EE28C9F84760C776">
|
||
<emphasis id="A599B7321C820C6565C9DC8490B304C1" italics="true">Marphysa</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from Sanguinea-group occurring in the same water body (South
|
||
<collectingCountry id="4469028B04F5472A5752B31F6463DDD4" name="China">China</collectingCountry>
|
||
Sea) as
|
||
<taxonomicName id="DFD5583B2E988A712D1401846928CB31" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="4E660B75F6ABB3B8DA87363160CA5AE3" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DF1E363F441FD2CC1715178E4D7A1827" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are
|
||
<taxonomicName id="0347C556D8870F37625E61F616685707" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="BBC1EB1B9141B0AF0CE094C0E01F4C72" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD53BEEBAA7BCDC0CC7B8C98BB75C58D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="1EE984FB92E54862C914B5A829AE77C0" authority="Wang, Zhang & Qiu, 2018" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="3794B31054C5915A257834A3AC7E96C2" italics="true">M. hongkongensa</emphasis>
|
||
<bibRefCitation id="3F63CE4E714C11A2338B05FCD48AF528" DOI="10.6620/ZS.2018.57-48" author="Wang Z & Zhang Y & Qiu JW" refId="B43" refString="Wang Z, Zhang Y, Qiu JW (2018) A new species in the Marphysa sanguinea complex (Annelida, Eunicidae) from Hong Kong. Zoological Studies 57. https: // doi. org / 10.6620 / ZS. 2018.57 - 48" year="2018">Wang, Zhang & Qiu, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality:
|
||
<collectingCountry id="DBE74900B3A1F18049951679722FA82D" name="Hong Kong">Hong Kong</collectingCountry>
|
||
),
|
||
<taxonomicName id="4090CE80ED154BA64A81C410BCBA3788" authority="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings, 2019" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="8FCADAB21E602E7D26F0268E14859ADF" italics="true">M. iloiloensis</emphasis>
|
||
<bibRefCitation id="4B2C75925449B034209FBC3B472A8395" DOI="10.11646/zootaxa.4674.2.7" author="Glasby CJ & Mandario MAE & Burghardt I & Kupriyanova E & Gunton LM & Hutchings P" issue="2" journalOrPublisher="Zootaxa" pagination="264-282" refId="B14" refString="Glasby CJ, Mandario MAE, Burghardt I, Kupriyanova E, Gunton LM, Hutchings P (2019) A new species of the sanguinea-group Quatrefages, 1866 (Annelida: Eunicidae: Marphysa) from the Philippines. Zootaxa 4674 (2): 264 - 282. https: // doi. org / 10.11646 / zootaxa. 4674.2. 7" title="A new species of the sanguinea-group Quatrefages, 1866 (Annelida: Eunicidae: Marphysa) from the Philippines." volume="4674" year="2019">Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality:
|
||
<collectingCountry id="20EA099EB50BEE2EAEA8F63E69F7B910" name="Philippines">Philippines</collectingCountry>
|
||
),
|
||
<taxonomicName id="E5FF2442305C5CA0D159E0B9CB87BC07" authority="Liu, Hutchings & Sun, 2017" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="multipectinata">
|
||
<emphasis id="4859EED8A81B95EAB2AC7D03D7B24C9C" italics="true">M. multipectinata</emphasis>
|
||
<bibRefCitation id="1325A395E316A2D07350BDEA80F3ECB1" DOI="10.11646/zootaxa.4263.2.2" author="Liu Y & Hutchings P & Sun S-C" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228 - 250. https: // doi. org / 10.11646 / zootaxa. 4263.2. 2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu, Hutchings & Sun, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Shimen,
|
||
<collectingCountry id="90E97A7D851C7985B38BA281E10FB587" name="Taiwan">Taiwan</collectingCountry>
|
||
of
|
||
<collectingCountry id="E914CBA74695EE56EEA665A06BDF4B22" name="China">China</collectingCountry>
|
||
),
|
||
<taxonomicName id="9C2535C29C3F1EB0DE72D55E9A8F8D41" authority="Treadwell, 1936" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="6BC53E98638F63B0F1FA088A5A5258ED" italics="true">M. orientalis</emphasis>
|
||
<bibRefCitation id="9473E85715E57F000B65CD7D558062B0" DOI="10.5479/si.00963801.83-2984.261" author="Treadwell AL" issue="2984" journalOrPublisher="Proceedings of the United States National Museum" pagination="261-278" refId="B41" refString="Treadwell AL (1936) Polychaetous annelids from Amoy, China. Proceedings of the United States National Museum 83 (2984): 261 - 278. https: // doi. org / 10.5479 / si. 00963801.83 - 2984.261" title="Polychaetous annelids from Amoy, China." volume="83" year="1936">Treadwell, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Xiamen,
|
||
<collectingCountry id="961FD4FDAE08B310688EC917854E856D" name="China">China</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C6F80EE695DAEEE89DF64792C61C4DE" authority="Liu, Hutchings & Sun, 2017" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="AC6BDDA7621CEBCF85C0B06000F1B906" italics="true">M. tribranchiata</emphasis>
|
||
<bibRefCitation id="991AE3FF85D830AE8FA0B67E6DA9A641" DOI="10.11646/zootaxa.4263.2.2" author="Liu Y & Hutchings P & Sun S-C" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228 - 250. https: // doi. org / 10.11646 / zootaxa. 4263.2. 2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu, Hutchings & Sun, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Wanli,
|
||
<collectingCountry id="CEBBF6D04858A1135D2EB272711153ED" name="Taiwan">Taiwan</collectingCountry>
|
||
of
|
||
<collectingCountry id="A5BCE4AD50BBF99B8D47E85100EAD43A" name="China">China</collectingCountry>
|
||
), and
|
||
<taxonomicName id="7A4CF9D29A39066BF4A81FD7063CA688" authority="Liu, Hutchings & Sun, 2017" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="707EC507C41C47D03ADE1465CEF6A815" italics="true">M. tripectinata</emphasis>
|
||
<bibRefCitation id="C7BF454DC26B647EB3D9CEB1CB6E9CD5" DOI="10.11646/zootaxa.4263.2.2" author="Liu Y & Hutchings P & Sun S-C" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228 - 250. https: // doi. org / 10.11646 / zootaxa. 4263.2. 2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu, Hutchings & Sun, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
(type locality: Beihai,
|
||
<collectingCountry id="985C674A869EDB069556ABFCC28DE329" name="China">China</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7B011AF10BCB4EA27EB8F2EBA6306783">
|
||
<taxonomicName id="7DDE4F7C2351DF4E330C9BB759FEDBF5" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="7EEDF26E4AF92556F85027A2D5A1766A" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5E1E1372F72259DEFA3F4BD88AA4A1D2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="0BC400858D4D3FB2CD5BEF6C2237201D" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="3F01E5CC8116E23BF1DAED260863E6D4" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DF6C8FD0068CB024984C8646FEC65099" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in having a pair of eyes and the absence of peduncle on the prostomial appendages. However, they can be differentiated by the number of types of pectinate chaetae, maxillary formula, chaetiger on which the branchiae and subacicular hooks occur, shape of dorsal cirri, chaetal lobes and subacicular hooks. Number of types of pectinate chaetae in
|
||
<taxonomicName id="531CB26CC8668D7CEBBFB6EBC4B74F35" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="697F6953B6E7CE21D83AC0CE964AE1A3" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="110BC76027A385D5B195A2FC58D1113F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is five (types 1, 4, 5, 6, 8), whereas in
|
||
<taxonomicName id="34113F0128E12561CE3CB1BB56BD7C7A" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="588F10F8E104F9102E6EB6B249AF815A" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D2B170AD05CA1126AE71D27F541B21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
there are four (types 1, 2, 7, 8), and they lack the thick, wide isodont pectinate chaetae (types 4, 5).
|
||
<taxonomicName id="C90433FDE804EE90909A35FBECEA5527" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="8A5FF390AD6DC593B31F629CBB38404B" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="38E04676327D9F2DE1B415CFA67DB73E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<abbrev id="ABBRID0EJ6AI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
: 5.25 (3.45–5.85) mm) has more denticles on MIII 7 (6–7) + 0 compared to
|
||
<taxonomicName id="B9D25DBB42B03ABA74DE2E88F7067A20" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="8434F29502DB650676FC55F4AEC21C59" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B8BAFD711FE62921DAD6FF5833121B79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<abbrev id="ABBRID0EY6AI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
: 2.7 (2.85–4.8) mm) which has MIII: 5 (4–6) + 0. Branchiae and subacicular hook of
|
||
<taxonomicName id="CC1800E1DEF67CBDC55870A8502B27E3" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="0BF20DE3F70F04AD96FD3B5A7FCF5250" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="67CA0C27097CB6E05C787A3004106549" rank="species">sp. nov.</taxonomicNameLabel>
|
||
occur later (chaetiger 24 (16–27) and 37 (26–42 )), respectively) compared to
|
||
<taxonomicName id="8277FD67CC5A0EE78E8846E6546D40C1" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="091632561ABA0D4F98003F132D015433" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CF8EEFCC6D7A81A45760ADE8E248CB0F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, where they occur from chaetiger 20 (15–25) and 25 (21–38), respectively.
|
||
<taxonomicName id="70AE05A5638F75A2ED56575BCE1C5263" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="760FEB9495D636110EAB821B4B711546" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66AE7F2F0C765E155D5088B91A0120EE" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has digitiform dorsal cirri along the whole body, while
|
||
<taxonomicName id="F9ABB6061DCB64FE413E8B1FC9DEEFAB" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="E04C3D15B4E408F2100897BA003DCE48" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C095BD9DE874AB6299166F95E6A1A226" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has both thumb-shaped and digitiform dorsal cirri.
|
||
<taxonomicName id="26709396850F872621845F575E4E33A6" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="5B7DD117A1F53AF610F9B8C962E9D269" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="775A42E4E7363F63ED01D41F3F1A94FF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has rounded shaped chaetal lobe in the anterior and posterior, and conical in the median region, whereas
|
||
<taxonomicName id="111A8F3FDA17E114AEFA142FD3937658" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="0330E64A7D0E4F3036601F152881F7EC" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2036E5B396CAEE5FC0720505BFF4485D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has rounded chaetal lobes on all parapodia. Finally,
|
||
<taxonomicName id="22B138C61D2A3F99CC7436C2995BD04A" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="75EFA760F4CFC0C487CC0D0039B1FDE8" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="830E91C435C570D6B5FD920254EDFED6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has unidentate subacicular hook, whereas
|
||
<taxonomicName id="33FFAF391035CAAE22C74D576CE51142" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="CD5ECF4D88F82EB6271A26F1E364F659" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5BFAE58E227AC2F703FD8CED1482F8BB" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has unidentate and a few bidentate subacicular hooks present in posterior chaetigers.
|
||
</paragraph>
|
||
<paragraph id="18FB042E48A4F8B1DD8E7E3030884587">
|
||
<taxonomicName id="E820EC0AB20331A68D8E438032E4C501" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="479A0DF2ED9057312040CD56FA06BD93" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E04FAAC3B133396B0387B58D742A6C18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="8A055F960DE3E08816CF28F3343D014E" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="33B4C75F16D5C03D852C927176428D75" italics="true">M. hongkongensa</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated by the presence or absence of eyes, number of types of pectinate chaetae, maximum number of branchial filaments, and the shape of subacicular hooks.
|
||
<taxonomicName id="C929B422C55E45F282AF756763484534" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="99FFE94E661DBD26A9AF6778D1E342BE" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CD808BF1A23DB59CD4B73491F368D382" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a pair of eyes but they are absent in
|
||
<taxonomicName id="F307D22CDFC72A5BA568B42152A07761" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="2A85A3F8A21EF1CB58EFF7BB9366D7D9" italics="true">M. hongkongensa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="6679552AF63964877A70508F18B8B9DE" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="493A14EE9AB6287688E7A9003A9E6F2A" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9308A787245A796CB90C22031A1E1A91" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has
|
||
<specimenCount id="67380BC45AF6242AA07A04A2B90415DD" count="5" type="generic" typeStatus="types">five types</specimenCount>
|
||
of pectinate chaetae (types 1, 4, 5, 6, 8) compared to
|
||
<specimenCount id="5E67BCFBE82B64B28E330442D098B8CF" count="4" type="generic" typeStatus="types">four types</specimenCount>
|
||
present in
|
||
<taxonomicName id="DCCEB71A1A4414C201D577887350975B" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="BC9A47627E795D7061FBFB7AC065C16A" italics="true">M. hongkongensa</emphasis>
|
||
</taxonomicName>
|
||
(types 1, 2, 7, 8).
|
||
<taxonomicName id="E02D5888D3532EC4B05F913CE3B866AD" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="F5F8B35B4E3FA583E540D7ED71ED9C43" italics="true">Marphysa hongkongensa</emphasis>
|
||
</taxonomicName>
|
||
lacks thick, wide isodont and thick, narrow anodont pectinate chaetae (types 4, 5, 6) which are present in the new species. The maximum number of branchial filaments in
|
||
<taxonomicName id="E75CBDE56EEF8338B408AA9308BFECDC" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="3D0ECC1BD3157CC4DDAA40E5B867BEAB" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="047828F6F250A4D2750040AFC66A33F1" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<abbrev id="ABBRID0EOFBI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
: 5.25 (3.45–5.85) mm) is six and they begin from chaetiger 24 (16–27) whereas
|
||
<taxonomicName id="525347E1CDA941660752E5F925344B85" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="46731E81D3F9C2AD572DCEC140209B1B" italics="true">M. hongkongensa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<abbrev id="ABBRID0E4FBI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
:
|
||
<quantity id="3427CBB45CDEEF586ED604742E70B5CA" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="7.0" metricValueMin="3.3" unit="mm" value="5.15" valueMax="7.0" valueMin="3.3">3.3–7 mm</quantity>
|
||
) has a maximum of ten branchial filaments, beginning from chaetiger 15–35. Finally,
|
||
<taxonomicName id="C8EAF6AD265B82B9DBC73FB393B51926" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="E6ABC9319749866AE93A3F1A24B9C476" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5541EA3BB1CB0C28FBEBB334B6C6E82E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
only has unidentate subacicular hooks while both unidentate and bidentate subacicular hooks are present in
|
||
<taxonomicName id="628027FB08F398606C6174FB4425EEE6" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="8EC8843AB2FEE4C11533D2268FEC99E5" italics="true">M. hongkongensa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DD251E85521628D11E27629148C2346F">
|
||
<taxonomicName id="20C83F587454841E64637101DC7CB6C8" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="D2EB54AFABC991086203CC3D0801BBAE" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="30B2B97085F5B2A7E4A8FBC91C936411" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="C60FC171F346666DD34DA553D1CD3519" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="9B1985D9CC628A6B6B079243A7EC82F7" italics="true">M. iloiloensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="388D07B48B7D60EBC5EB5219A396559B" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="multipectinata">
|
||
<emphasis id="EBB3680DF2D629F216800C2E26D7EC70" italics="true">M. multipectinata</emphasis>
|
||
</taxonomicName>
|
||
in having a pair of eyes. However, they can be distinguished by the number of types of pectinate chaetae present, the chaetiger on which branchiae and subacicular hooks occur, number of branchial filaments, shape of subacicular hooks and the maxillae formula.
|
||
<taxonomicName id="764F0B5D369C6DE0F5387C6619C4CD69" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="4BD42579002FDA944938F1296AD5DBE9" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0A50DC3E9E639C3FBA435D3DF0FF0A9C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has
|
||
<specimenCount id="BE7DE5296E9871FE4ECE90CFB1DB5E6C" count="5" type="generic" typeStatus="types">five types</specimenCount>
|
||
of pectinate chaetae (types 1, 4, 5, 6, 8) whereas
|
||
<taxonomicName id="1EF7433086CC5E5D17830B694D51193A" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="B0C87D0BCD961CB51B2229ADD96141B7" italics="true">M. iloiloensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="89835C820C7BC18DE696752DCA1EA23C" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="multipectinata">
|
||
<emphasis id="784EE53CDAB02A136C5A21FF20242411" italics="true">M. multipectinata</emphasis>
|
||
</taxonomicName>
|
||
have three (types 1, 4, 6) and four (types 1, 4, 7, 8) respectively.
|
||
<taxonomicName id="539D193CA66D93B5548F34254525222C" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="53465398B52946B35CFBD6AC5880E4D7" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9C640E886F8E4E5FB7B95E580C8F0B22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6815140AE69F245C9A25D3A606873E28" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="84F5FB7E8B13D0E660D1B3CBC671B359" italics="true">M. iloiloensis</emphasis>
|
||
</taxonomicName>
|
||
have the same type of pectinate branchiae (beginning on the same chaetiger with different range of variation) (chaetiger 24 (16–27) for the new species, and chaetiger 19 (16–20) for
|
||
<taxonomicName id="10279A4FC078377B319FE7369F694F20" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="1E5158EEFB93294AB4C512B2A8CDD8A1" italics="true">M. iloiloensis</emphasis>
|
||
</taxonomicName>
|
||
). The maximum number of branchial filaments in
|
||
<taxonomicName id="7EB579E7F943174CD38135145152DBEF" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="76865A54251F29F187E38D6BCB546561" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA3B2FAF968681293055C0494BD28102" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(TL: 94 (37–144) mm) is six, while
|
||
<taxonomicName id="A590A1F9EE9A8F360124C11CE54E331A" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="776176DCAC62A8FDE9D72FD911A92658" italics="true">M. iloiloensis</emphasis>
|
||
</taxonomicName>
|
||
(TL: 99 (95–165 +) mm) has a maximum of seven branchial filaments.
|
||
<taxonomicName id="A1A3FBF5E692EADBD1D8BBD0D9B2FC52" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="multipectinata">
|
||
<emphasis id="1722BC0756EFCA3F250F49E9172F0B96" italics="true">Marphysa multipectinata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<abbrev id="ABBRID0E4KBI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
:
|
||
<quantity id="714356E6DAD56C5B8E4BD2CD57FB4DD9" metricMagnitude="-2" metricUnit="m" metricValue="1.39" unit="mm" value="13.9">13.9 mm</quantity>
|
||
) has palmate branchiae with maximum of five branchial filaments from chaetiger 32. Finally, all these species have different formulae for MII, MIII and MIV (see Table
|
||
<tableCitation id="BFC78FF3FA4F59781EF40ADA3CBF22AA" captionStart="Table 6" captionStartId="T6" captionText="Table 6. Morphological features comparison between Marphysa Group B (Sanguinea) described in this study and species occurring within Malaysian water bodies (South China Sea). The features for new species are based on the holotype, with variation in parentheses for paratypes. Abbreviations: MF: maxillary formula, roman numerals refer to number of maxilla; PR-I: first peristomial ring; PR-II: second peristomial ring; p / a: present / absent; NIA: no information available. The major differences between the species are marked with asterisk (*)." httpUri="http://table.plazi.org/id/F5C127FC0BE3E3F582A959B969674F28" tableUuid="F5C127FC0BE3E3F582A959B969674F28">6</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="F5C127FC0BE3E3F582A959B969674F28" ID-Table-UUID="F5C127FC0BE3E3F582A959B969674F28" httpUri="http://table.plazi.org/id/F5C127FC0BE3E3F582A959B969674F28" startId="T6">
|
||
<paragraph id="E65DA4ADBE27BBDBC643A9D5FA467EFB">
|
||
<label id="33074F7E1FDC10F78560BDB2A4F91E88">Table 6.</label>
|
||
</paragraph>
|
||
<paragraph id="5AA4063CBA1369D1509FEEF44EE7F787">
|
||
Morphological features comparison between
|
||
<taxonomicName id="B16201BFDBB6BE506B0F547511725870" authorityName="de Quatrefages" authorityYear="1866" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="genus">
|
||
<emphasis id="70BB4A77A25F68D61478704E66948B91" italics="true">Marphysa</emphasis>
|
||
</taxonomicName>
|
||
Group B (Sanguinea) described in this study and species occurring within Malaysian water bodies (South China Sea). The features for new species are based on the holotype, with variation in parentheses for paratypes. Abbreviations:
|
||
<abbrev id="ABBRID0EVLBI" xlink_title="maxillary formula, roman numerals refer to number of maxilla">MF</abbrev>
|
||
: maxillary formula, roman numerals refer to number of maxilla;
|
||
<abbrev id="ABBRID0EZLBI" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
: first peristomial ring;
|
||
<abbrev id="ABBRID0E4LBI" xlink_title="second peristomial ring">PR-II</abbrev>
|
||
: second peristomial ring;
|
||
<abbrev id="ABBRID0EBMBI" xlink_title="present/absent">p / a</abbrev>
|
||
: present / absent;
|
||
<abbrev id="ABBRID0EFMBI" xlink_title="no information available">NIA</abbrev>
|
||
: no information available. The major differences between the species are marked with asterisk (*).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BEEC758AE7D1159724D09610693EF225">
|
||
<table id="TID0EFKBI" rules="all">
|
||
<tbody id="1F5C99EC1B7299D940174298F1DD7E35">
|
||
<tr id="A737BCE66ACF3B96E1E2E53DA2BE982E">
|
||
<th id="3BB6AEFEC411D18F2D21CA29686DAD38" colspan="1" rowspan="1">Morphological feature</th>
|
||
<th id="8A2BF2F6C7006E796ED4CEE52B6AA764" colspan="1" rowspan="1">
|
||
<taxonomicName id="331AABD20C33970D3EBD99DD2EBF816E" authority="Wang, Zhang & Qiu, 2018" authorityName="Wang, Zhang & Qiu" authorityYear="2018" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="hongkongensa">
|
||
<emphasis id="773002B0EDF021D36B4BC52C168F88D8" italics="true">M. hongkongensa</emphasis>
|
||
<bibRefCitation id="8A3D4E50EA9DF8EC4E4E185075FE1FA8" DOI="10.6620/ZS.2018.57-48" author="Wang Z & Zhang Y & Qiu JW" refId="B43" refString="Wang Z, Zhang Y, Qiu JW (2018) A new species in the Marphysa sanguinea complex (Annelida, Eunicidae) from Hong Kong. Zoological Studies 57. https: // doi. org / 10.6620 / ZS. 2018.57 - 48" year="2018">Wang, Zhang & Qiu, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="DC4B11D29055B3AF1112D4E39B8C3171" colspan="1" rowspan="1">
|
||
<taxonomicName id="606CF5D5423BD87C1067D6DE1E1BB4CA" authority="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings, 2019" authorityName="Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings" authorityYear="2019" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="iloiloensis">
|
||
<emphasis id="B06C995A5B20DA5EDBEBD9916F511653" italics="true">M. iloiloensis</emphasis>
|
||
<bibRefCitation id="A90A8ABE4E3D6E0084F28BD973B188E0" DOI="10.11646/zootaxa.4674.2.7" author="Glasby CJ & Mandario MAE & Burghardt I & Kupriyanova E & Gunton LM & Hutchings P" issue="2" journalOrPublisher="Zootaxa" pagination="264-282" refId="B14" refString="Glasby CJ, Mandario MAE, Burghardt I, Kupriyanova E, Gunton LM, Hutchings P (2019) A new species of the sanguinea-group Quatrefages, 1866 (Annelida: Eunicidae: Marphysa) from the Philippines. Zootaxa 4674 (2): 264 - 282. https: // doi. org / 10.11646 / zootaxa. 4674.2. 7" title="A new species of the sanguinea-group Quatrefages, 1866 (Annelida: Eunicidae: Marphysa) from the Philippines." volume="4674" year="2019">Glasby, Mandario, Burghardt, Kupriyanova, Gunton & Hutchings, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="6DA15E38EB560082F07B05D5C8A1343A" colspan="1" rowspan="1">
|
||
<taxonomicName id="0910868F67F03EA1B96AD8FA41DC4F4C" authority="Liu, Hutchings & Sun, 2017" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="multipectinata">
|
||
<emphasis id="CA4236E802AED6EF257F773C18B85B2B" italics="true">M. multipectinata</emphasis>
|
||
<bibRefCitation id="250DD4F2763EB59D927FE40BA97CCEE8" DOI="10.11646/zootaxa.4263.2.2" author="Liu Y & Hutchings P & Sun S-C" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228 - 250. https: // doi. org / 10.11646 / zootaxa. 4263.2. 2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu, Hutchings & Sun, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="CABBC44823B898796AA64BF48312F97A" colspan="1" rowspan="1">
|
||
<taxonomicName id="14AD3372F8FBC35A17511BC6E9E73050" authority=", Treadwell, 1936" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="390338F2F65A8E191838078BC49A8C29" italics="true">M. orientalis</emphasis>
|
||
,
|
||
<bibRefCitation id="C7FE7D1A80CAC3B1EF2A5131813209BE" DOI="10.5479/si.00963801.83-2984.261" author="Treadwell AL" issue="2984" journalOrPublisher="Proceedings of the United States National Museum" pagination="261-278" refId="B41" refString="Treadwell AL (1936) Polychaetous annelids from Amoy, China. Proceedings of the United States National Museum 83 (2984): 261 - 278. https: // doi. org / 10.5479 / si. 00963801.83 - 2984.261" title="Polychaetous annelids from Amoy, China." volume="83" year="1936">Treadwell, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="E5ACE2904952256C9A3DE45AD1E6830E" colspan="1" rowspan="1">
|
||
<taxonomicName id="02C9CCF088B759FC73696203C8A2855D" authority="Liu, Hutchings & Sun, 2017" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="8E01C4DDC842CAD164EA440999C55993" italics="true">M. tribranchiata</emphasis>
|
||
<bibRefCitation id="B3FA422485F7E4990DFB73C8D7962B9A" DOI="10.11646/zootaxa.4263.2.2" author="Liu Y & Hutchings P & Sun S-C" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228 - 250. https: // doi. org / 10.11646 / zootaxa. 4263.2. 2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu, Hutchings & Sun, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="BEE333BFBBBDB39827516B3AB3AA046E" colspan="1" rowspan="1">
|
||
<taxonomicName id="82038C1501A094CB69CAD622D2D03CCC" authority="Liu, Hutchings & Sun, 2017" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="2D892A5A880E1D5F378AD44F1C1F6549" italics="true">M. tripectinata</emphasis>
|
||
<bibRefCitation id="0DBABE05279EC4A5C5496C1E33A62C13" DOI="10.11646/zootaxa.4263.2.2" author="Liu Y & Hutchings P & Sun S-C" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228 - 250. https: // doi. org / 10.11646 / zootaxa. 4263.2. 2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu, Hutchings & Sun, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="1D344754DC37DF31A73939C0EC0B7F1B" colspan="1" rowspan="1">
|
||
<taxonomicName id="870D06A917B1012CC411C32C69D3FCAB" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="AFFCE86D2E2E9D7D1F1EE6887DDC787F" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="03DC35707394E428A3F9CA24A1B3D6DA" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="0D9048A2CF1282303ED8D4E0D1203066" colspan="1" rowspan="1">
|
||
<taxonomicName id="C686E682011D35906A9B0E03586CD17D" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="setiuense" status="sp. nov.">
|
||
<emphasis id="9369C6EFB7D1996056664925DA1A5620" italics="true">M. setiuense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CB93D42B88456E88269F45BD728AA675" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr id="FDB59351D521069AE559EED4AE9F0D4D">
|
||
<td id="8859578BC3A67698E29326EA483BDACF" colspan="1" rowspan="1">Source of Information</td>
|
||
<td id="26FED6C0AAA1CF3D70E9094EA28586ED" colspan="1" rowspan="1">
|
||
Holotype: SWIMS-ANN- 18-012; Paratypes: SWIMS-ANN- 18-013 - (
|
||
<bibRefCitation id="CF5356C033467BBEF7C31CD8B01B709F" DOI="10.6620/ZS.2018.57-48" author="Wang" etAl="et al." firstAuthor="Wang" refId="B43" refString="Wang Z, Zhang Y, Qiu JW (2018) A new species in the Marphysa sanguinea complex (Annelida, Eunicidae) from Hong Kong. Zoological Studies 57. https://doi.org/10.6620/ZS.2018.57-48" year="2018">Wang et al. 2018</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="3BBF664B10787AC6C80F24CB4ADBA651" colspan="1" rowspan="1">
|
||
Holotype: NTM W 29624; Paratypes: NTM W 29619 – NTM 29623 (
|
||
<bibRefCitation id="8DB5D6E8CD540FB3DBCDCD9FD3A8C96A" DOI="10.11646/zootaxa.4674.2.7" author="Glasby" etAl="et al." firstAuthor="Glasby" issue="2" journalOrPublisher="Zootaxa" pagination="264-282" refId="B14" refString="Glasby CJ, Mandario MAE, Burghardt I, Kupriyanova E, Gunton LM, Hutchings P (2019) A new species of the sanguinea-group Quatrefages, 1866 (Annelida: Eunicidae: Marphysa) from the Philippines. Zootaxa 4674 (2): 264–282. https://doi.org/10.11646/zootaxa.4674.2.7" title="A new species of the sanguinea-group Quatrefages, 1866 (Annelida: Eunicidae: Marphysa) from the Philippines." volume="4674" year="2019">Glasby et al. 2019</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="C86CA898215ACC4270758E81CF5BB588" colspan="1" rowspan="1">
|
||
Holotype: ASIZW 0000345-1 (
|
||
<bibRefCitation id="CF6B66A119DDD296BE9F0A3054A54D8C" DOI="10.11646/zootaxa.4263.2.2" author="Liu" etAl="et al." firstAuthor="Liu" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228–250. https://doi.org/10.11646/zootaxa.4263.2.2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu et al. 2017</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="65D90A257D5A7AEFA4D949333A08FE36" colspan="1" rowspan="1">
|
||
Type: USNM. No. 20114 (
|
||
<bibRefCitation id="02822C18029658A0C08AD331C341A7AA" DOI="10.5479/si.00963801.83-2984.261" author="Treadwell" firstAuthor="Treadwell" issue="2984" journalOrPublisher="Proceedings of the United States National Museum" pagination="261-278" refId="B41" refString="Treadwell AL (1936) Polychaetous annelids from Amoy, China. Proceedings of the United States National Museum 83 (2984): 261–278. https://doi.org/10.5479/si.00963801.83-2984.261" title="Polychaetous annelids from Amoy, China." volume="83" year="1936">Treadwell 1936</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="9F9271E9747A5B9417689B5021E18E7D" colspan="1" rowspan="1">
|
||
Holotype: ASIZW 0000348-2 (
|
||
<bibRefCitation id="C1CC2E10549A9C86C2263E93E252AE0B" DOI="10.11646/zootaxa.4263.2.2" author="Liu" etAl="et al." firstAuthor="Liu" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228–250. https://doi.org/10.11646/zootaxa.4263.2.2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu et al. 2017</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="A3B2177BB42472DF4428F40996BB3E9C" colspan="1" rowspan="1">
|
||
Holotype:
|
||
<named-content id="85B440BE58F5343739F91D15724FC4EB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/australian-museum" xlink_title="Australian Museum, Sydney, Australia">AM</named-content>
|
||
W. 49069 (
|
||
<bibRefCitation id="53A407DFC4C4A5B12034229C68B33AED" DOI="10.11646/zootaxa.4263.2.2" author="Liu" etAl="et al." firstAuthor="Liu" issue="2" journalOrPublisher="Zootaxa" pagination="228-250" refId="B25" refString="Liu Y, Hutchings P, Sun S-C (2017) Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934. Zootaxa 4263 (2): 228–250. https://doi.org/10.11646/zootaxa.4263.2.2" title="Three new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from the south coast of China and redescription of Marphysa sinensis Monro, 1934." volume="4263" year="2017">Liu et al. 2017</bibRefCitation>
|
||
)
|
||
</td>
|
||
<td id="D5BA131DF57FC3408A4A5D64E955EC21" colspan="1" rowspan="1">Holotype: UMTAnn 2149 (this study)</td>
|
||
<td id="89E7213E9933A013E578D9BA17E1931F" colspan="1" rowspan="1">Holotype: UMTAnn 2177 (this study)</td>
|
||
</tr>
|
||
<tr id="0F5D4816C516329A27D8B19DA530A054">
|
||
<td id="CAB0FFBEB2BDFE47FE5BCBBECEC37472" colspan="1" rowspan="1">
|
||
Size (mm):
|
||
<abbrev id="ABBRID0ENSBI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
,
|
||
<abbrev id="ABBRID0ERSBI" xlink_title="width at chaetiger 10">W 10</abbrev>
|
||
</td>
|
||
<td id="2548815A90921F57BE0087F535892DCC" colspan="1" rowspan="1">3.3–7.0, 2.2–5.3</td>
|
||
<td id="B9A752B5D0FFC508A88EC59D5789728A" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E2SBI" xlink_title="no information available">NIA</abbrev>
|
||
, 2.6
|
||
</td>
|
||
<td id="FE2B53765AB8C93CFFCF6063D94F0815" colspan="1" rowspan="1">13.9, 5.7</td>
|
||
<td id="76DA0EFBADA42E6D33BAE43273141CEC" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EGTBI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="7B9D91BE6FDDA6F042556E1204F07501" colspan="1" rowspan="1">8.7, 3.9</td>
|
||
<td id="BBA9C182EC508C9A43347FB1DE6DA6B2" colspan="1" rowspan="1">12.7, 5.95</td>
|
||
<td id="BF9290832FCDC7944E0D9BC6F08B8E8C" colspan="1" rowspan="1">5.25 (3.45–5.85), 2.85 (1.95–3.15)</td>
|
||
<td id="21E7A704E88FFED6F8927E371FF6A4E7" colspan="1" rowspan="1">2.7 (2.85–4.8), 1.8 (1.65–2.55)</td>
|
||
</tr>
|
||
<tr id="0DC0B695FAAC4A3311EC0D0EBCD55FE6">
|
||
<td id="9E78100C3209BDE9A2E96D84CF48B9B9" colspan="1" rowspan="1">Prostomium: shape</td>
|
||
<td id="E86662C2E12F5AA03FCA474431979F7A" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="4511989D73F62D4A70A46855E15FB285" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="AB813F2620EFF1349AAE3DEC24CEDD63" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="C7B764BB28AB90D6326E05F2F840CA3F" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="B30331688052635C93DF43C60D2E5C15" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="B3EAC99756EEB70CD9A403796963D057" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="22FA6C60E5EEE872710398EEA02F6359" colspan="1" rowspan="1">Bilobed</td>
|
||
<td id="72275F16C5C6C16F9D4372B4B85E4C74" colspan="1" rowspan="1">Bilobed</td>
|
||
</tr>
|
||
<tr id="9BB58DF4794C947EC9E7B8AAA9881E4B">
|
||
<td id="89E5A941C05667ED27EBF7C70CF8A5B1" colspan="1" rowspan="1">Palps: reaching</td>
|
||
<td id="20227E0CD54247795B878B59678B73D4" colspan="1" rowspan="1">Chaetiger 1</td>
|
||
<td id="58A98851F0D4D126CB4CF44B0F69183F" colspan="1" rowspan="1">Chaetiger 1</td>
|
||
<td id="2D8E989AD092EB76F37B4948B1A15642" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EAVBI" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
<td id="25C07423AD8FBF8A53B2C0116BDCADEF" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EIVBI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="46403AC8A0C4DC275EFB7128BC0A018F" colspan="1" rowspan="1">Chaetiger 1</td>
|
||
<td id="9661419203B78AA3D4354F8A9049D5D4" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ETVBI" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
<td id="953BF53E42FE4AC9E0A9876899E6C528" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E2VBI" xlink_title="second peristomial ring">PR-II</abbrev>
|
||
</td>
|
||
<td id="D96F6D0E5C30B7BC0B65132949AE6475" colspan="1" rowspan="1">Chaetiger 3</td>
|
||
</tr>
|
||
<tr id="E3B73988953E9F361A91A1E7B9B7F51E">
|
||
<td id="A8D305A0C040AAF486AD22867286C06F" colspan="1" rowspan="1">Lateral antennae: reaching</td>
|
||
<td id="E90E0F1ED233F8A019CFE525D38817D2" colspan="1" rowspan="1">Chaetiger 1</td>
|
||
<td id="9CBD094B9046C556649E13D8C3960E21" colspan="1" rowspan="1">Chaetiger 1 or 2</td>
|
||
<td id="7B40CBCEA7CAE198EE736E0D18DBA4C6" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EQWBI" xlink_title="first peristomial ring">PR-I</abbrev>
|
||
</td>
|
||
<td id="2446BBAD5884837E891144CE3A6CF819" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EYWBI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="0E7D773A053B8547BEFD4E5D28E4982F" colspan="1" rowspan="1">Chaetiger 2</td>
|
||
<td id="F5DE13B558FED0786DAFC4414BA1233B" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EDXBI" xlink_title="second peristomial ring">PR-II</abbrev>
|
||
</td>
|
||
<td id="F941FA5FA0DE8794BDC043C4AFB33F6B" colspan="1" rowspan="1">Chaetiger 2</td>
|
||
<td id="A4E892C3A6218CDE35D3EB0433FB6914" colspan="1" rowspan="1">Chaetiger 4</td>
|
||
</tr>
|
||
<tr id="0E29765D55A457624EF3E483C86CA33C">
|
||
<td id="4E5E71FD7C4EFF675E797A288EF7415B" colspan="1" rowspan="1">Median antennae: reaching</td>
|
||
<td id="12F832D67EB1BFD7D4CAAAA0F46872B8" colspan="1" rowspan="1">Chaetiger 1 or 2</td>
|
||
<td id="F6FD875DFF4C3409410D2CE9C22E6011" colspan="1" rowspan="1">Chaetiger 1 or 2</td>
|
||
<td id="A3826AE862D2CC33B0ADB625954B26D1" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E2XBI" xlink_title="second peristomial ring">PR-II</abbrev>
|
||
</td>
|
||
<td id="CEB8D2DADF5F300B3B1ED2BAE9F1620C" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EDYBI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="4DE83587A35B57B9CF5D1EFA6B104AD8" colspan="1" rowspan="1">Chaetiger 3</td>
|
||
<td id="1926EBB860AB1A33B1A3FCA4C12F4D6A" colspan="1" rowspan="1">Chaetiger 1</td>
|
||
<td id="3885EC576894A94B680473FE9F80DA2F" colspan="1" rowspan="1">Chaetiger 3</td>
|
||
<td id="84C396064AE6BA54E63BD96AF2F60582" colspan="1" rowspan="1">Chaetiger 5</td>
|
||
</tr>
|
||
<tr id="CB8051ACD12F1DD4EAD23C3D7AC02AE6">
|
||
<td id="DACB9F5F90C0B2B545A2105727205BD9" colspan="1" rowspan="1">Peduncle in prostomial appendages</td>
|
||
<td id="CE8659CC774D39A8596C00CAECFAA593" colspan="1" rowspan="1">Absent</td>
|
||
<td id="49E8E0802370ABAAF7BD87A3C19729C6" colspan="1" rowspan="1">Absent</td>
|
||
<td id="FA1B39970C534E0E09A4C0D9A574319F" colspan="1" rowspan="1">Absent</td>
|
||
<td id="60DCDC2E224A33A21DD5031B20CCF6F2" colspan="1" rowspan="1">Absent</td>
|
||
<td id="641C5DBEFDDDB746488B75A963F64545" colspan="1" rowspan="1">Absent</td>
|
||
<td id="E10E15D09BB07204EC2BF9336C97B7E9" colspan="1" rowspan="1">Absent</td>
|
||
<td id="2D7F091989CFD5206ACC8D42EE7C066A" colspan="1" rowspan="1">Absent</td>
|
||
<td id="B87109E070F9D1DFF2C8483EBBB23466" colspan="1" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr id="737278C4E92918AD9B0EDC133D4B7A1E">
|
||
<td id="8CE2EC5D805F8435DD0CC37DF6863C0F" colspan="1" rowspan="1">Eyes *</td>
|
||
<td id="3EC61EE5075E096D4F79FD65721149B6" colspan="1" rowspan="1">Absent</td>
|
||
<td id="780ED095DCB13FAE47426448EC96E48A" colspan="1" rowspan="1">Present</td>
|
||
<td id="46F92F57F0FC9BFDD534E5B610AB70B5" colspan="1" rowspan="1">Present</td>
|
||
<td id="5F419116128DA966DFFCEEC503B22FEC" colspan="1" rowspan="1">Absent</td>
|
||
<td id="15FF38EBCA32E7004325BC8FA51C1C08" colspan="1" rowspan="1">Absent</td>
|
||
<td id="CB5CBA18233E16C46C69CD7547C795A1" colspan="1" rowspan="1">Absent</td>
|
||
<td id="1C19071787F0C71571D638EDF8158D64" colspan="1" rowspan="1">Present</td>
|
||
<td id="7C26B721704B748989FAB247F225BE6C" colspan="1" rowspan="1">Present</td>
|
||
</tr>
|
||
<tr id="634FC090EF07911F3C26C28DE781B8C9">
|
||
<td id="AAC2C09CC2A76CBDB2E96805A2FD2319" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EQ1BI" xlink_title="maxillary formula, roman numerals refer to number of maxilla">MF</abbrev>
|
||
: MII, MIII, MIV *
|
||
</td>
|
||
<td id="FF47E7ABD6C4B1A070B9A29457A401C2" colspan="1" rowspan="1">5-6 + 5 – 6, 7 + 0, 4 + 8</td>
|
||
<td id="AF04C930621E89C75699431629274A44" colspan="1" rowspan="1">4 + 5, 4–5 + 0, 3–4 + 5 – 6</td>
|
||
<td id="06BBD4B23A277B491FFE0110DA3B0BDC" colspan="1" rowspan="1">3 + 3, 4 + 0, 4 + 5</td>
|
||
<td id="396B19B789C7E20CFA8FA40F2786ADF2" colspan="1" rowspan="1">3 + 3, 4 + 0, 4 + 3</td>
|
||
<td id="8C335C6AF82A0F68CA281F32F8B62DD5" colspan="1" rowspan="1">4 + 4, 5 + 0, 4 + 8</td>
|
||
<td id="6ED24DF981ACFF0D2D2616F8EB03E7DE" colspan="1" rowspan="1">5 + 5, 5 + 0, 4 + 8</td>
|
||
<td id="AF4582CE1E34E12418BBF93817A95290" colspan="1" rowspan="1">5 (4–5) + 5 (5–6), 7 (6–7) + 0, 4 (4–5) + 8 (5–8)</td>
|
||
<td id="BC785A02CC0216883F3F299D23461503" colspan="1" rowspan="1">5 (4–5) + 5 (4–6), 5 (4–6) + 0, 3 (3–4) + 6 (7–8)</td>
|
||
</tr>
|
||
<tr id="71C99563262872B9A4F72DBF542AFB8E">
|
||
<td id="A6F09EB5982726EEC9E071C2EB390D6C" colspan="1" rowspan="1">Branchiae: shape</td>
|
||
<td id="3E947203CB4D6557D03CA34F6722480E" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="128E909E46489F1B7CC95B690BB0B4BB" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="1FF53512C6CA5644489E565E6D173F3B" colspan="1" rowspan="1">Palmate</td>
|
||
<td id="A5F56BEA4AF0C6D2FD76F3CBF3C3EFC9" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E42BI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="5C6F2C053B122CA4AA194961ED305D84" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="2EAD0827D7F8FC90D36DA8A5923E8C39" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="4FC5AC759D43D0362BB86C75A5AFA74C" colspan="1" rowspan="1">Pectinate</td>
|
||
<td id="0B2CEF1DA61BE05564B851F13A9B5369" colspan="1" rowspan="1">Pectinate</td>
|
||
</tr>
|
||
<tr id="EE332F0F8AEACF09E99693DEC5967A71">
|
||
<td id="28C758A51CC7A9C398A0F360C69AC85A" colspan="1" rowspan="1">Branchiae: start chaetiger; last chaetiger before pygidium</td>
|
||
<td id="24A85C8E98555D22A38B1C656B1573E6" colspan="1" rowspan="1">15–35; until pygidium</td>
|
||
<td id="7AEDBC48082AD7F755C06E9297378072" colspan="1" rowspan="1">16–20; until pygidium</td>
|
||
<td id="AC0902DCD8712BC21912356C5228D306" colspan="1" rowspan="1">32; end at chaetiger 281</td>
|
||
<td id="50098397FDE92350567307FC803EFC5D" colspan="1" rowspan="1">45; end ~ 30 last chaetiger</td>
|
||
<td id="76CDE75C24024FAB322AA69BE7992E52" colspan="1" rowspan="1">26; end at chaetiger 181</td>
|
||
<td id="813E254C51E23E3BD0420D0118822BA1" colspan="1" rowspan="1">15; end at chaetiger 399</td>
|
||
<td id="6DA2FAEDE0188A9FB34586A1E1433049" colspan="1" rowspan="1">24 (16–27); end ~ 10 last chaetiger</td>
|
||
<td id="54A51471E9761731165D323B879FCBF0" colspan="1" rowspan="1">20 (15–25); until pygidium</td>
|
||
</tr>
|
||
<tr id="E8929C75A8C3EB13DFDA3E08A86DE7D5">
|
||
<td id="06FA60A572827A22558B26F2C58A78F0" colspan="1" rowspan="1">Branchial filaments: numbers</td>
|
||
<td id="46DFDC74AE8167216AA111F892BA7F79" colspan="1" rowspan="1">5–10</td>
|
||
<td id="5620E57289F0FAC3F41455AD4ABEE17E" colspan="1" rowspan="1">6–7</td>
|
||
<td id="8ECFF9042C8D77D06E6BA91ECC3ACE5F" colspan="1" rowspan="1">5</td>
|
||
<td id="C5F35A40D211E3C4FC7EA0E667FD365E" colspan="1" rowspan="1">3</td>
|
||
<td id="A931CFE0D184835034B0AB06B46B7C93" colspan="1" rowspan="1">3</td>
|
||
<td id="35F77D42DC01DE187651C45F2EE092E4" colspan="1" rowspan="1">8</td>
|
||
<td id="87D98CF4B5F51FF684860B7C569CAD59" colspan="1" rowspan="1">6</td>
|
||
<td id="7E9A33D718549AFE4639F78750238480" colspan="1" rowspan="1">5</td>
|
||
</tr>
|
||
<tr id="17EAD0A06629AD5D6AEB7C9A8FDD0346">
|
||
<td id="A9717366F7D67CE1EC0C27B66446ECF6" colspan="1" rowspan="1">Dorsal cirri: shaped</td>
|
||
<td id="6C4D45478F805DD0EC11AE3C38D580FB" colspan="1" rowspan="1">Conical</td>
|
||
<td id="3EA8FA9A83254BEEB1B81E32851B2805" colspan="1" rowspan="1">Conical</td>
|
||
<td id="3EB6B0E9FB3B0DACCF34CA6784C76AA2" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ET5BI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="988A5F2FC5AEAC2797233D67B42E0516" colspan="1" rowspan="1">Conical</td>
|
||
<td id="EAC449029F2BF8FD3F86B2242E1C363B" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E55BI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="54607A2606A1420DE2BEACF484478BC3" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EG6BI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="C93304624EF2F2632AF921404FFE2E59" colspan="1" rowspan="1">Digitiform</td>
|
||
<td id="762334C5D1F2D161350017680F371EA0" colspan="1" rowspan="1">Thumb-shape, digitiform</td>
|
||
</tr>
|
||
<tr id="595CAC77A45881BBE972E0D284BC1018">
|
||
<td id="B8E88DB9C902F82A63BE538C9D8A1A84" colspan="1" rowspan="1">Prechaetal lobe: shaped</td>
|
||
<td id="AE4E8820A9F844FD71F7251D842CEA6D" colspan="1" rowspan="1">Transverse fold</td>
|
||
<td id="6C0F56B7AB1EA006C92D2642A8DBB234" colspan="1" rowspan="1">Transverse fold</td>
|
||
<td id="9E9F5184333D01BB18AC7CC47D3CB27C" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E56BI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="0497F1E1EF1B46399EDA186E9CDEB1F9" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EGACI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="57666760CE2F29884154F8ED2A2B4010" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EOACI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="8BBD1ED883D207F023A2978C3A485A18" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EWACI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="731159502FF83FF94E19F70B7D4B7FDE" colspan="1" rowspan="1">Transverse fold</td>
|
||
<td id="B47826DDCA2C4CEFCFDF2406B7A7A891" colspan="1" rowspan="1">Transverse fold</td>
|
||
</tr>
|
||
<tr id="BA857357F38A9070169472997A7B752E">
|
||
<td id="E05BC489CEBA2997104790A8BA439758" colspan="1" rowspan="1">Chaetal lobe: shaped</td>
|
||
<td id="6CD8CC94A995377D67496AAA4073D062" colspan="1" rowspan="1">Rounded</td>
|
||
<td id="06F80CCFF60C1CBBF51AEC7D7283F3C6" colspan="1" rowspan="1">Rounded</td>
|
||
<td id="3F66FCE9BC1C3B352D054679BD143D93" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EOBCI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="18E7292314CFB5988FFBF984B0FEA4F9" colspan="1" rowspan="1">Rounded</td>
|
||
<td id="83B3AA9B6A899A161EEE9734A6FD4259" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EZBCI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="85E561307153067D9859ABA1D2BD8085" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EBCCI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="4AF0129BC74FA71C4145A4398B7AB7DA" colspan="1" rowspan="1">Rounded and conical</td>
|
||
<td id="720158BC69755F6E20BAA2B5E2D16641" colspan="1" rowspan="1">Rounded</td>
|
||
</tr>
|
||
<tr id="1D52F1B1D6AFA3B6B7C6640D3DDD6D50">
|
||
<td id="D01122F53D3BCAC3687D291EA0DB825A" colspan="1" rowspan="1">Aciculae: shape; colour</td>
|
||
<td id="57FE3D9A0BE7FA6704C843619CF0FB52" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ETCCI" xlink_title="no information available">NIA</abbrev>
|
||
; black with paler tips
|
||
</td>
|
||
<td id="392B24FEBD0A49043A3C197A27D996A0" colspan="1" rowspan="1">Blunt; black with paler tips</td>
|
||
<td id="9F6F955068F4A3570997C2AE66BE828E" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0E5CCI" xlink_title="no information available">NIA</abbrev>
|
||
; brown
|
||
</td>
|
||
<td id="33959D0478F8A51A4E105C9E1EE00947" colspan="1" rowspan="1">Blunt; black</td>
|
||
<td id="79A18DC431C463BDADD42BE3610FD500" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EJDCI" xlink_title="no information available">NIA</abbrev>
|
||
; brown
|
||
</td>
|
||
<td id="5580BFF7634F51C4BAE3048D0EA6C2A3" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ERDCI" xlink_title="no information available">NIA</abbrev>
|
||
; black
|
||
</td>
|
||
<td id="63E9BC51E88B6CEA4B2CF77EC7C62B55" colspan="1" rowspan="1">Blunt; black and translucent at distal end</td>
|
||
<td id="3494CE5A53846B1552FB6C5C6D1103D6" colspan="1" rowspan="1">Blunt; black and translucent at distal end</td>
|
||
</tr>
|
||
<tr id="FBEA30D16889DBBA83DE7A2B2BF7A364">
|
||
<td id="55256C680FAEE8C009D79A124289E792" colspan="1" rowspan="1">
|
||
Subacicular limbate chaetae: (
|
||
<abbrev id="ABBRID0EAECI" xlink_title="present/absent">p / a</abbrev>
|
||
); distribution
|
||
</td>
|
||
<td id="7EF6ACE5C443583B53FF7CD3CBF928FF" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="ABDD0192DEC6B780EF55A73302BBB941" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="F0B84F47D463B8BFB9293950C477BD2E" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="94F35A1E347AB518A76025A240308534" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="6017AC7E3FEDB1D6BE973BCDC600363D" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="96635CC094A59642DF88C19BAC25C382" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="C5AC08D071F16E3241118F4F7BE9FA4E" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
<td id="EF27EB5120A8F3D5A89D64317E223AEA" colspan="1" rowspan="1">Absent; all chaetigers</td>
|
||
</tr>
|
||
<tr id="E6A3BB0BFF3F03AC4CCD973B4625F85A">
|
||
<td id="00EA0E2DEC0755C15FE9C8B67F0DB160" colspan="1" rowspan="1">Pectinate chaetae: number of type *</td>
|
||
<td id="9DCE1A452CD09C59988C3A7F756E5279" colspan="1" rowspan="1">4</td>
|
||
<td id="FB53E9C4459CC582D5012CD5EA49C635" colspan="1" rowspan="1">3</td>
|
||
<td id="FADD75D39F4A85812F0EEE1E04183AE5" colspan="1" rowspan="1">4</td>
|
||
<td id="C354DAB23DFDCFD712D0567C48EB8D13" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0ENFCI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="86FE5ED7C5F06D3B33F01DBF13519841" colspan="1" rowspan="1">3</td>
|
||
<td id="1FFA4A583F8EB0537C1A9288B7CC539A" colspan="1" rowspan="1">3</td>
|
||
<td id="A7A5FD41FF17E4541D51118BD4759831" colspan="1" rowspan="1">5</td>
|
||
<td id="BFF076958AE402CCCD7FC5EBCAD98589" colspan="1" rowspan="1">4</td>
|
||
</tr>
|
||
<tr id="6C2B9E33A4D0D81BB859BA7C8347A330">
|
||
<td id="820D6384A547AFA592FA2CAE8F7248EA" colspan="1" rowspan="1">Subacicular hook: shape; colour *</td>
|
||
<td id="7CC6541366E6C78EBC7DCB0068ED3D21" colspan="1" rowspan="1">Unidentate and bidentate; amber</td>
|
||
<td id="8F8D2EA8E9709FEA6037DD9C12A04CB4" colspan="1" rowspan="1">Unidentate; amber to black</td>
|
||
<td id="A672B060ACEEF423043DBA293BEA327E" colspan="1" rowspan="1">Unidentate and bidentate; yellow</td>
|
||
<td id="53AAC4A15415D299093BE77E68D0CD31" colspan="1" rowspan="1">
|
||
Unidentate;
|
||
<abbrev id="ABBRID0EOGCI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="BF205B69F4CD595002F572C317D86D63" colspan="1" rowspan="1">Unidentate and bidentate; brown</td>
|
||
<td id="9520C3E7035E6696E5B9E67F79F71BE4" colspan="1" rowspan="1">Unidentate</td>
|
||
<td id="BEEB470BE4FFC1E39E303569D1444C50" colspan="1" rowspan="1">Unidentate; light brown and translucent at distal end</td>
|
||
<td id="69EC29D2CF3937B4AD9427E9C0853CE7" colspan="1" rowspan="1">Unidentate and bidentate; light brown and translucent at distal end</td>
|
||
</tr>
|
||
<tr id="2C40C529868BEA0A249C834C1F25FEBE">
|
||
<td id="31C3AF04AAEF0EAD002A1D2B68EAEC4F" colspan="1" rowspan="1">Subacicular hook: start chaetiger *</td>
|
||
<td id="BC47B44C9529FD74931C8D3545941901" colspan="1" rowspan="1">26–58</td>
|
||
<td id="38CB50B98CB49C3535BED4E12BA82C04" colspan="1" rowspan="1">30–38</td>
|
||
<td id="0533DD5144F1A97992130519FA5006A8" colspan="1" rowspan="1">20</td>
|
||
<td id="00F4979109D36DE8DC4DEF7952A8F903" colspan="1" rowspan="1">Present in posterior region (No information on start chaetiger)</td>
|
||
<td id="6BDEA09E90B437F61D96810F8FC46ABA" colspan="1" rowspan="1">20</td>
|
||
<td id="ED8B9763496292CEB4C2C225A40FA40F" colspan="1" rowspan="1">170</td>
|
||
<td id="995F646D3EA9B3657D245147F4963BF1" colspan="1" rowspan="1">37 (26–42)</td>
|
||
<td id="518599DC1F1A60A1A33EFB01C1635C6E" colspan="1" rowspan="1">25 (21–38)</td>
|
||
</tr>
|
||
<tr id="C80D86C06486D5D8AB449D840BD6AD7B">
|
||
<td id="4AF08DAAED7C0D0D9273AFFD960B809E" colspan="1" rowspan="1">Subacicular hook: distribution</td>
|
||
<td id="2E8DE059F98F4873F3DD37ACAAE57873" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="9A8A7608C96F2696DD46FCE3C4B6ED5F" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="7F0836EBDA551EC56D2F98BEB52E8187" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="370E36F2F1D7A79921F30AB07D6B3929" colspan="1" rowspan="1">
|
||
<abbrev id="ABBRID0EKICI" xlink_title="no information available">NIA</abbrev>
|
||
</td>
|
||
<td id="2B8D81C6600616335122264CFE738436" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="9548E1EF5B7ACB2CE005978310269130" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="6BC0CE7A841F7141584593BB91E1FE65" colspan="1" rowspan="1">Continuous</td>
|
||
<td id="86B5132902CBF4D7662547539E8C3DB3" colspan="1" rowspan="1">Continuous</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="60BD4E0D8F980BBAAA2A07F8F04587AC">
|
||
The other species from the Sanguinea complex,
|
||
<taxonomicName id="AF4DA9363648EF9A53889789D844D3EB" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="5F914D869C80679D5C48DB64E1AF6184" italics="true">M. tribranchiata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4F0846C9F743BEE871A016CE14C6D40F" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="9CE6BDC0BC22A743AE7E603168D98B07" italics="true">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="1D2FE050B56533CD9DA47377800129C9" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="C7EBCA343B031970C7E634A93CCDEA72" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7C69EAE66220F34C7B5C0B6469496433" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by the absence of eyes. Both
|
||
<taxonomicName id="1171706156BA7CA4AF5531C22229D821" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="90263318CF8279C091F029525591122C" italics="true">M. tribranchiata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="693ADC93C6FD6010E6D436AD7A51584B" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="040744DB75CEDB53FC350FA22E455BCB" italics="true">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
have
|
||
<specimenCount id="9B43192C8F0F4C35659C0F1EEB1C84E7" count="3" type="generic" typeStatus="types">three types</specimenCount>
|
||
of pectinate chaetae, whereas
|
||
<taxonomicName id="E03BFB1B82D15D0DA3509431E74F9EC7" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="F3E3D5D9862094511737B34B73C7768D" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="192F431623B47232977D34D16ADACAB6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has
|
||
<specimenCount id="F4E572CF0A657B789AE162D85D4F47CA" count="5" type="generic" typeStatus="types">five types</specimenCount>
|
||
.
|
||
<taxonomicName id="A2EE1BABDCFBAF781D07C110E525A181" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="83D54EC6819775532DF603A1CE9D57CC" italics="true">Marphysa tribranchiata</emphasis>
|
||
</taxonomicName>
|
||
lacks thick, wide isodont and thick, narrow anodont pectinate chaetae (types 4, 6), while
|
||
<taxonomicName id="8C2146DCC2CEDAC19EAA792C92B245E1" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="7E990B766E79565BCC9825AD79045426" italics="true">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
lacks thin, narrow isodont pectinate chaetae (type 1) which are present in the new species (types 1, 4, 5, 6, 8). While
|
||
<taxonomicName id="D25069594E3ED886A8B7F3D61FD2980B" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="2B225FC0C4278D7046508B7D58731AF2" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="943EDB98561CD1A88A2AD2BFE5630CE6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6F15E6A464CD919895900FE91E2162C7" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="287F6AF77344E748FC5C9C667DE23653" italics="true">M. tripectinata</emphasis>
|
||
</taxonomicName>
|
||
only have unidentate subacicular hooks, they begin much later (chaetiger 170) in the latter species.
|
||
<taxonomicName id="77D34E1A2B7267A5BB3A5412153B746B" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="D77D5DCAD4872B918FEB38180C3A1308" italics="true">Marphysa tribranchiata</emphasis>
|
||
</taxonomicName>
|
||
has both unidentate and bidentate subacicular hooks whereas only unidentate hooks are present in
|
||
<taxonomicName id="526476B35141D3501C6F54AFF7B1D618" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="20FC97F3A4C9CE71AEE669946A9CE648" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0FE0F2E1E189D3BECADE0EECD0A43B0F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The maximum number of branchiae filaments present in
|
||
<taxonomicName id="DD37DE9C5030C1C70C81D35DEB97CBA4" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tribranchiata">
|
||
<emphasis id="A3BC004DDCA4B5F3F15B145A7FDD3834" italics="true">M. tribranchiata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<abbrev id="ABBRID0ELNCI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
:
|
||
<quantity id="4E7654B31579B1400D259FE9B82818D1" metricMagnitude="-3" metricUnit="m" metricValue="8.7" unit="mm" value="8.7">8.7 mm</quantity>
|
||
) and
|
||
<taxonomicName id="B753C7A3DB50D4D4C4C3D4E0EF3EED92" authorityName="Liu, Hutchings & Sun" authorityYear="2017" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="tripectinata">
|
||
<emphasis id="0287D4F425BE3375B40BFFF969BE900A" italics="true">M tripectinata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<abbrev id="ABBRID0E1NCI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
:
|
||
<quantity id="9710408591E9B3D7C3F3305B36CBB272" metricMagnitude="-2" metricUnit="m" metricValue="1.27" unit="mm" value="12.7">12.7 mm</quantity>
|
||
) are three and eight respectively, differs from
|
||
<taxonomicName id="D46F87A32EC48D06EF7668AD026C6605" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="0DC4069D357608FF39EBB128D08EAEEF" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0B9271551B523F9441F5ECEAB39972D6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<abbrev id="ABBRID0EJOCI" xlink_title="Length at chaetiger 10">L 10</abbrev>
|
||
: 5.25 (3.45–5.85) mm), which has a maximum of six.
|
||
</paragraph>
|
||
<paragraph id="8C7BAB03472645F476B052F136995C0E">
|
||
Finally,
|
||
<taxonomicName id="6004477E52327F159A6DC0FC15E6A493" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="F95F34B49EF0DC931E6EE51946DF0B0D" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9DA5CCEF6C4EB439908BBD800C675FB5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is similar to
|
||
<taxonomicName id="C052293FD5A895FD179A6FF614EBA204" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="D8314E1A3F767AD9DCAC3EF006CEDB02" italics="true">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
by having unidentate subacicular hooks.
|
||
<taxonomicName id="F188E139EE155119797D57F14A0C2518" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="7E574B8BCF25FC7BF6053A0F48DDA46C" italics="true">Marphysa merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71089DE10E6EB241CBF6D7B025E9D47F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a pair of eyes and two pairs of anal cirri, while
|
||
<taxonomicName id="C4AF6079831A434BC4C828B1D2E9859B" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="C76F89EB90021D068179CC2DA886860F" italics="true">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
has no eyes and only one pair of anal cirri. Also, branchiae in
|
||
<taxonomicName id="223FA5FB7DB8C98BDF7AF713FAD83300" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="1A4430FCC4D28638442A7CF6EECB3AFA" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85781434FA0E4F9A1EE07AA8FDDD4886" rank="species">sp. nov.</taxonomicNameLabel>
|
||
begin earlier from chaetiger 24 (16–27) compared to
|
||
<taxonomicName id="09A6A18376A9FA4E8C16CA946081D7FA" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="89172F27472396E77EC30727DBC816A1" italics="true">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
(chaetiger 45). The maximum number of branchial filaments in
|
||
<taxonomicName id="0E77EF63BC3CC76072A25E8BB6CF4A41" authority="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings, 2024" authorityName="Che Engku Abdullah & Idris & Fahmi & Flaxman & Hutchings" authorityYear="2024" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="merchangensis" status="sp. nov.">
|
||
<emphasis id="7E368ADDFDA9ABFC5E40CD432E21C9F3" italics="true">M. merchangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FA2E055E9932DF355EBAB1705A8C996C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is six, while
|
||
<taxonomicName id="82C108086396B990F2A6CF00C8249F83" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="5A30AB8C7D68139585A5FD7AB7D81902" italics="true">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
has a maximum of three branchial filaments. Nevertheless, the original description of
|
||
<taxonomicName id="B796A2589DE3C8E9DDAE34EDEBCCDA2E" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="E7B1282308E9512D4FC010F8F89BAE6B" italics="true">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
is incomplete and does not include certain important features such as the number and type of pectinate chaetae. Fresh material of
|
||
<taxonomicName id="5386F62CD3F909B9081DEB9CB2FEC30A" authorityName=", Treadwell" authorityYear="1936" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" phylum="Annelida" rank="species" species="orientalis">
|
||
<emphasis id="0B24985E4E32FFDE225BA6E368FC215E" italics="true">M. orientalis</emphasis>
|
||
</taxonomicName>
|
||
should be collected and redescribed from the type locality at Gulf of Mannar,
|
||
<collectingCountry id="861E33D000CF2D7D5666F7B222FB6065" name="Sri Lanka">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |