treatments-xml/data/03/84/87/038487E62F2D2434FF12B7EAFD6BFF0D.xml
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<document id="C7F32F3C13CADA2F6F936492D8DFBA4F" ID-CLB-Dataset="6241" ID-DOI="10.11646/zootaxa.4973.1.1" ID-GBIF-Dataset="d0967b08-98d1-4ccd-b472-a678e0e57eda" ID-ISSN="1175-5326" ID-Zenodo-Dep="4771958" ID-ZooBank="42EB4BF2-A571-4894-9EEF-783649A27E4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1621409551899" checkinUser="plazi" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander &amp; Auffenberg, Kurt" docDate="2021" docId="038487E62F2D2434FF12B7EAFD6BFF0D" docLanguage="en" docName="zootaxa.4973.1.1.pdf" docOrigin="Zootaxa 4973 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chamalycaeus irmatallus Pall-Gergely 2021, n. sp." docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="FFBDFF9E2F20243AFF85B51EFF9BFFD8" masterDocTitle="Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)" masterLastPageNumber="61" masterPageNumber="1" pageNumber="14" updateTime="1698940034724" updateUser="ExternalLinkService">
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<mods:title id="336F3CD0E23A1945AEB5FF4CC84FFF63">Nineteen new species of Alycaeidae from Myanmar and Thailand (Gastropoda: Caenogastropoda: Cyclophoroidea)</mods:title>
</mods:titleInfo>
<mods:name id="B14620B2CDC04EDF51C9E704A5ED6F44" type="personal">
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<mods:namePart id="32F18253E0B6C890B0D91365AAC11680">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="E095475CE29BBAE7EDEDAD095DF39D6F">Hunyadi, András</mods:namePart>
<mods:nameIdentifier id="3DA2EDC1085D1F551D27CF6D37013214" type="ORCID">0000-0003-3488-2817</mods:nameIdentifier>
<mods:nameIdentifier id="D3F349E06445A03A735A01CEE115FD72" type="email">hunand@freemail.hu</mods:nameIdentifier>
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<mods:namePart id="F607042C15C8249AF9FA843197DEC31F">Grego, Jozef</mods:namePart>
<mods:nameIdentifier id="8C7AD27EE433D1CDD925B0B9F640DE0E" type="ORCID">0000-0002-4977-0415</mods:nameIdentifier>
<mods:nameIdentifier id="ECF0C50E6A809E4BC2D3E2FD89CDE6DD" type="email">jozef.grego@gmail.com</mods:nameIdentifier>
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<mods:namePart id="1BBED30BB0FE8699F2E58E26450D3FCB">Reischütz, Alexander</mods:namePart>
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<mods:nameIdentifier id="BE4EBAAC64D2CF973697E247E16D142D" type="email">alexander.reischuetz@gmx.at</mods:nameIdentifier>
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<mods:namePart id="35A282B22219F4ED9DB90D895FDD516A">Auffenberg, Kurt</mods:namePart>
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<mods:nameIdentifier id="0E467B8626CD8DF5F36D66307B930024" type="email">kurtauffenberg@gmail.com</mods:nameIdentifier>
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<mods:title id="361B683FF15353C531AAA8CC1618AD79">Zootaxa</mods:title>
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<mods:date id="87158193525AA27638B165534530AC0D">2021</mods:date>
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<mods:number id="64FFDCF3EEA6ED26A7BB7D59318AB9A3">2021-05-18</mods:number>
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<mods:number id="59F4B5B94DF41CC6E66D93FD7B2E8ACD">4973</mods:number>
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<treatment id="038487E62F2D2434FF12B7EAFD6BFF0D" ID-DOI="http://doi.org/10.5281/zenodo.4771756" ID-GBIF-Taxon="182640515" ID-Zenodo-Dep="4771756" LSID="urn:lsid:plazi:treatment:038487E62F2D2434FF12B7EAFD6BFF0D" httpUri="http://treatment.plazi.org/id/038487E62F2D2434FF12B7EAFD6BFF0D" lastPageId="14" lastPageNumber="14" pageId="13" pageNumber="14">
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<heading id="D0DA819C2F2D2437FF12B7EAFD20FCD7" bold="true" box="[151,699,756,783]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="4">
<emphasis id="B959EAE22F2D2437FF12B7EAFD20FCD7" bold="true" box="[151,699,756,783]" pageId="13" pageNumber="14">
<taxonomicName id="4C2D4D732F2D2437FF12B7EAFDEAFCD7" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2021" box="[151,625,756,783]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="irmatallus" status="sp. nov.">
<emphasis id="B959EAE22F2D2437FF12B7EAFE57FCD7" bold="true" box="[151,460,756,783]" italics="true" pageId="13" pageNumber="14">Chamalycaeus irmatallus</emphasis>
Páll-Gergely
</taxonomicName>
<taxonomicNameLabel id="A26A57992F2D2437FDFCB7EBFD20FCD7" box="[633,699,757,783]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C337657B2F2D2437FF12B606FE81FCEB" box="[151,282,792,819]" pageId="13" pageNumber="14" type="description">
<paragraph id="8B9236F02F2D2437FF12B606FE81FCEB" blockId="13.[151,699,756,819]" box="[151,282,792,819]" pageId="13" pageNumber="14">
<figureCitation id="13162A752F2D2437FF12B606FF45FCEB" box="[151,222,792,819]" captionStart="FIGURE 9" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[151,1436,237,1672]" captionTargetId="figure-56@14.[151,1436,237,1672]" captionTargetPageId="14" captionText="FIGURE 9. Chamalycaeus irmatallus Páll-Gergely, n. sp. A: holotype (UF 279508); B: paratype (UF 551216)." figureDoi="http://doi.org/10.5281/zenodo.4771980" httpUri="https://zenodo.org/record/4771980/files/figure.png" pageId="13" pageNumber="14">Figs 9</figureCitation>
,
<figureCitation id="13162A752F2D2437FF6FB607FE81FCEB" box="[234,282,793,819]" captionStart="FIGURE 10" captionStartId="15.[151,250,1811,1835]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-18@15.[151,1436,181,1787]" captionTargetPageId="15" captionText="FIGURE 10. Shells of Chamalycaeus Möllendorff, 1897 species. A: Chamalycaeus irmatallus Páll-Gergely, n. sp., NHMUK 1906.4.4.71; B: Chamalycaeus (?) armillatus (Benson, 1856), holotype (UMZC 102995)." figureDoi="http://doi.org/10.5281/zenodo.4771982" httpUri="https://zenodo.org/record/4771982/files/figure.png" pageId="13" pageNumber="14">10A</figureCitation>
</paragraph>
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<subSubSection id="C337657B2F2D2437FF12B67EFA82FBD3" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8B9236F02F2D2437FF12B67EFA82FBD3" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF12B67EFE19FCA2" bold="true" box="[151,386,864,890]" pageId="13" pageNumber="14">Material examined.</emphasis>
<materialsCitation id="3B453CAD2F2D2437FE0EB67FFA94FC47" ID-GBIF-Occurrence="3128117390" accessionNumber="UF 279508" collectingDate="1987-06-03" collectionCode="UF" collectorName="F. G. Thompson" country="Thailand" elevation="100" latitude="8.659717" location="Na San" longLatPrecision="1" longitude="99.39805" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Surat Thani" typeStatus="holotype">
<emphasis id="B959EAE22F2D2437FE0EB67FFD9AFCA3" bold="true" box="[395,513,865,891]" pageId="13" pageNumber="14">
<typeStatus id="549688522F2D2437FE0EB67FFE67FCA3" box="[395,508,865,891]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F33A76602F2D2437FD8CB67EFDF4FCA2" box="[521,623,864,890]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
,
<collectingRegion id="49E9F8122F2D2437FDF9B67FFCF4FCA2" box="[636,879,864,891]" country="Thailand" name="Surat Thani" pageId="13" pageNumber="14">Surat Thani Province</collectingRegion>
, limestone mtn.,
<quantity id="4CD59B152F2D2437FBB1B67EFBF5FCA2" box="[1076,1134,864,890]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="14" unit="km" value="6.0">6 km</quantity>
S of
<location id="8EF2602B2F2D2437FB2AB67FFA9EFCA3" LSID="urn:lsid:plazi:treatment:038487E62F2D2434FF12B7EAFD6BFF0D:8EF2602B2F2D2437FB2AB67FFA9EFCA3" box="[1199,1285,865,891]" country="Thailand" latitude="8.659717" longLatPrecision="1" longitude="99.39805" name="Na San" pageId="13" pageNumber="14" stateProvince="Surat Thani">Na San</location>
,
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,
<geoCoordinate id="EE1950372F2D2437FF12B69BFEB3FC47" box="[151,296,901,927]" degrees="99" direction="east" minutes="23.883" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="99.39805">99°23.883E</geoCoordinate>
,
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<quantity id="4CD59B152F2D2437FEB0B69BFEE5FC47" box="[309,382,901,927]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
,
<date id="FF9310302F2D2437FE4CB69BFDCAFC47" box="[457,593,901,927]" pageId="13" pageNumber="14" value="1987-06-03">
<collectingDate id="EFD7E9D82F2D2437FE4CB69BFDCAFC47" box="[457,593,901,927]" pageId="13" pageNumber="14" value="1987-06-03">3 Jun. 1987</collectingDate>
</date>
,
<collectorName id="26D853262F2D2437FDDBB69BFC96FC46" box="[606,781,900,927]" pageId="13" pageNumber="14">F.G. Thompson</collectorName>
leg.,
<collectionCode id="ED3CAE352F2D2437FCCBB69BFCE9FC47" box="[846,882,901,927]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="13" pageNumber="14" type="Museum">UF</collectionCode>
279508 (D:
<quantity id="4CD59B152F2D2437FB81B69BFBF7FC47" box="[1028,1132,901,927]" metricMagnitude="-3" metricUnit="m" metricValue="2.02" pageId="13" pageNumber="14" unit="mm" value="2.02">2.02 mm</quantity>
, H:
<quantity id="4CD59B152F2D2437FB18B69BFA9CFC47" box="[1181,1287,901,927]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="13" pageNumber="14" unit="mm" value="1.32">1.32 mm</quantity>
)
</materialsCitation>
.
<materialsCitation id="3B453CAD2F2D2437FA9EB69BFD0DFC1B" ID-GBIF-Occurrence="3128117368" accessionNumber="UF 551216" collectingDate="1987-06-03" collectionCode="UF" collectorName="F. G. Thompson" country="Thailand" elevation="100" latitude="8.659717" location="Na San" longLatPrecision="1" longitude="99.39805" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Surat Thani" typeStatus="paratype">
<emphasis id="B959EAE22F2D2437FA9EB69BFA07FC47" bold="true" box="[1307,1436,901,927]" pageId="13" pageNumber="14">
<typeStatus id="549688522F2D2437FA9EB69BFA0CFC47" box="[1307,1431,901,927]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
:
</emphasis>
11 shells, same data as holotype,
<collectionCode id="ED3CAE352F2D2437FD97B6B7FDADFC1B" box="[530,566,937,963]" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="13" pageNumber="14" type="Museum">UF</collectionCode>
551216
</materialsCitation>
.
<emphasis id="B959EAE22F2D2437FD24B6B6FCC4FC1A" bold="true" box="[673,863,936,962]" pageId="13" pageNumber="14">Other material:</emphasis>
<materialsCitation id="3B453CAD2F2D2437FCE2B6B6FEDAFC3F" ID-GBIF-Occurrence="3128117372" collectionCode="NHMUK" collectorName="Blanford" country="Myanmar" location="Pegu" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Bago">
6 shells,
<collectingCountry id="F33A76602F2D2437FC49B6B7FBA0FC1B" box="[972,1083,937,963]" name="Myanmar" pageId="13" pageNumber="14">Myanmar</collectingCountry>
, Thayet-myo,
<collectingRegion id="49E9F8122F2D2437FB5BB6B7FA82FC1B" box="[1246,1305,937,963]" country="Myanmar" name="Bago" pageId="13" pageNumber="14">Pegu</collectingRegion>
, coll.
<collectorName id="26D853262F2D2437FADBB6B6FF52FC3E" pageId="13" pageNumber="14">Blanford</collectorName>
,
<collectionCode id="ED3CAE352F2D2437FF52B6D3FEDAFC3F" box="[215,321,973,999]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="14" type="Museum">NHMUK</collectionCode>
</materialsCitation>
<materialsCitation id="3B453CAD2F2D2437FECFB6D2FA8EFBD3" ID-GBIF-Occurrence="3128117336" collectingDate="2015-02-20" collectionCode="HA" collectorName="A. Hunyadi" country="Thailand" elevation="40" latitude="8.463817" location="Tham Nam Phud" longLatPrecision="1" longitude="98.54362" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Phangnga">
1906.4.4.71; 1 shell,
<collectingCountry id="F33A76602F2D2437FDB8B6D2FD38FC3E" box="[573,675,972,998]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
,
<collectingRegion id="49E9F8122F2D2437FD34B6D2FC04FC3E" box="[689,927,972,999]" country="Thailand" name="Phangnga" pageId="13" pageNumber="14">Phang Nga Province</collectingRegion>
,
<collectingRegion id="49E9F8122F2D2437FC2BB6D2FBB5FC3F" box="[942,1070,972,999]" country="Thailand" name="Phangnga" pageId="13" pageNumber="14">Phang Nga</collectingRegion>
,
<location id="8EF2602B2F2D2437FBB9B6D2FA98FC3E" LSID="urn:lsid:plazi:treatment:038487E62F2D2434FF12B7EAFD6BFF0D:8EF2602B2F2D2437FBB9B6D2FA98FC3E" box="[1084,1283,972,999]" country="Thailand" latitude="8.463817" longLatPrecision="1" longitude="98.54362" name="Tham Nam Phud" pageId="13" pageNumber="14" stateProvince="Phangnga">Tham Nam Phud</location>
,
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,
<geoCoordinate id="EE1950372F2D2437FF12B6EEFEB2FBD2" box="[151,297,1008,1034]" degrees="98" direction="east" minutes="32.617" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="98.54362">98°32.617E</geoCoordinate>
,
<elevation id="0000D1C32F2D2437FEB1B6EFFE33FBD2" box="[308,424,1008,1035]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="14" unit="m" value="40.0">
<quantity id="4CD59B152F2D2437FEB1B6EFFEF6FBD3" box="[308,365,1009,1035]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="14" unit="m" value="40.0">40 m</quantity>
a.s.l.
</elevation>
(locality code: 2015/33),
<date id="FF9310302F2D2437FD4FB6EFFCFBFBD3" box="[714,864,1008,1035]" pageId="13" pageNumber="14" value="2015-02-20">
<collectingDate id="EFD7E9D82F2D2437FD4FB6EFFCFBFBD3" box="[714,864,1008,1035]" pageId="13" pageNumber="14" value="2015-02-20">20 Feb. 2015</collectingDate>
</date>
,
<collectorName id="26D853262F2D2437FCECB6EFFC70FBD2" box="[873,1003,1008,1035]" pageId="13" pageNumber="14">A. Hunyadi</collectorName>
leg., around the cave,
<collectionCode id="ED3CAE352F2D2437FB6CB6EFFA8EFBD3" box="[1257,1301,1009,1035]" pageId="13" pageNumber="14">HA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F2D2437FF42B10BFDA0FB8B" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8B9236F02F2D2437FF42B10BFDA0FB8B" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B10BFED5FBF7" bold="true" box="[199,334,1045,1071]" pageId="13" pageNumber="14">Etymology.</emphasis>
The specific epithet is an anagram for
<taxonomicName id="4C2D4D732F2D2437FC82B10AFCE1FBF6" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[775,890,1044,1070]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="armillatus">
<emphasis id="B959EAE22F2D2437FC82B10AFCE1FBF6" box="[775,890,1044,1070]" italics="true" pageId="13" pageNumber="14">armillatus</emphasis>
</taxonomicName>
, a species with which this new species has been confused (see Differential diagnosis).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F2D2437FF42B142FAD1FB43" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8B9236F02F2D2437FF42B142FAD1FB43" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B142FED9FBAE" bold="true" box="[199,322,1116,1142]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
A tiny
<taxonomicName id="4C2D4D732F2D2437FE1EB142FD01FBAE" box="[411,666,1116,1142]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B959EAE22F2D2437FE1EB142FDDBFBAE" box="[411,576,1116,1142]" italics="true" pageId="13" pageNumber="14">Chamalycaeus</emphasis>
species
</taxonomicName>
with glossy shell, irregularly, finely ribbed R1, very short R2 with blunt, low ribs, and somewhat longer R3 (the two latter regions are shorter than a quarter whorl combined).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F2D2437FF42B1BAFE4FF992" pageId="13" pageNumber="14" type="description">
<paragraph id="8B9236F02F2D2437FF42B1BAFB38F9DA" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B1BAFEC2FB66" bold="true" box="[199,345,1188,1214]" pageId="13" pageNumber="14">Description.</emphasis>
Shell off-white, rather glossy, although available shells are somewhat eroded; shell outline slightly oval in dorsal view; spire slightly elevated, conical-domed; body whorl somewhat “parabolic” in apertural view; protoconch rather low, glossy, spiral striae lacking, 1.25 whorls; R1 of 1.51.75 whorls, with very indistinct, irregular, low and rounded, relatively wide ribs, and some weak spiral striation; boundary between R1 and R2 distinct due to change in intensity and strength of ribbing; R2 short, only ca. 1620 ribs; R2+R3 less than quarter whorl (ca. 7080˚); R3 up to twice as long as R2; boundary between R2 and R3 clear due to change in rib morphology, although constriction very shallow; R3 with very fine, rounded ribs, and low, convex, elongated central swelling; aperture round, slightly oblique to shell axis; boundary between inner and outer peristomes distinct, inner peristome strongly protruding, thickened, slightly expanded; outer peristome slightly weaker or as strong, also expanded, only slightly reflected toward umbilicus; umbilicus relatively narrow, ca. one third of shell width.
</paragraph>
<paragraph id="8B9236F02F2D2437FF42B313FCBEF9FF" blockId="13.[151,1437,864,2043]" box="[199,805,1548,1575]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B313FEE5F9FF" bold="true" box="[199,382,1549,1575]" pageId="13" pageNumber="14">Measurements.</emphasis>
D:
<quantity id="4CD59B152F2D2437FE22B312FDD7F9FF" box="[423,588,1548,1575]" metricMagnitude="-3" metricUnit="m" metricValue="1.885" metricValueMax="2.02" metricValueMin="1.75" pageId="13" pageNumber="14" unit="mm" value="1.885" valueMax="2.02" valueMin="1.75">1.752.02 mm</quantity>
, H:
<quantity id="4CD59B152F2D2437FDFEB312FCBAF9FF" box="[635,801,1548,1575]" metricMagnitude="-3" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.32" metricValueMin="1.15" pageId="13" pageNumber="14" unit="mm" value="1.2349999999999999" valueMax="1.32" valueMin="1.15">1.151.32 mm</quantity>
.
</paragraph>
<paragraph id="8B9236F02F2D2437FF42B32EFE4FF992" blockId="13.[151,1437,864,2043]" box="[199,468,1584,1610]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B32EFECCF992" bold="true" box="[199,343,1584,1610]" pageId="13" pageNumber="14">Operculum.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F2D2437FF42B34AFCDCF96E" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8B9236F02F2D2437FF42B34AFCDCF96E" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B34AFE49F9B6" bold="true" box="[199,466,1620,1646]" pageId="13" pageNumber="14">Differential diagnosis.</emphasis>
<taxonomicName id="4C2D4D732F2D2437FE59B34AFC21F9B7" authority="(Benson, 1856)" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[476,954,1620,1647]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="armillatus">
<emphasis id="B959EAE22F2D2437FE59B34AFD64F9B6" box="[476,767,1620,1646]" italics="true" pageId="13" pageNumber="14">Chamalycaeus armillatus</emphasis>
(
<bibRefCitation id="EFBC4B012F2D2437FC97B34BFC29F9B7" author="Benson, W. H." box="[786,946,1620,1647]" pageId="13" pageNumber="14" pagination="225 - 233" refId="ref24490" refString="Benson, W. H. (1856) Characters of seventeen new forms of the Cyclostomacea from the British provinces of Burmah, collected by W. Theobald, Jun., Esq. The Annals and Magazine of Natural History, Series 2, 17, 225 - 233. https: // doi. org / 10.1080 / 00222935608697501" type="journal article" year="1856">Benson, 1856</bibRefCitation>
)
</taxonomicName>
(material examined: Thayet-Mio, UMZC 102995,
<typeStatus id="549688522F2D2437FF7DB366FEC6F94A" box="[248,349,1656,1682]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
,
<figureCitation id="13162A752F2D2437FEE2B366FE53F94B" box="[359,456,1656,1683]" captionStart="FIGURE 10" captionStartId="15.[151,250,1811,1835]" captionTargetBox="[151,1436,181,1787]" captionTargetId="figure-18@15.[151,1436,181,1787]" captionTargetPageId="15" captionText="FIGURE 10. Shells of Chamalycaeus Möllendorff, 1897 species. A: Chamalycaeus irmatallus Páll-Gergely, n. sp., NHMUK 1906.4.4.71; B: Chamalycaeus (?) armillatus (Benson, 1856), holotype (UMZC 102995)." figureDoi="http://doi.org/10.5281/zenodo.4771982" httpUri="https://zenodo.org/record/4771982/files/figure.png" pageId="13" pageNumber="14">Fig. 10B</figureCitation>
) has a more rounded body whorl, longer R2 and R3, and a thicker peristome with greater distance between the edges of the inner and outer peristomes.
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F2D2437FF42B3DEFAC7F8FB" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B9236F02F2D2437FF42B3DEFAC7F8FB" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B3DEFEFAF902" bold="true" box="[199,353,1728,1754]" pageId="13" pageNumber="14">Distribution.</emphasis>
The
<typeStatus id="549688522F2D2437FE1CB3DFFE52F903" box="[409,457,1729,1755]" pageId="13" pageNumber="14">type</typeStatus>
locality is in Thailands Surat Thani Province. A single shell was collected in the neighbouring Phang Nga Province (locality code: 2015/33, coll. HA). Shells collected more than
<quantity id="4CD59B152F2D2437FB4CB3FBFAACF926" box="[1225,1335,1764,1791]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="km" value="1000.0">1,000 km</quantity>
north in Thayet,
<collectingCountry id="F33A76602F2D2437FF76B217FEFBF8FB" box="[243,352,1801,1827]" name="Myanmar" pageId="13" pageNumber="14">Myanmar</collectingCountry>
are so similar to the Thai populations that they could not be separated (see below) (
<figureCitation id="13162A752F2D2437FA8FB216FAD5F8FB" box="[1290,1358,1800,1827]" captionStart="FIGURE 3" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1436,181,1461]" captionTargetId="figure-18@7.[151,1436,181,1461]" captionTargetPageId="7" captionText="FIGURE 3. Distribution of alycaeid species in southern Thailand. Filled triangle: Dicharax ovatus Páll-Gergely &amp; Auffenberg, n. sp., inverted filled triangle: Dicharax subroseus Páll-Gergely, n. sp., inverted empty triangles: Chamalycaeus irmatallus Páll-Gergely, n. sp., empty circles: C. krabiensis Páll-Gergely &amp; A. Reischütz, n. sp., filled circle: Chamalycaeus serratus Páll-Gergely &amp; Auffenberg, n. sp., empty triangles: Chamalycaeus hirpex Páll-Gergely &amp; A. Reischütz, n. sp., filled square: Chamalycaeus buaboke Páll-Gergely &amp; A. Reischütz, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4771964" httpUri="https://zenodo.org/record/4771964/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C337657B2F2D2434FF42B233FD6BFF0D" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8B9236F02F2D2437FF42B233FD1DF857" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
<emphasis id="B959EAE22F2D2437FF42B233FEA0F89F" bold="true" box="[199,315,1837,1863]" pageId="13" pageNumber="14">Remarks.</emphasis>
<taxonomicName id="4C2D4D732F2D2437FEC4B232FDFBF89E" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[321,608,1836,1862]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="armillatus">
<emphasis id="B959EAE22F2D2437FEC4B232FDFBF89E" box="[321,608,1836,1862]" italics="true" pageId="13" pageNumber="14">Chamalycaeus armillatus</emphasis>
</taxonomicName>
was described from an eroded specimen. Some spiral striations are visible on the
<typeStatus id="549688522F2D2437FF66B24EFED3F8B2" box="[227,328,1872,1898]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, but they might be the part of the lower shell layer. Thus, this species was provisionally assigned to
<taxonomicName id="4C2D4D732F2D2437FF12B26AFEA7F856" authorityName="Mollendorff" authorityYear="1897" box="[151,316,1908,1934]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="B959EAE22F2D2437FF12B26AFEA7F856" box="[151,316,1908,1934]" italics="true" pageId="13" pageNumber="14">Chamalycaeus</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFBC4B012F2D2437FEE5B26AFD19F857" author="Pall-Gergely, B. &amp; Sajan, S. &amp; Tripathy, B. &amp; Meng, K. &amp; Asami, T. &amp; Ablett, J." box="[352,642,1908,1935]" pageId="13" pageNumber="14" pagination="1 - 220" refId="ref26053" refString="Pall-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T. &amp; Ablett, J. (2020) Genus-level revision of the Alycaeidae, with a catalog of the species (Gastropoda: Cyclophoroidea). ZooKeys, 981, 1 - 220. https: // doi. org / 10.3897 / zookeys. 981.53583" type="journal article" year="2020">
Páll-Gergely
<emphasis id="B959EAE22F2D2437FE70B26BFDB5F856" box="[501,558,1908,1934]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
(2020)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B9236F02F2D2437FF42B286FDB2F80F" blockId="13.[151,1437,864,2043]" pageId="13" pageNumber="14">
Specimens from Thayet (NHMUK 1906.4.4.71, labelled as
<taxonomicName id="4C2D4D732F2D2437FCEDB286FC40F86A" baseAuthorityName="Benson" baseAuthorityYear="1856" box="[872,987,1944,1970]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="armillatus">
<emphasis id="B959EAE22F2D2437FCEDB286FC40F86A" box="[872,987,1944,1970]" italics="true" pageId="13" pageNumber="14">armillatus</emphasis>
</taxonomicName>
) have slightly less densely arranged R2 ribs than typical
<taxonomicName id="4C2D4D732F2D2437FED6B2A3FE7DF80E" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg, 2021" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg" authorityYear="2021" box="[339,486,1980,2007]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="irmatallus" status="sp. nov.">
<emphasis id="B959EAE22F2D2437FED6B2A3FE7DF80E" box="[339,486,1980,2007]" italics="true" pageId="13" pageNumber="14">C. irmatallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57992F2D2437FE68B2A3FDB2F80F" box="[493,553,1981,2007]" pageId="13" pageNumber="14" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9236F02F2D2434FF42B2FFFD6BFF0D" blockId="13.[151,1437,864,2043]" lastBlockId="14.[151,1436,151,213]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
A single, eroded shell from Tham Nam Phud (
<collectingCountry id="F33A76602F2D2437FD4BB2FEFCAEF823" box="[718,821,2016,2043]" name="Thailand" pageId="13" pageNumber="14">Thailand</collectingCountry>
) is somewhat larger than the
<typeStatus id="549688522F2D2437FB05B2FEFB79F822" box="[1152,1250,2016,2042]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2D4D732F2D2437FA8DB2FFFA07F822" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg, 2021" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg" authorityYear="2021" box="[1288,1436,2016,2043]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="irmatallus" status="sp. nov.">
<emphasis id="B959EAE22F2D2437FA8DB2FFFA07F822" box="[1288,1436,2016,2043]" italics="true" pageId="13" pageNumber="14">C. irmatallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57992F2E2434FF12B589FF4DFF69" box="[151,214,151,177]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
, and has a longer R2. Since all other shell characters are identical, we provisionally identify the Tham Nam Phud and Thayet specimens as
<taxonomicName id="4C2D4D732F2E2434FE70B5A5FD36FF0D" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg, 2021" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Auffenberg" authorityYear="2021" box="[501,685,187,213]" class="Gastropoda" family="Cyclophoridae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="irmatallus" status="sp. nov.">
<emphasis id="B959EAE22F2E2434FE70B5A5FD94FF0D" box="[501,527,187,213]" italics="true" pageId="14" pageNumber="15">C.</emphasis>
cf.
<emphasis id="B959EAE22F2E2434FDBFB5A5FD36FF0D" box="[570,685,187,213]" italics="true" pageId="14" pageNumber="15">irmatallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57992F2E2434FD31B5A5FD6BFF0D" box="[692,752,187,213]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>