treatments-xml/data/37/5B/87/375B87BEFFD8A61C278ECF92FB31FB77.xml
2024-06-21 12:33:32 +02:00

223 lines
28 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="7EB69C2B636501CE2DBC60B98CB341E9" ID-DOI="10.1080/00222933.2018.1437231" ID-ISSN="1464-5262" ID-Zenodo-Dep="5178489" ID-ZooBank="9FC2A016-1295-4047-877E-BC9030BD2BD2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628626551251" checkinUser="felipe" docAuthor="Santarém, Maria Clara Alves, Borkent, Art, Spinelli, Gustavo &amp; Felippe-Bauer, Maria Luiza" docDate="2018" docId="375B87BEFFD8A61C278ECF92FB31FB77" docLanguage="en" docName="JNatHist.52.9-10.509-540.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (9 - 10)" docSource="http://dx.doi.org/10.1080/00222933.2018.1437231" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Downeshelea marambaia Santarém &amp; Borkent &amp; Spinelli &amp; Felippe-Bauer 2018, sp. nov." docType="treatment" docUuid="B4E7D002-4F93-4922-B5D6-576F0444E678" docUuidSource="ZooBank" docVersion="5" lastPageNumber="536" masterDocId="CB62FFC6FFC3A600264FCC1AFFB5FFA5" masterDocTitle="New Neotropical species of Downeshelea Wirth and Grogan and redescription of D. multilineata (Lutz) (Diptera: Ceratopogonidae)" masterLastPageNumber="540" masterPageNumber="509" pageNumber="535" updateTime="1699048774547" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="8B667F169E2B5C3618B6250E0573CAA7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9D4C5986D64427B7447BFBC6729A5507">
<mods:title id="A17C0356000E2F9F5EAB9F61870FFDCA">New Neotropical species of Downeshelea Wirth and Grogan and redescription of D. multilineata (Lutz) (Diptera: Ceratopogonidae)</mods:title>
</mods:titleInfo>
<mods:name id="26E1FE9A1AC83D83578AA4BA5C03AE52" type="personal">
<mods:role id="CCC9BF584F66A635E2B00AF4ED4A835E">
<mods:roleTerm id="CE109132307A6C36201A047A36546060">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="814A49CE4FC3B90704D411EF942BB832">Santarém, Maria Clara Alves</mods:namePart>
</mods:name>
<mods:name id="6DDF4D0097404F58159A023F9DC43814" type="personal">
<mods:role id="050345F2F4E440082922ED219982EE26">
<mods:roleTerm id="24E75ECDF1369C4F32F3C7BA13CEB8CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BA2AEAABA9517AF8AF8C538E00548C2">Borkent, Art</mods:namePart>
</mods:name>
<mods:name id="C6AB0BCA07D48C72C1354F95FA3AC3A0" type="personal">
<mods:role id="3AD36E50F6681DCDD04F06D25580214B">
<mods:roleTerm id="982B960DB17042F16CC0DF5E9FB8B283">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC45B41640385EB041FB2A7E691FDB5C">Spinelli, Gustavo</mods:namePart>
</mods:name>
<mods:name id="8CA36F295410A76F29D30DD8EB10F096" type="personal">
<mods:role id="C269B81D4E8C0B8ACDDAB195DEC25302">
<mods:roleTerm id="778289F002916E6FADA036A371E5585B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64E3DF1FE3C1131437FB59056C714D60">Felippe-Bauer, Maria Luiza</mods:namePart>
</mods:name>
<mods:typeOfResource id="E894DC587122487C80C57C5A080EE0FB">text</mods:typeOfResource>
<mods:relatedItem id="F9AB6EC9A91775801FB6C2EFCEDB94D3" type="host">
<mods:titleInfo id="A76262803E117114F043BA061482638E">
<mods:title id="2FEB43B31481ACA6BDB7183741E9A99C">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="02C233DE4BF2166361E1050038D68165">
<mods:date id="F1843A03B65BC17B800662F68A6C300F">2018</mods:date>
<mods:detail id="699F7B902A1052B1D10DA02D1853F45D" type="series">
<mods:title id="D05539A78BE1E7C7E8F6A2C5FDDC8542">J. Nat. Hist.</mods:title>
</mods:detail>
<mods:detail id="01F264CEC83FB0A2792781A84B9BD22F" type="pubDate">
<mods:number id="842A1743338DB9265612DBFB35222343">2018-03-07</mods:number>
</mods:detail>
<mods:detail id="32F34C82F2F27332F246CBC6CDE88BCB" type="volume">
<mods:number id="0E68BAB4203A684193F7A2C65787BDD4">52</mods:number>
</mods:detail>
<mods:detail id="5777C90954B51A7A4571DA8770DBB1AF" type="issue">
<mods:number id="58693C462821855B6428E3282AA940A6">9 - 10</mods:number>
</mods:detail>
<mods:extent id="0D8B590AF9F3DE32BD44C66723DEC64B" unit="page">
<mods:start id="6189E97D0CAE2D1359D4E6DE5489181C">509</mods:start>
<mods:end id="D839CA82940C170178197D8F1C1DA763">540</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="51EE6294D820BA76B28CDA237CAC6D8F">
<mods:url id="088DAAE9F8C9CF880091D4742A135774">http://dx.doi.org/10.1080/00222933.2018.1437231</mods:url>
</mods:location>
<mods:classification id="0700418F2E06554B0FF3505D3956401A">journal article</mods:classification>
<mods:identifier id="22C0517F9B1CF63671A42ECFC9A83023" type="DOI">10.1080/00222933.2018.1437231</mods:identifier>
<mods:identifier id="EE91FBFFEB88F5A9375E89ADF5CD82E4" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="34A554B9FA38E8DDEB17A1FFB9E4C6AE" type="Zenodo-Dep">5178489</mods:identifier>
<mods:identifier id="45C5F061B489B5D13C6B7A613EBBA9F7" type="ZooBank">9FC2A016-1295-4047-877E-BC9030BD2BD2</mods:identifier>
</mods:mods>
<treatment id="375B87BEFFD8A61C278ECF92FB31FB77" ID-DOI="http://doi.org/10.5281/zenodo.5189618" ID-Zenodo-Dep="5189618" LSID="urn:lsid:zoobank.org:act:B4E7D002-4F93-4922-B5D6-576F0444E678" httpUri="http://treatment.plazi.org/id/375B87BEFFD8A61C278ECF92FB31FB77" lastPageId="28" lastPageNumber="536" pageId="27" pageNumber="535">
<subSubSection id="F7E86523FFD8A61B278ECF92FCD7FC04" box="[449,866,904,929]" pageId="27" pageNumber="535" type="nomenclature">
<paragraph id="BF4D36A8FFD8A61B278ECF92FCD7FC04" blockId="27.[449,866,904,964]" box="[449,866,904,929]" pageId="27" pageNumber="535">
<heading id="E40581C4FFD8A61B278ECF92FCD7FC04" box="[449,866,904,929]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="535" reason="5">
<taxonomicName id="78F24D2BFFD8A61B278ECF92FD49FC04" authority="Santarém &amp; Borkent &amp; Spinelli &amp; Felippe-Bauer, 2018" authorityName="Santarém &amp; Borkent &amp; Spinelli &amp; Felippe-Bauer" authorityYear="2018" box="[449,764,904,929]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="535" phylum="Arthropoda" rank="species" species="marambaia" status="sp. nov.">
<emphasis id="8D86EABAFFD8A61B278ECF92FD49FC04" bold="true" box="[449,764,904,929]" italics="true" pageId="27" pageNumber="535">Downeshelea marambaia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96B557C1FFD8A61B254ACF92FCD7FC04" box="[773,866,904,929]" pageId="27" pageNumber="535" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F7E86523FFD8A61B244CCFB1FC95FC61" box="[515,800,939,964]" pageId="27" pageNumber="535" type="description">
<paragraph id="BF4D36A8FFD8A61B244CCFB1FC95FC61" blockId="27.[449,866,904,964]" box="[515,800,939,964]" pageId="27" pageNumber="535">
(
<figureCitation id="27C92A2DFFD8A61B2444CFB1FD11FC61" box="[523,676,939,964]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (ad). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (eh). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="27" pageNumber="535">Figures 7ad</figureCitation>
,
<figureCitation id="27C92A2DFFD8A61B24FDCFB1FD58FC61" box="[690,749,939,964]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="27" pageNumber="535">9eg</figureCitation>
,
<figureCitation id="27C92A2DFFD8A61B24B3CFB1FCACFC61" box="[764,793,939,964]" captionStart="Figure 10" captionStartId="25.[160,225,1002,1025]" captionTargetBox="[162,1153,142,971]" captionTargetId="figure-177@25.[162,1153,142,971]" captionTargetPageId="25" captionText="Figure 10. Geographic distribution of species in the Downeshelea multilineata species group." figureDoi="http://doi.org/10.5281/zenodo.5178509" httpUri="https://zenodo.org/record/5178509/files/figure.png" pageId="27" pageNumber="535">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7E86523FFD8A61B26EFCFE3FECAFAA8" pageId="27" pageNumber="535" type="diagnosis">
<paragraph id="BF4D36A8FFD8A61B26EFCFE3FEA1FBB7" blockId="27.[160,1156,1017,1215]" box="[160,276,1017,1042]" pageId="27" pageNumber="535">
<emphasis id="8D86EABAFFD8A61B26EFCFE3FEA1FBB7" bold="true" box="[160,276,1017,1042]" italics="true" pageId="27" pageNumber="535">Diagnosis</emphasis>
</paragraph>
<paragraph id="BF4D36A8FFD8A61B26EFC806FB86FB1A" blockId="27.[160,1156,1017,1215]" pageId="27" pageNumber="535">
<emphasis id="8D86EABAFFD8A61B26EFC806FF55FB90" bold="true" box="[160,224,1052,1077]" italics="true" pageId="27" pageNumber="535">Male.</emphasis>
The only species of
<taxonomicName id="78F24D2BFFD8A61B279FC806FDD4FB90" box="[464,609,1052,1077]" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="535" phylum="Arthropoda" rank="genus">
<emphasis id="8D86EABAFFD8A61B279FC806FDD4FB90" box="[464,609,1052,1077]" italics="true" pageId="27" pageNumber="535">Downeshelea</emphasis>
</taxonomicName>
in the Americas with the following combination of features: gonostylus 0.650.66 length of gonocoxite; parameres fused for 0.12 of total length; stem slightly curved, convergent distally, slightly enlarged apically, distal portion strongly curved (
<figureCitation id="27C92A2DFFD8A61B2710C89FFE7FFB38" box="[351,458,1157,1181]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="27" pageNumber="535">
Figure
<quantity id="780A9B4DFFD8A61B27E1C89FFE7FFB38" box="[430,458,1157,1181]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="27" pageNumber="535" unit="g" value="9.0">9g</quantity>
</figureCitation>
); aedeagus rectangular, basal arch shallow, extending to 0.17 of total length with two large elliptical sclerotised anteromesal areas (
<figureCitation id="27C92A2DFFD8A61B258EC8BDFB93FB1A" box="[961,1062,1191,1215]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="27" pageNumber="535">Figure 9f</figureCitation>
).
</paragraph>
<paragraph id="BF4D36A8FFD8A61B26EFC8EFFECAFAA8" blockId="27.[160,383,1269,1294]" box="[160,383,1269,1294]" pageId="27" pageNumber="535">
<emphasis id="8D86EABAFFD8A61B26EFC8EFFF49FAA8" bold="true" box="[160,252,1269,1294]" italics="true" pageId="27" pageNumber="535">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="F7E86523FFD8A61C26EFC959FD00FC63" lastPageId="28" lastPageNumber="536" pageId="27" pageNumber="535" type="description">
<paragraph id="BF4D36A8FFD8A61B26EFC959FE93FAF9" blockId="27.[160,1155,1347,1441]" box="[160,294,1347,1372]" pageId="27" pageNumber="535">
<emphasis id="8D86EABAFFD8A61B26EFC959FE93FAF9" bold="true" box="[160,294,1347,1372]" italics="true" pageId="27" pageNumber="535">Description</emphasis>
</paragraph>
<paragraph id="BF4D36A8FFD8A61B26EFC97CFC46FA04" blockId="27.[160,1155,1347,1441]" pageId="27" pageNumber="535">
<emphasis id="8D86EABAFFD8A61B26EFC97CFF55FADB" bold="true" box="[160,224,1382,1407]" italics="true" pageId="27" pageNumber="535">Male.</emphasis>
<emphasis id="8D86EABAFFD8A61B26BFC97FFE9CFADB" box="[240,297,1381,1406]" italics="true" pageId="27" pageNumber="535">Head</emphasis>
. Eyes separated by a distance shorter than one ommatidia (
<figureCitation id="27C92A2DFFD8A61B2247C97CFBC3FADB" box="[1032,1142,1382,1406]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (ad). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (eh). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="27" pageNumber="535">Figure 7c</figureCitation>
); antennal ratio 1.00 (n = 1) (
<figureCitation id="27C92A2DFFD8A61B2793C993FDFCFA04" box="[476,585,1417,1441]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (ad). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (eh). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="27" pageNumber="535">Figure 7b</figureCitation>
); palpal ratio 2.50 (n = 1) (
<figureCitation id="27C92A2DFFD8A61B2532C993FC53FA04" box="[893,998,1417,1441]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (ad). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (eh). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="27" pageNumber="535">Figure 7c</figureCitation>
).
</paragraph>
<paragraph id="BF4D36A8FFD8A61C26EFC9CCFDB0FF6C" blockId="27.[160,1156,1494,1727]" lastBlockId="28.[160,1155,142,201]" lastPageId="28" lastPageNumber="536" pageId="27" pageNumber="535">
<emphasis id="8D86EABAFFD8A61B26EFC9CCFF4CFA4A" bold="true" box="[160,249,1494,1519]" italics="true" pageId="27" pageNumber="535">Thorax.</emphasis>
Brown, without definite pattern in slide-mounted specimens. Wing (
<figureCitation id="27C92A2DFFD8A61B225DC9CDFBC9FA4A" box="[1042,1148,1495,1519]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (ad). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (eh). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="27" pageNumber="535">Figure 7a</figureCitation>
) with greyish spot over CuA
<subScript id="237634EDFFD8A61B279ACA1EFE6AF9B1" attach="left" box="[469,479,1540,1556]" fontSize="7" pageId="27" pageNumber="535">2</subScript>
reaching wing margin; 2
<superScript id="48879BE0FFD8A61B24B2C9EEFCA6F9A1" attach="left" box="[765,787,1524,1540]" fontSize="7" pageId="27" pageNumber="535">nd</superScript>
radial cell twice longer than 1
<superScript id="48879BE0FFD8A61B2220C9EEFBCBF9A1" attach="none" box="[1135,1150,1524,1540]" fontSize="7" pageId="27" pageNumber="535">st</superScript>
; wing length 1.05 (n = 1) mm; breadth 0.37 (n = 1) mm; costal ratio 0.74 (n = 1). Halter knob brown, stem pale. Legs (
<figureCitation id="27C92A2DFFD8A61B244FCA25FDD8F9F2" box="[512,621,1599,1623]" captionStart="Figure 7" captionStartId="22.[160,225,1582,1605]" captionTargetBox="[163,1150,144,1550]" captionTargetId="figure-1@22.[163,1152,142,1551]" captionTargetPageId="22" captionText="Figure 7. Downeshelea marambaia sp. nov., male (ad). (a) wing; (b) antenna; (c) head, anterior view; (d) fore, mid, hind legs (left to right). Downeshelea moravia sp. nov., male (eh). (e) wing; (f) head, anterior view; (g) palpus; (h) fore, mid, hind legs (left to right)." figureDoi="http://doi.org/10.5281/zenodo.5178503" httpUri="https://zenodo.org/record/5178503/files/figure.png" pageId="27" pageNumber="535">Figure 7d</figureCitation>
) brown, hind femur darker apically, hind tibia with subbasal dark band. Fore-, hind tarsomere 1 with one basal, one apical spine; midtarsomere 1 with two basal, two apical, three other ventral spines; apical spines of tarsomeres 24 of fore-, mid-, hind legs: 223, 222, 111; fore tarsal ratio 2.25 (n = 1), mid tarsal ratio 2.52 (n = 1), hind tarsal ratio 2.24 (n = 1); claws 0.40 (n = 1) length of their respective tarsomere 5.
</paragraph>
<paragraph id="BF4D36A8FFDFA61C26EFCD18FBC1FD3D" blockId="28.[160,1156,258,664]" pageId="28" pageNumber="536">
<emphasis id="8D86EABAFFDFA61C26EFCD18FEADFEBE" bold="true" box="[160,280,258,283]" italics="true" pageId="28" pageNumber="536">Abdomen.</emphasis>
Dark brown. Terminalia (
<figureCitation id="27C92A2DFFDFA61C240ACD19FD04FEBE" box="[581,689,259,283]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="28" pageNumber="536">Figure 9e</figureCitation>
): tergite 9 with quadrate apex, apicolateral process elongate, slender, in bad condition in slide-mounted specimen (not illustrated); sternite 9 concave anteriorly, posterior margin in bad condition in slidemounted specimen (not illustrated). Gonocoxite 1.912.17 (2.04, n = 2) times longer than basal width; gonostylus delicate, 0.650.66 (0.65, n = 2) length of gonocoxite. Parameres (
<figureCitation id="27C92A2DFFDFA61C2767CDAAFE2DFE6D" box="[296,408,432,456]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="28" pageNumber="536">
Figure
<quantity id="780A9B4DFFDFA61C2734CDAAFE2DFE6D" box="[379,408,432,456]" metricMagnitude="-3" metricUnit="kg" metricValue="9.0" pageId="28" pageNumber="536" unit="g" value="9.0">9g</quantity>
</figureCitation>
) 1.161.17 (1.16, n = 2) times longer than aedeagus, fused for 0.12 (n = 1) of total length; knob bulbous; stem nearly straight, convergent distally, slightly expanded apically, distal portion strongly curved, tapering to tip, 0.56 (n = 1) of total length. Aedeagus (
<figureCitation id="27C92A2DFFDFA61C278BCE02FD9AFD95" box="[452,559,536,560]" captionStart="Figure 9" captionStartId="24.[160,225,1645,1668]" captionTargetBox="[198,1117,142,1613]" captionTargetId="figure-1@24.[198,1117,142,1614]" captionTargetPageId="24" captionText="Figure 9. Downeshelea jarina sp. nov., (a) male terminalia with aedeagus; (b) parameres. Downeshelea litorale sp. nov., (c) male terminalia with aedeagus; (d) parameres. Downeshelea marambaia sp. nov., (e) male terminalia; (f) aedeagus; (g) parameres. Downeshelea moravia sp. nov., (h) male terminalia with aedeagus; (i) parameres." figureDoi="http://doi.org/10.5281/zenodo.5178507" httpUri="https://zenodo.org/record/5178507/files/figure.png" pageId="28" pageNumber="536">Figure 9f</figureCitation>
) rectangular, enlarged distally, heavily sclerotised laterally, basal arch shallow, extending to 0.17 (n = 1) of total length with large elliptical sclerotised anteromesal areas reaching midlength of aedeagus; distal portion with deep mesal excavation terminating in two prominent sclerotised pointed serrate processes.
</paragraph>
<paragraph id="BF4D36A8FFDFA61C26EFCECBFECAFD4F" blockId="28.[160,383,721,746]" box="[160,383,721,746]" pageId="28" pageNumber="536">
<emphasis id="8D86EABAFFDFA61C26EFCECBFF49FD4F" bold="true" box="[160,252,721,746]" italics="true" pageId="28" pageNumber="536">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="BF4D36A8FFDFA61C26EFCF39FE2FFC99" blockId="28.[160,1156,803,966]" box="[160,410,803,828]" pageId="28" pageNumber="536">
<emphasis id="8D86EABAFFDFA61C26EFCF39FE2FFC99" bold="true" box="[160,410,803,828]" italics="true" pageId="28" pageNumber="536">Specimens examined</emphasis>
</paragraph>
<paragraph id="BF4D36A8FFDFA61C26EFCF5CFD00FC63" blockId="28.[160,1156,803,966]" pageId="28" pageNumber="536">
<typeStatus id="6049880AFFDFA61C26EFCF5CFEB2FCFB" box="[160,263,838,862]" pageId="28" pageNumber="536" type="holotype">Holotype</typeStatus>
male adult, labeled
<materialsCitation id="0F9A3CF5FFDFA61C2445CF5CFEDAFC01" collectingDate="1990-03-22" collectionCode="CCER" country="Brazil" location="Itaguai" pageId="28" pageNumber="536" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<typeStatus id="6049880AFFDFA61C2445CF5CFDC4FCFB" box="[522,625,838,862]" pageId="28" pageNumber="536" type="holotype">Holotype</typeStatus>
<taxonomicName id="78F24D2BFFDFA61C2431CF5CFE62FC24" authority="Santarem, Borkent, Spinelli and Felippe-Bauer" authorityName="Santarem, Borkent, Spinelli and Felippe-Bauer" class="Insecta" family="Ceratopogonidae" genus="Downeshelea" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="536" phylum="Arthropoda" rank="species" species="marambaia">
<emphasis id="8D86EABAFFDFA61C2431CF5CFC2EFCFA" box="[638,923,838,863]" italics="true" pageId="28" pageNumber="536">Downeshelea marambaia</emphasis>
Santarém, Borkent, Spinelli and Felippe-Bauer
</taxonomicName>
,
<collectingCountry id="C7E57638FFDFA61C27A7CF73FD8FFC24" box="[488,570,873,897]" name="Brazil" pageId="28" pageNumber="536">BRAZIL</collectingCountry>
,
<collectingRegion id="7D36F84AFFDFA61C2404CF73FD4EFC24" box="[587,763,873,897]" country="Brazil" name="Rio de Janeiro" pageId="28" pageNumber="536">Rio de Janeiro</collectingRegion>
,
<location id="BA2D6073FFDFA61C2543CF73FCEEFC24" LSID="urn:lsid:plazi:treatment:375B87BEFFD8A61C278ECF92FB31FB77:BA2D6073FFDFA61C2543CF73FCEEFC24" box="[780,859,873,897]" country="Brazil" name="Itaguai" pageId="28" pageNumber="536" stateProvince="Rio de Janeiro">Itaguaí</location>
,
<date id="CB4C1068FFDFA61C2522CF73FB91FC24" box="[877,1060,873,897]" pageId="28" pageNumber="536" value="1990-03-22">
<collectingDate id="DB08E980FFDFA61C2522CF73FB91FC24" box="[877,1060,873,897]" pageId="28" pageNumber="536" value="1990-03-22">22 March 1990</collectingDate>
</date>
, FEEMA team col. (
<collectionCode id="D9E3AE6DFFDFA61C2767CF96FEDFFC01" box="[296,362,908,932]" pageId="28" pageNumber="536">CCER</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0F9A3CF5FFDFA61C27CECF96FBCAFC01" box="[385,1151,908,932]" country="Brazil" location="Marambaia island" pageId="28" pageNumber="536" specimenCount="2" stateProvince="Rio de Janeiro" typeStatus="paratype">
<typeStatus id="6049880AFFDFA61C27CECF96FE51FC01" box="[385,484,908,932]" pageId="28" pageNumber="536" type="paratype">Paratype</typeStatus>
male labeled as
<typeStatus id="6049880AFFDFA61C248FCF96FC91FC01" box="[704,804,908,932]" pageId="28" pageNumber="536" type="holotype">holotype</typeStatus>
except
<location id="BA2D6073FFDFA61C25C2CF96FBE8FC01" LSID="urn:lsid:plazi:treatment:375B87BEFFD8A61C278ECF92FB31FB77:BA2D6073FFDFA61C25C2CF96FBE8FC01" box="[909,1117,908,932]" country="Brazil" name="Marambaia island" pageId="28" pageNumber="536" stateProvince="Rio de Janeiro">Marambaia island</location>
, P
</materialsCitation>
. Armas,
<date id="CB4C1068FFDFA61C26BACFB4FE03FC63" box="[245,438,942,966]" pageId="28" pageNumber="536" value="1992-10-29">29 October 1992</date>
, Quintelas col. (CCER).
</paragraph>
</subSubSection>
<subSubSection id="F7E86523FFDFA61C26EFCFE5FE07FBFB" pageId="28" pageNumber="536" type="distribution">
<paragraph id="BF4D36A8FFDFA61C26EFCFE5FE50FBBD" blockId="28.[160,1155,1023,1118]" box="[160,485,1023,1048]" pageId="28" pageNumber="536">
<emphasis id="8D86EABAFFDFA61C26EFCFE5FE50FBBD" bold="true" box="[160,485,1023,1048]" italics="true" pageId="28" pageNumber="536">Distribution and bionomics</emphasis>
</paragraph>
<paragraph id="BF4D36A8FFDFA61C26EFC839FE07FBFB" blockId="28.[160,1155,1023,1118]" pageId="28" pageNumber="536">
This species is restricted to
<collectingRegion id="7D36F84AFFDFA61C27B4C839FD1CFB9E" box="[507,681,1059,1083]" country="Brazil" name="Rio de Janeiro" pageId="28" pageNumber="536">Rio de Janeiro</collectingRegion>
(
<collectingCountry id="C7E57638FFDFA61C24F1C839FD48FB9E" box="[702,765,1059,1083]" name="Brazil" pageId="28" pageNumber="536">Brazil</collectingCountry>
) in coastal areas of the Itaguai municipality (
<figureCitation id="27C92A2DFFDFA61C2775C85CFE10FBFB" box="[314,421,1094,1118]" captionStart="Figure 10" captionStartId="25.[160,225,1002,1025]" captionTargetBox="[162,1153,142,971]" captionTargetId="figure-177@25.[162,1153,142,971]" captionTargetPageId="25" captionText="Figure 10. Geographic distribution of species in the Downeshelea multilineata species group." figureDoi="http://doi.org/10.5281/zenodo.5178509" httpUri="https://zenodo.org/record/5178509/files/figure.png" pageId="28" pageNumber="536">Figure 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7E86523FFDFA61C26EFC88DFB31FB77" box="[160,1156,1175,1234]" pageId="28" pageNumber="536" type="etymology">
<paragraph id="BF4D36A8FFDFA61C26EFC88DFEA8FB15" blockId="28.[160,1156,1175,1234]" box="[160,285,1175,1200]" pageId="28" pageNumber="536">
<emphasis id="8D86EABAFFDFA61C26EFC88DFEA8FB15" bold="true" box="[160,285,1175,1200]" italics="true" pageId="28" pageNumber="536">Etymology</emphasis>
</paragraph>
<paragraph id="BF4D36A8FFDFA61C26EFC8A0FB31FB77" blockId="28.[160,1156,1175,1234]" box="[160,1156,1210,1234]" pageId="28" pageNumber="536">
The name of this species refers to its distribution in Marambaia island,
<collectingRegion id="7D36F84AFFDFA61C25D1C8A0FB8FFB77" box="[926,1082,1210,1234]" country="Brazil" name="Rio de Janeiro" pageId="28" pageNumber="536">Rio de Janeiro</collectingRegion>
,
<collectingCountry id="C7E57638FFDFA61C220CC8A0FB35FB77" box="[1091,1152,1210,1234]" name="Brazil" pageId="28" pageNumber="536">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>