190 lines
23 KiB
XML
190 lines
23 KiB
XML
<document id="27AE770F75CF460C6654895331A4454D" ID-CLB-Dataset="31005" ID-DOI="10.11646/phytotaxa.599.3.4" ID-GBIF-Dataset="df7cb3e2-17f0-4060-a457-687ddd68b01d" ID-ISSN="1179-3163" ID-Zenodo-Dep="8012553" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1686118506052" checkinUser="plazi" docAuthor="Hoo, Pui Kiat, Sugumaran, Manickam & Wong, Khoon Meng" docDate="2023" docId="03E3D725C97A860262C8F9A6F8BEF981" docLanguage="en" docName="phytotaxa.599.3.4.pdf" docOrigin="Phytotaxa 599 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.599.3.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Timonius Candolle 1830" docType="treatment" docVersion="7" lastPageNumber="175" masterDocId="FFDAAF5DC97986066240FF96FC63FFEC" masterDocTitle="Two new species of Timonius (Rubiaceae: Guettardeae) from the Malay Peninsula" masterLastPageNumber="182" masterPageNumber="171" pageNumber="174" updateTime="1699521378453" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="C85E8A3A8704F73542C0126A899841D6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="76D038E2C963C4C164143FA27345B4F5">
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<mods:title id="FA134E95BE3CCF72BAA559EDC7B531DB">Two new species of Timonius (Rubiaceae: Guettardeae) from the Malay Peninsula</mods:title>
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<mods:name id="F8A47FD623369C7633A1E4CDBB6B2AB3" type="personal">
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<mods:roleTerm id="7298B43581BAC9378A8C51645CD27A1E">Author</mods:roleTerm>
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<mods:namePart id="A0711C35331E111CB7DB70CB8ED7A751">Hoo, Pui Kiat</mods:namePart>
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<mods:affiliation id="EF801689AB8DEC23EEBE42258989D524">Institute of Biological Sciences, University of Malaya, 50603 Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:name id="B5F70A91BDBB71C4A2240CC0EC193E9E" type="personal">
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<mods:roleTerm id="373536F82258476B37213A28E75393F1">Author</mods:roleTerm>
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<mods:namePart id="271D5E18C4D70BE4197D5F1C620830DF">Sugumaran, Manickam</mods:namePart>
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<mods:affiliation id="4EAA8A58F7E120BA07621F8E5791E9D4">Institute of Biological Sciences, University of Malaya, 50603 Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:roleTerm id="E197F9A8285FDBDCF86B823399DF39E8">Author</mods:roleTerm>
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<mods:namePart id="D6411E03609F33CA5E2498612EB3F523">Wong, Khoon Meng</mods:namePart>
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<mods:affiliation id="2EF2A0E925D1611F7510A51D044387CE">Singapore Botanic Gardens, 1 Cluny Road, Singapore 259569, Republic of Singapore</mods:affiliation>
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<mods:title id="2424C7D20F0155031FBC38E6857953E0">Phytotaxa</mods:title>
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<mods:part id="F558019AB02BCDB54746271CCEAF1FB2">
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<mods:date id="694BB6E8FA3C6F9E3CE6CADEF7498A6E">2023</mods:date>
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<mods:number id="1543176D8C1804C414CAA0C27FED9D2F">2023-06-07</mods:number>
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<mods:number id="7AC3844468C0F5C3A015AA3B7EB669D0">599</mods:number>
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<mods:start id="5613B32A6E04742F557A68BD93954ADD">171</mods:start>
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<mods:url id="C2DFC856B68CD871BBC9665A89517D3A">http://dx.doi.org/10.11646/phytotaxa.599.3.4</mods:url>
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<treatment id="03E3D725C97A860262C8F9A6F8BEF981" ID-DOI="http://doi.org/10.5281/zenodo.8012555" ID-GBIF-Taxon="209670392" ID-Zenodo-Dep="8012555" LSID="urn:lsid:plazi:treatment:03E3D725C97A860262C8F9A6F8BEF981" httpUri="http://treatment.plazi.org/id/03E3D725C97A860262C8F9A6F8BEF981" lastPageId="4" lastPageNumber="175" pageId="3" pageNumber="174">
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<subSubSection id="C35035B8C97A860562C8F9A6FFEBF9A6" box="[136,904,1584,1611]" pageId="3" pageNumber="174" type="nomenclature">
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<paragraph id="8BF56633C97A860562C8F9A6FFEBF9A6" blockId="3.[136,904,1584,1611]" box="[136,904,1584,1611]" pageId="3" pageNumber="174">
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<heading id="D0BDD15FC97A860562C8F9A6FFEBF9A6" bold="true" box="[136,904,1584,1611]" fontSize="11" level="1" pageId="3" pageNumber="174" reason="1">
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<emphasis id="B93EBA21C97A860562C8F9A6FFEBF9A6" bold="true" box="[136,904,1584,1611]" pageId="3" pageNumber="174">
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An unusual
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<taxonomicName id="4C4A1DB0C97A86056359F9A7FDE5F9A7" ID-CoL="8W47W" ID-ENA="88289" authorityName="Candolle" authorityYear="1830" box="[281,390,1585,1611]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="174" phylum="Tracheophyta" rank="genus">
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<emphasis id="B93EBA21C97A86056359F9A7FDE5F9A7" bold="true" box="[281,390,1585,1611]" italics="true" pageId="3" pageNumber="174">Timonius</emphasis>
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with black bark from the Malay Peninsula
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35035B8C97A860562C8F9EFFF76F7F3" pageId="3" pageNumber="174" type="reference_group">
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<paragraph id="8BF56633C97A860562C8F9EFFFB3F863" blockId="3.[136,1452,1657,2079]" pageId="3" pageNumber="174">
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<treatmentCitationGroup id="AB5A411DC97A860562C8F9EFFF7BF97F" box="[136,792,1657,1683]" pageId="3" pageNumber="174">
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<treatmentCitation id="0AEB4022C97A860562C8F9EFFF7BF97F" author="Baker, J. G." box="[136,792,1657,1683]" page="127" pageId="3" pageNumber="174" year="1823">
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<taxonomicName id="4C4A1DB0C97A860562C8F9EFFF7BF97F" ID-CoL="56ZHD" authority="(Jack 1823: 127) Baker (1877: 144)" authorityName="Baker" authorityPageNumber="144" authorityYear="1877" baseAuthorityName="Jack" baseAuthorityPageNumber="127" baseAuthorityYear="1823" box="[136,792,1657,1683]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="174" phylum="Tracheophyta" rank="species" species="flavescens">
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<emphasis id="B93EBA21C97A860562C8F9EFFD0FF97F" box="[136,364,1657,1683]" italics="true" pageId="3" pageNumber="174">Timonius flavescens</emphasis>
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(
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<bibRefCitation id="EFDB1BC2C97A8605633EF9EFFE51F97F" author="Jack, W." box="[382,562,1657,1683]" pageId="3" pageNumber="174" pagination="114 - 130" refId="ref6336" refString="Jack, W. (1823) Account of the Lansium and some other genera of Malayan plants. Transactions of the Linnean Society 14: 114 - 130. https: // doi. org / 10.1111 / j. 1095 - 8339.1823. tb 00083. x" type="journal article" year="1823">Jack 1823: 127</bibRefCitation>
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)
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<bibRefCitation id="EFDB1BC2C97A86056004F9EFFF7BF97F" author="Baker, J. G." box="[580,792,1657,1683]" pageId="3" pageNumber="174" refId="ref5681" refString="Baker, J. G. (1877) Flora of Mauritius and the Seychelles: A description of the flowering plants and ferns of those islands. L. Reeve & Co, London, 557 pp. https: // doi. org / 10.5962 / bhl. title. 139960" type="book" year="1877">Baker (1877: 144)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
</treatmentCitationGroup>
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is commonly found in peat swamp forests mostly along coastal fringes including on coastal mountains where peaty substrates form. In the Malay Peninsula, it is one of the most widespread species, and although leaf shape and size can be variable, it is easily recognised in the field by its purplish midribs and secondary veins on the lower leaf surface, shagreened lower leaf lamina surface and 4-lobed fruits. While sorting through specimens in herbaria, a couple of collections (
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<emphasis id="B93EBA21C97A860561AAF89FF9C8F8CF" box="[1002,1451,1801,1827]" italics="true" pageId="3" pageNumber="174">Md. Shah, Ahmad & Mahmud MS 3317</emphasis>
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and
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<emphasis id="B93EBA21C97A860562F8F8BBFD06F8AB" box="[184,357,1837,1863]" italics="true" pageId="3" pageNumber="174">Loh FRI 13500</emphasis>
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) seemed to be a distinctive species but yet possible to be included in the general circumscription of
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<taxonomicName id="4C4A1DB0C97A860562E7F8C7FD55F887" ID-CoL="56ZHD" authorityName="Baker" authorityYear="1877" baseAuthorityName="Jack" baseAuthorityYear="1823" box="[167,310,1873,1899]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="174" phylum="Tracheophyta" rank="species" species="flavescens">
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<emphasis id="B93EBA21C97A860562E7F8C7FD55F887" box="[167,310,1873,1899]" italics="true" pageId="3" pageNumber="174">T. flavescens</emphasis>
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</taxonomicName>
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in
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<bibRefCitation id="EFDB1BC2C97A8605631AF8C7FD91F887" author="Wong, K. M." box="[346,498,1873,1899]" pageId="3" pageNumber="174" pagination="491 - 518" refId="ref7378" refString="Wong, K. M. (1988) The Antirheoideae (Rubiaceae) of the Malay Peninsula. Kew Bulletin 43: 491 - 518. https: // doi. org / 10.2307 / 4119769" type="journal article" year="1988">Wong (1988)</bibRefCitation>
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. These two collections are here deemed distinct from
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<taxonomicName id="4C4A1DB0C97A86056611F8C7F883F887" ID-CoL="56ZHD" authorityName="Baker" authorityYear="1877" baseAuthorityName="Jack" baseAuthorityYear="1823" box="[1105,1248,1873,1899]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="174" phylum="Tracheophyta" rank="species" species="flavescens">
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<emphasis id="B93EBA21C97A86056611F8C7F883F887" box="[1105,1248,1873,1899]" italics="true" pageId="3" pageNumber="174">T. flavescens</emphasis>
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</taxonomicName>
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based on ecology and general morphology including one unexpected character—bark colour.
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</paragraph>
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<paragraph id="8BF56633C97A860562FDF80FFF76F7F3" blockId="3.[136,1452,1657,2079]" pageId="3" pageNumber="174">
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All Malay Peninsula
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<taxonomicName id="4C4A1DB0C97A860563E7F80FFE6DF85E" ID-CoL="8W47W" authorityName="Candolle" authorityYear="1830" box="[423,526,1945,1970]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="174" phylum="Tracheophyta" rank="genus">
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<emphasis id="B93EBA21C97A860563E7F80FFE6DF85E" box="[423,526,1945,1970]" italics="true" pageId="3" pageNumber="174">Timonius</emphasis>
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</taxonomicName>
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hitherto known share a common bark colour which ranges from grey to brown (
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<figureCitation id="13717AB6C97A8605673EF80FFCCFF83B" captionStart="FIGURE 3" captionStartId="4.[136,229,1155,1177]" captionTargetBox="[151,1436,190,1112]" captionTargetId="figure-23@4.[151,1436,190,1130]" captionTargetPageId="4" captionText="FIGURE 3.A.Pale brown barkof Timonius flavescens, as in many other Malay Peninsula Timonius species.B. Black bark ofT.denigricans. Photos A by R.C.J. Lim and B by P.K. Hoo." figureDoi="http://doi.org/10.5281/zenodo.8012561" httpUri="https://zenodo.org/record/8012561/files/figure.png" pageId="3" pageNumber="174">Fig. 3A</figureCitation>
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) and so this character was not thought of as being important in species delimitation. However, the new species was noted as having black bark, a character that thus stands out among the species in the Malay Peninsula. We were able to ascertain its bark colour from field observations (
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<figureCitation id="13717AB6C97A860560F0F793FF6BF7F3" box="[688,776,2053,2079]" captionStart="FIGURE 3" captionStartId="4.[136,229,1155,1177]" captionTargetBox="[151,1436,190,1112]" captionTargetId="figure-23@4.[151,1436,190,1130]" captionTargetPageId="4" captionText="FIGURE 3.A.Pale brown barkof Timonius flavescens, as in many other Malay Peninsula Timonius species.B. Black bark ofT.denigricans. Photos A by R.C.J. Lim and B by P.K. Hoo." figureDoi="http://doi.org/10.5281/zenodo.8012561" httpUri="https://zenodo.org/record/8012561/files/figure.png" pageId="3" pageNumber="174">Fig. 3B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<caption id="DF3536BBC97D860262C8FB15FE4FFB50" ID-DOI="http://doi.org/10.5281/zenodo.8012561" ID-Zenodo-Dep="8012561" httpUri="https://zenodo.org/record/8012561/files/figure.png" pageId="4" pageNumber="175" startId="4.[136,229,1155,1177]" targetBox="[151,1436,190,1112]" targetPageId="4" targetType="figure">
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<paragraph id="8BF56633C97D860262C8FB15FE4FFB50" blockId="4.[136,1451,1155,1213]" pageId="4" pageNumber="175">
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<emphasis id="B93EBA21C97D860262C8FB15FC98FB74" bold="true" box="[136,251,1155,1177]" pageId="4" pageNumber="175">FIGURE 3.</emphasis>
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A. Pale brown bark of
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<taxonomicName id="4C4A1DB0C97D8602638EFB15FEEEFB74" authorityName="Baker" authorityYear="1877" baseAuthorityName="Jack" baseAuthorityYear="1823" box="[462,653,1155,1176]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="flavescens">
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<emphasis id="B93EBA21C97D8602638EFB15FEEEFB74" box="[462,653,1155,1176]" italics="true" pageId="4" pageNumber="175">Timonius flavescens</emphasis>
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</taxonomicName>
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, as in many other Malay Peninsula
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<taxonomicName id="4C4A1DB0C97D86026197FB15F8F7FB74" box="[983,1172,1155,1177]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="undefined-b">
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<emphasis id="B93EBA21C97D86026197FB15F84CFB74" box="[983,1071,1155,1176]" italics="true" pageId="4" pageNumber="175">Timonius</emphasis>
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species. B
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</taxonomicName>
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. Black bark of
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<taxonomicName id="4C4A1DB0C97D8602675EFB12F9C6FB74" box="[1310,1445,1155,1177]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="denigricans">
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<emphasis id="B93EBA21C97D8602675EFB12F9C6FB74" box="[1310,1445,1155,1177]" italics="true" pageId="4" pageNumber="175">T. denigricans</emphasis>
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</taxonomicName>
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. Photos A by R.C.J. Lim and B by P.K. Hoo.
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</paragraph>
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</caption>
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<subSubSection id="C35035B8C97D860262FDFB7DF8BEF981" pageId="4" pageNumber="175" type="description">
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<paragraph id="8BF56633C97D860262FDFB7DF8BEF981" blockId="4.[136,1452,1259,1646]" pageId="4" pageNumber="175">
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The new species, here named
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<taxonomicName id="4C4A1DB0C97D8602605AFB7AFF1BFAEA" authority="P. K. Hoo" authorityName="P. K. Hoo" box="[538,888,1259,1286]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="denigricans">
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<emphasis id="B93EBA21C97D8602605AFB7AFF6CFAE9" box="[538,783,1259,1285]" italics="true" pageId="4" pageNumber="175">Timonius denigricans</emphasis>
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P.K.Hoo
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</taxonomicName>
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due to the blackening of the bark, appears to be endemic to the Malay Peninsula. Its bark colour, although highly unusual for
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<taxonomicName id="4C4A1DB0C97D86026663FA86F8E9FAC5" authorityName="Candolle" authorityYear="1830" box="[1059,1162,1296,1321]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="genus">
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<emphasis id="B93EBA21C97D86026663FA86F8E9FAC5" box="[1059,1162,1296,1321]" italics="true" pageId="4" pageNumber="175">Timonius</emphasis>
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</taxonomicName>
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in the Malay Peninsula, apparently does occur in the genus, albeit rarely, in other regions. One such species is an endemic of the
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<collectingCountry id="F35D26A3C97D86026706FAA5FCB5FA9D" name="Solomon Islands" pageId="4" pageNumber="175">Solomon Islands</collectingCountry>
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—
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<taxonomicName id="4C4A1DB0C97D860262B3FACEFF5EFA9E" authority="Merrill & L. M. Perry (1945: 251)" authorityName="Merrill & L. M. Perry" authorityPageNumber="251" authorityYear="1945" box="[243,829,1367,1394]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="melanophloeus">
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<emphasis id="B93EBA21C97D860262B3FACEFDD9FA9D" box="[243,442,1367,1393]" italics="true" pageId="4" pageNumber="175">T. melanophloeus</emphasis>
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Merrill & L.M.Perry (1945: 251)
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</taxonomicName>
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, in which even the bark of the branchlets is black (the bark of the branchlets in
|
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<taxonomicName id="4C4A1DB0C97D860263E9FAEAFE28FA79" box="[425,587,1403,1429]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="denigricans">
|
||
<emphasis id="B93EBA21C97D860263E9FAEAFE28FA79" box="[425,587,1403,1429]" italics="true" pageId="4" pageNumber="175">T. denigricans</emphasis>
|
||
</taxonomicName>
|
||
is grey). Some other species appear to be more varied in their bark colour, often ranging from grey or brown to black such as
|
||
<taxonomicName id="4C4A1DB0C97D860260F6FA36F8B2FA55" authority="(Sprengel 1813: 18) Merrill (1937: 131)" authorityName="Merrill" authorityPageNumber="131" authorityYear="1937" baseAuthorityName="Sprengel" baseAuthorityPageNumber="18" baseAuthorityYear="1813" box="[694,1233,1439,1465]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="timon">
|
||
<emphasis id="B93EBA21C97D860260F6FA36FF73FA55" box="[694,784,1440,1465]" italics="true" pageId="4" pageNumber="175">T. timon</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDB1BC2C97D8602615DFA09FF88FA55" author="Sprengel, C. P. J." box="[797,1003,1439,1465]" pageId="4" pageNumber="175" refId="ref7245" refString="Sprengel, C. P. J. (1813) Plantarum Minus Cognitarum Pugillus, vol. 1. Apud C. A. Kummelium, Halae, 66 pp." type="book" year="1813">Sprengel 1813: 18</bibRefCitation>
|
||
)
|
||
<bibRefCitation id="EFDB1BC2C97D860261B9FA09F8B2FA55" author="Merrill, E. D." box="[1017,1233,1439,1465]" pageId="4" pageNumber="175" pagination="130 - 132" refId="ref6668" refString="Merrill, E. D. (1937) New names in Timonius. Journal of the Arnold Arboretum 18: 130 - 132. [https: // www. jstor. org / stable / 43780822] https: // doi. org / 10.5962 / p. 185362" type="journal article" year="1937">Merrill (1937: 131)</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4A1DB0C97D86026745FA36FD98FA31" authority="Merrill & L. M. Perry (1994: 78)" authorityName="Merrill & L. M. Perry" authorityPageNumber="78" authorityYear="1994" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="trichocladus">
|
||
<emphasis id="B93EBA21C97D86026745FA36F9C8FA55" box="[1285,1451,1439,1465]" italics="true" pageId="4" pageNumber="175">T. trichocladus</emphasis>
|
||
Merrill & L.M.Perry (1994: 78)
|
||
</taxonomicName>
|
||
from the Papuasian region as well as
|
||
<taxonomicName id="4C4A1DB0C97D86026181FA52F93AFA31" authority="Junhao Chen (2014: 139)" authorityName="Junhao Chen" authorityPageNumber="139" authorityYear="2014" box="[961,1369,1475,1501]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="abanii">
|
||
<emphasis id="B93EBA21C97D86026181FA52F84BFA31" box="[961,1064,1475,1501]" italics="true" pageId="4" pageNumber="175">T. abanii</emphasis>
|
||
Junhao
|
||
<bibRefCitation id="EFDB1BC2C97D860266CCFA55F93AFA31" author="Chen, J. & Wong, K. M. & van der Ent, A. & Tan, H. T. W." box="[1164,1369,1475,1501]" pageId="4" pageNumber="175" pagination="138 - 150" refId="ref5765" refString="Chen, J., Wong, K. M., van der Ent, A. & Tan, H. T. W. (2014) Nine new species of Timonius (Rubiaceae) from Kinabalu Park, Borneo. Phytotaxa 181: 138 - 150. https: // doi. org / 10.11646 / phytotaxa. 181.3.2" type="journal article" year="2014">Chen (2014: 139)</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4A1DB0C97D860267D7FA52FE5FF9ED" authority="Junhao Chen (2014: 146)" authorityName="Junhao Chen" authorityPageNumber="146" authorityYear="2014" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="species" species="ophioliticus">
|
||
<emphasis id="B93EBA21C97D860267D7FA52FD6FF9ED" italics="true" pageId="4" pageNumber="175">T. ophioliticus</emphasis>
|
||
Junhao
|
||
<bibRefCitation id="EFDB1BC2C97D86026330FA71FE5FF9ED" author="Chen, J. & Wong, K. M. & van der Ent, A. & Tan, H. T. W." box="[368,572,1511,1537]" pageId="4" pageNumber="175" pagination="138 - 150" refId="ref5765" refString="Chen, J., Wong, K. M., van der Ent, A. & Tan, H. T. W. (2014) Nine new species of Timonius (Rubiaceae) from Kinabalu Park, Borneo. Phytotaxa 181: 138 - 150. https: // doi. org / 10.11646 / phytotaxa. 181.3.2" type="journal article" year="2014">Chen (2014: 146)</bibRefCitation>
|
||
</taxonomicName>
|
||
from Borneo island. However, these species either belong to very different subgenera, or have been recently revised and display different characteristics (
|
||
<bibRefCitation id="EFDB1BC2C97D8602664AF99DF894F9C9" author="Chen, J. & Wong, K. M." box="[1034,1271,1547,1574]" pageId="4" pageNumber="175" refId="ref5828" refString="Chen, J. & Wong, K. M. (2023) Timonius in Borneo, with 68 new species: A little-studied, hyper-diverse rainforest plant genus in the Guettardeae (Rubiaceae) and implications for the taxonomy of an alliance widespread throughout Southeast Asia. Natural History Publications (Borneo), Kota Kinabalu, 424 pp." type="book" year="2023">Chen & Wong 2023</bibRefCitation>
|
||
). Besides going through taxonomic literature, the search for
|
||
<taxonomicName id="4C4A1DB0C97D860260C4F9A6FE88F9A5" authorityName="Candolle" authorityYear="1830" box="[644,747,1584,1609]" class="Magnoliopsida" family="Rubiaceae" genus="Timonius" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="175" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93EBA21C97D860260C4F9A6FE88F9A5" box="[644,747,1584,1609]" italics="true" pageId="4" pageNumber="175">Timonius</emphasis>
|
||
</taxonomicName>
|
||
with black bark was done through IPNI, focusing on species epithets that contain Latin or Greek terms for ‘black’ which are ‘niger’ and ‘melano-’, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |