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<document id="9401EE296E2571A63AF9217BA52E99EC" ID-DOI="10.5281/zenodo.191495" ID-GBIF-Dataset="bc9380c5-d4ec-4609-8b91-3e55e1da1b03" ID-ISSN="1175-5326" ID-Zenodo-Dep="191495" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461129398802" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M. &amp; Walter, David E." docDate="2009" docId="E24E87E38E41C83165DB5037FCF98A57" docLanguage="en" docName="zt02294p061.pdf" docOrigin="Zootaxa 2294" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Unduloribates Balogh 1943" docType="treatment" docVersion="8" lastPageNumber="50" masterDocId="1E77FF9B8E40C832654C5666FFF98B24" masterDocTitle="Unduloribates from North America (Acari, Oribatida, Unduloribatidae)" masterLastPageNumber="61" masterPageNumber="47" pageNumber="48" updateTime="1698597348471" updateUser="plazi">
<mods:mods id="C0B43A24786C13D613429BD7A50188ED" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B369C691655DFA30DF2B521EC417F89F">Unduloribates from North America (Acari, Oribatida, Unduloribatidae)</mods:title>
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<mods:namePart id="A28B564E3512B187AC5BBDB5C79C1F50">Behan-Pelletier, Valerie M.</mods:namePart>
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<mods:namePart id="81EA07E16228A4B436DA9F4CA770389E">Walter, David E.</mods:namePart>
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<mods:date id="D3D3E608D4F2CD0A4166771335E245C4">2009</mods:date>
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<mods:number id="45453725E3CEA7386B8358AD087B0B43">2294</mods:number>
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<mods:classification id="B9E0245E5CDD0484D06B64E2068E1643">journal article</mods:classification>
<mods:identifier id="68C9C10B25B68FD092F50EA1C96867E0" type="DOI">10.5281/zenodo.191495</mods:identifier>
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<treatment id="E24E87E38E41C83165DB5037FCF98A57" ID-DOI="http://doi.org/10.5281/zenodo.5658737" ID-GBIF-Taxon="119626699" ID-Zenodo-Dep="5658737" LSID="urn:lsid:plazi:treatment:E24E87E38E41C83165DB5037FCF98A57" httpUri="http://treatment.plazi.org/id/E24E87E38E41C83165DB5037FCF98A57" lastPageId="3" lastPageNumber="50" pageId="1" pageNumber="48">
<subSubSection id="22FD657E8E41C83365DB5037FD638DFF" pageId="1" pageNumber="48" type="nomenclature">
<paragraph id="6A5836F58E41C83365DB5037FE158D4E" blockId="1.[151,492,1616,1643]" box="[151,492,1616,1643]" pageId="1" pageNumber="48">
<heading id="311081998E41C83365DB5037FE158D4E" bold="true" box="[151,492,1616,1643]" fontSize="11" level="1" pageId="1" pageNumber="48" reason="1">
<taxonomicName id="ADE74D768E41C83365DB5037FE158D4E" ID-CoL="84VF" authority="Balogh, 1943" authorityName="Balogh" authorityYear="1943" box="[151,492,1616,1643]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="5893EAE78E41C83365DB5037FE158D4E" bold="true" box="[151,492,1616,1643]" pageId="1" pageNumber="48">
<emphasis id="5893EAE78E41C83365DB5037FEBD8D4F" bold="true" box="[151,324,1617,1643]" italics="true" pageId="1" pageNumber="48">Unduloribates</emphasis>
<bibRefCitation id="0E764B048E41C83364075036FE158D4E" author="Balogh" box="[331,492,1616,1642]" pageId="1" pageNumber="61" refString="Balogh, J. (1943) Systematische Studien uber Siebenburgische Moosmilben. Annales Historico-Naturales Musei Nationalis Hungarici, 36, 34 - 42." type="journal article" year="1943">Balogh, 1943</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="6A5836F58E41C83365DB50FCFD638DFF" blockId="1.[151,1000,1690,1948]" pageId="1" pageNumber="48">
<emphasis id="5893EAE78E41C83365DB50FCFEC78D90" bold="true" box="[151,318,1690,1716]" pageId="1" pageNumber="48">Type-species:</emphasis>
<taxonomicName id="ADE74D768E41C833640950FDFC118D90" authority="Berlese, 1914: 129" authorityName="Berlese" authorityPageNumber="129" authorityYear="1914" box="[325,1000,1690,1716]" class="Arachnida" family="Ceratozetidae" genus="Sphaerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="undulatus" subGenus="Tectoribates">
<emphasis id="5893EAE78E41C833640950FDFCFC8D90" box="[325,773,1691,1716]" italics="true" pageId="1" pageNumber="48">Sphaerozetes (Tectoribates) undulatus</emphasis>
<bibRefCitation id="0E764B048E41C833664050FCFC488D90" author="Berlese" box="[780,945,1690,1716]" pageId="1" pageNumber="61" refString="Berlese, A. (1914) Acari nuovi. Manipulus IX - XIII. Redia, 10, 113 - 150." type="journal article" year="1914">Berlese, 1914</bibRefCitation>
: 129
</taxonomicName>
<taxonomicName id="ADE74D768E41C83365DB50A7FD638DFF" authority="Willmann 1951: 153" authorityName="Willmann" authorityPageNumber="153" authorityYear="1951" box="[151,666,1729,1755]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="5893EAE78E41C83365DB50A7FE5B8DFE" box="[151,418,1729,1754]" italics="true" pageId="1" pageNumber="48">Tectoribates undulatus</emphasis>
<bibRefCitation id="0E764B048E41C83364E550A7FD9A8DFF" author="Willmann" box="[425,611,1729,1755]" pageId="1" pageNumber="61" refString="Willmann, C. (1951) Die hochalpine Milbenfauna der mittleren Hohen Tauern, insbesondere des Großglocknergebietes (Acari). Bonner Zoologische Beitrage, 1, 141 - 176." type="journal article" year="1951">Willmann 1951</bibRefCitation>
: 153
</taxonomicName>
</paragraph>
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<subSubSection id="22FD657E8E41C83065DB508EFB8B890B" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48" type="reference_group">
<paragraph id="6A5836F58E41C83365DB508EFD7C8C26" blockId="1.[151,1000,1690,1948]" box="[151,645,1768,1794]" pageId="1" pageNumber="48">
<treatmentCitationGroup id="4AF711DB8E41C83365DB508EFD7C8C26" box="[151,645,1768,1794]" pageId="1" pageNumber="48">
<taxonomicName id="ADE74D768E41C83365DB508EFD7C8C26" ID-CoL="7DK47" authority="Balogh 1943: 39" authorityName="Balogh" authorityPageNumber="39" authorityYear="1943" box="[151,645,1768,1794]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="5893EAE78E41C83365DB508EFE428C25" box="[151,443,1768,1793]" italics="true" pageId="1" pageNumber="48">Unduloribates undulatus</emphasis>
<treatmentCitation id="EB4610E48E41C833648E508EFD7C8C26" author="Balogh" box="[450,645,1768,1794]" page="39" pageId="1" pageNumber="48" year="1943">
<bibRefCitation id="0E764B048E41C833648E508EFDA48C26" author="Balogh" box="[450,605,1768,1794]" pageId="1" pageNumber="61" refString="Balogh, J. (1943) Systematische Studien uber Siebenburgische Moosmilben. Annales Historico-Naturales Musei Nationalis Hungarici, 36, 34 - 42." type="journal article" year="1943">Balogh 1943</bibRefCitation>
: 39
</treatmentCitation>
</taxonomicName>
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</paragraph>
<paragraph id="6A5836F58E41C83365DB5168FE438C0C" blockId="1.[151,1000,1690,1948]" box="[151,442,1806,1832]" pageId="1" pageNumber="48">
<emphasis id="5893EAE78E41C83365DB5168FE4B8C0C" bold="true" box="[151,434,1806,1832]" pageId="1" pageNumber="48">Other included species</emphasis>
:
</paragraph>
<paragraph id="6A5836F58E41C83365DB5153FDB08C6B" blockId="1.[151,1000,1690,1948]" box="[151,585,1845,1871]" pageId="1" pageNumber="48">
<treatmentCitationGroup id="4AF711DB8E41C83365DB5153FDB08C6B" box="[151,585,1845,1871]" pageId="1" pageNumber="48">
<taxonomicName id="ADE74D768E41C83365DB5153FDB08C6B" ID-CoL="7DJS7" authority="Aoki 1965: 298" authorityName="Aoki" authorityPageNumber="298" authorityYear="1965" box="[151,585,1845,1871]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="hebes">
<emphasis id="5893EAE78E41C83365DB5153FE738C6A" box="[151,394,1845,1870]" italics="true" pageId="1" pageNumber="48">Unduloribates hebes</emphasis>
<treatmentCitation id="EB4610E48E41C83364DD5153FDB08C6B" author="Aoki" box="[401,585,1845,1871]" page="298" pageId="1" pageNumber="48" year="1965">
<bibRefCitation id="0E764B048E41C83364DD5153FDEB8C6B" author="Aoki" box="[401,530,1845,1871]" pageId="1" pageNumber="61" refString="Aoki, J. (1965) Oribatid mites (Acarina: Oribatei) from Himalaya with descriptions of several new species. Journal of the College of Arts and Sciences, Chiba, 4, 289 - 302." type="journal article" year="1965">Aoki 1965</bibRefCitation>
: 298
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6A5836F58E41C83365DB513AFDA08C52" blockId="1.[151,1000,1690,1948]" box="[151,601,1884,1910]" pageId="1" pageNumber="48">
<treatmentCitationGroup id="4AF711DB8E41C83365DB513AFDA08C52" box="[151,601,1884,1910]" pageId="1" pageNumber="48">
<taxonomicName id="ADE74D768E41C83365DB513AFDA08C52" ID-CoL="7DK3V" authority="Piffl 1972: 270" authorityName="Piffl" authorityPageNumber="270" authorityYear="1972" box="[151,601,1884,1910]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="medusa">
<emphasis id="5893EAE78E41C83365DB513AFE588C51" box="[151,417,1884,1909]" italics="true" pageId="1" pageNumber="48">Unduloribates medusa</emphasis>
<treatmentCitation id="EB4610E48E41C83364E4513AFDA08C52" author="Piffl" box="[424,601,1884,1910]" page="270" pageId="1" pageNumber="48" year="1972">
<bibRefCitation id="0E764B048E41C83364E4513AFDDB8C52" author="Piffl" box="[424,546,1884,1910]" pageId="1" pageNumber="61" refString="Piffl, E. (1972). Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl 1972</bibRefCitation>
: 270
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6A5836F58E41C83065DB51E5FEF78B95" blockId="1.[151,1000,1690,1948]" lastBlockId="2.[151,1437,152,559]" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48">
<treatmentCitationGroup id="4AF711DB8E41C83065DB51E5FEF78B95" lastPageId="2" lastPageNumber="49" pageId="1" pageNumber="48">
<taxonomicName id="ADE74D768E41C83365DB51E5FC628CB8" ID-CoL="7DJRV" authority="Nubel-Reidelbach and Woas 1992: 99" authorityName="Nubel-Reidelbach and Woas" authorityPageNumber="99" authorityYear="1992" box="[151,923,1922,1948]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="brevisetosus">
<emphasis id="5893EAE78E41C83365DB51E5FE2E8CB8" box="[151,471,1923,1948]" italics="true" pageId="1" pageNumber="48">Unduloribates brevisetosus</emphasis>
<treatmentCitation id="EB4610E48E41C833649251E4FC628CB8" author="Nubel-Reidelbach" box="[478,923,1922,1948]" page="99" pageId="1" pageNumber="48" year="1992">
<bibRefCitation id="0E764B048E41C833649251E4FC8A8CB8" author="Nubel-Reidelbach" box="[478,883,1922,1948]" pageId="1" pageNumber="61" refString="Nubel-Reidelbach, E. &amp; Woas, S. (1992) Einige basale Arten der cepheiden und der pterogasterinen Entwicklungslinie der Hoheren Oribatiden (Acari, Oribatei). Andrias, Karlsruhe, 9, 75 - 119." type="journal article" year="1992">Nübel-Reidelbach and Woas 1992</bibRefCitation>
: 99
</treatmentCitation>
</taxonomicName>
<emphasis id="5893EAE78E42C83065DB56FEFEF78B95" box="[151,270,152,177]" italics="true" pageId="2" pageNumber="49">Diagnosis</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6A5836F58E42C830658A56D8FB8B890B" blockId="2.[151,1437,152,559]" pageId="2" pageNumber="49">
Species in this genus have the following unique combination of character states: adult with irregularly tuberculate, thick, blocky, birefringent cerotegument; prodorsum with pair of deep vertical incisions in rostral tectum, ending near seta
<emphasis id="5893EAE78E42C8306482576CFE1E8A07" box="[462,487,266,291]" italics="true" pageId="2" pageNumber="49">ro</emphasis>
; with large, broad lamellae, almost touching medially, extending anterior of rostrum; bothridium with internal ring-like thickenings; notogaster without octotaxic system; with lenticulus; with 10 pairs of notogastral setae; pteromorph without hinge; posterior notogastral tectum absent; palpal eupathidium
<emphasis id="5893EAE78E42C830647E571BFE9A8AB2" box="[306,355,381,406]" italics="true" pageId="2" pageNumber="49">acm</emphasis>
attached to solenidion at least along distal third of length; axillary saccule of subcapitulum present; coxisternal setal formula 3-1-2-3, 3-1-3-3 or 3-2-7-7; 1 pair aggenital, 3 pairs of adanal and 2 pairs of anal setae; genital setae neotrichous or not, with 6 to 10 pairs; postanal porose area absent; immatures apheredermous, plicate; larva with 11, nymphs with 15 pairs of gastronotal setae; seta
<emphasis id="5893EAE78E42C83061EB5789FB4F892C" box="[1191,1206,495,520]" italics="true" pageId="2" pageNumber="49">d</emphasis>
absent from genua and tibiae in immatures and adults. Paraproctal atrichosy from larva to deutonymph.
</paragraph>
</subSubSection>
<subSubSection id="22FD657E8E42C83065DB5407FC0C8DDD" pageId="2" pageNumber="49" type="description">
<paragraph id="6A5836F58E42C83065DB5407FEDB895E" blockId="2.[151,1437,609,1785]" box="[151,290,609,634]" pageId="2" pageNumber="49">
<emphasis id="5893EAE78E42C83065DB5407FEDB895E" box="[151,290,609,634]" italics="true" pageId="2" pageNumber="49">Description</emphasis>
</paragraph>
<paragraph id="6A5836F58E42C830658A54EEFC158EBB" blockId="2.[151,1437,609,1785]" pageId="2" pageNumber="49">
<emphasis id="5893EAE78E42C830658A54EEFEF78985" box="[198,270,648,673]" italics="true" pageId="2" pageNumber="49">Adult.</emphasis>
Color of mature adults dark brown. Notogaster and ventral plate with thick, blocky, birefringent cerotegument (
<figureCitation id="F2DC2A708E42C830640254CBFE5C89E3" box="[334,421,685,711]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,272,867,891]" captionTargetBox="[151,1435,151,859]" captionTargetId="figure@5.[151,1436,139,897]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Unduloribates dianae sp. nov., scanning electron micrographs of adult females: 4, ventral aspect; 5, dorsal aspect; 6, notogaster with much of cerotegument removed. Scale bars: 100 µm." httpUri="https://zenodo.org/record/191497/files/figure.png" pageId="2" pageNumber="49">Figs. 5</figureCitation>
,
<figureCitation id="F2DC2A708E42C83064FF54CBFE3D89E3" box="[435,452,685,711]" captionStart="FIGURES 7 – 12" captionStartId="6.[151,271,1895,1919]" captionTargetBox="[151,1436,194,1871]" captionTargetId="figure@6.[151,1436,194,1871]" captionTargetPageId="6" captionText="FIGURES 7 – 12. Unduloribates dianae sp. nov., scanning electron micrographs; figures 7 - 10, 12 of adult females; 7, detail of bothridium and bothridial seta; 8, detail of blocky cerotegument on notogaster; 9, detail of prodorsum and anterior of notogaster; 10, prodorsum, lateral aspect; 11, egg in female hysterosoma; 12, tarsus I and anterior of tibia I. Scale bars: 7, 8, 11, 12, 10 µm; 9, 10, 20 µm." httpUri="https://zenodo.org/record/191498/files/figure.png" pageId="2" pageNumber="49">8</figureCitation>
). Lenticulus present. Legs with weakly developed, tuberculate, non-birefringent cerotegument. All sclerotized integument microtuberculate, with convex polygonal pattern on notogaster (
<figureCitation id="F2DC2A708E42C830602054B2FF508830" captionStart="FIGURES 4 – 6" captionStartId="5.[151,272,867,891]" captionTargetBox="[151,1435,151,859]" captionTargetId="figure@5.[151,1436,139,897]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Unduloribates dianae sp. nov., scanning electron micrographs of adult females: 4, ventral aspect; 5, dorsal aspect; 6, notogaster with much of cerotegument removed. Scale bars: 100 µm." httpUri="https://zenodo.org/record/191497/files/figure.png" pageId="2" pageNumber="49">Fig. 6</figureCitation>
). Dorsophragmata and pleurophragmata present. Prodorsum with large, broad lamellae, almost touching medially, extending anterior of rostrum (
<figureCitation id="F2DC2A708E42C83067365546FD35881E" box="[634,716,800,826]" captionStart="FIGURES 1 – 3" captionStartId="4.[151,271,1925,1949]" captionTargetBox="[207,1392,160,1911]" captionTargetId="figure@4.[197,1403,139,1911]" captionTargetPageId="4" captionText="FIGURES 1 – 3. Unduloribates dianae sp. nov., adult female: 1, dorsal aspect, dorsophragmata indicated by arrow; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed). Scale bars: 100 µm." httpUri="https://zenodo.org/record/191496/files/figure.png" pageId="2" pageNumber="49">Figs. 1</figureCitation>
,
<figureCitation id="F2DC2A708E42C83067975546FD15881E" box="[731,748,800,826]" captionStart="FIGURES 7 – 12" captionStartId="6.[151,271,1895,1919]" captionTargetBox="[151,1436,194,1871]" captionTargetId="figure@6.[151,1436,194,1871]" captionTargetPageId="6" captionText="FIGURES 7 – 12. Unduloribates dianae sp. nov., scanning electron micrographs; figures 7 - 10, 12 of adult females; 7, detail of bothridium and bothridial seta; 8, detail of blocky cerotegument on notogaster; 9, detail of prodorsum and anterior of notogaster; 10, prodorsum, lateral aspect; 11, egg in female hysterosoma; 12, tarsus I and anterior of tibia I. Scale bars: 7, 8, 11, 12, 10 µm; 9, 10, 20 µm." httpUri="https://zenodo.org/record/191498/files/figure.png" pageId="2" pageNumber="49">9</figureCitation>
). Rostrum broadly rounded (
<figureCitation id="F2DC2A708E42C830610A5546FB63881E" box="[1094,1178,800,826]" captionStart="FIGURES 1 – 3" captionStartId="4.[151,271,1925,1949]" captionTargetBox="[207,1392,160,1911]" captionTargetId="figure@4.[197,1403,139,1911]" captionTargetPageId="4" captionText="FIGURES 1 – 3. Unduloribates dianae sp. nov., adult female: 1, dorsal aspect, dorsophragmata indicated by arrow; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed). Scale bars: 100 µm." httpUri="https://zenodo.org/record/191496/files/figure.png" pageId="2" pageNumber="49">Figs. 2</figureCitation>
,
<figureCitation id="F2DC2A708E42C83061E45546FB40881E" box="[1192,1209,800,826]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,272,867,891]" captionTargetBox="[151,1435,151,859]" captionTargetId="figure@5.[151,1436,139,897]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Unduloribates dianae sp. nov., scanning electron micrographs of adult females: 4, ventral aspect; 5, dorsal aspect; 6, notogaster with much of cerotegument removed. Scale bars: 100 µm." httpUri="https://zenodo.org/record/191497/files/figure.png" pageId="2" pageNumber="49">4</figureCitation>
,
<figureCitation id="F2DC2A708E42C830618A5546FB1F881E" box="[1222,1254,800,826]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,272,1296,1320]" captionTargetBox="[150,1436,741,1284]" captionTargetId="figure@7.[145,1440,741,1286]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Unduloribates dianae sp. nov., scanning electron micrographs of adult females; 13, gnathosoma, ventral aspect; 14, gnathosoma, anterior aspect; 15, pedipalp with lateral detail of gena and rutellum. Scale bars: 20 µm." httpUri="https://zenodo.org/record/191499/files/figure.png" pageId="2" pageNumber="49">13</figureCitation>
). Deep vertical incisions present laterally on rostral tectum, ending near seta
<emphasis id="5893EAE78E42C83066C95520FC67887B" box="[901,926,838,863]" italics="true" pageId="2" pageNumber="49">ro</emphasis>
; oblique genal notch absent (
<figureCitation id="F2DC2A708E42C830604B5520FAA78844" box="[1287,1374,838,864]" captionStart="FIGURES 1 – 3" captionStartId="4.[151,271,1925,1949]" captionTargetBox="[207,1392,160,1911]" captionTargetId="figure@4.[197,1403,139,1911]" captionTargetPageId="4" captionText="FIGURES 1 – 3. Unduloribates dianae sp. nov., adult female: 1, dorsal aspect, dorsophragmata indicated by arrow; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed). Scale bars: 100 µm." httpUri="https://zenodo.org/record/191496/files/figure.png" pageId="2" pageNumber="49">Figs. 3</figureCitation>
,
<figureCitation id="F2DC2A708E42C83060205520FA778844" box="[1388,1422,838,864]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,272,1296,1320]" captionTargetBox="[150,1436,741,1284]" captionTargetId="figure@7.[145,1440,741,1286]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Unduloribates dianae sp. nov., scanning electron micrographs of adult females; 13, gnathosoma, ventral aspect; 14, gnathosoma, anterior aspect; 15, pedipalp with lateral detail of gena and rutellum. Scale bars: 20 µm." httpUri="https://zenodo.org/record/191499/files/figure.png" pageId="2" pageNumber="49">14</figureCitation>
). Bothridium with internal ring-like thickenings. Porose areas Am and Ah present, Ad and Al not evident. Tutorium present with tutorial cusp (
<figureCitation id="F2DC2A708E42C830671F55F4FD518888" box="[595,680,914,940]" captionStart="FIGURES 1 – 3" captionStartId="4.[151,271,1925,1949]" captionTargetBox="[207,1392,160,1911]" captionTargetId="figure@4.[197,1403,139,1911]" captionTargetPageId="4" captionText="FIGURES 1 – 3. Unduloribates dianae sp. nov., adult female: 1, dorsal aspect, dorsophragmata indicated by arrow; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed). Scale bars: 100 µm." httpUri="https://zenodo.org/record/191496/files/figure.png" pageId="2" pageNumber="49">Figs. 3</figureCitation>
,
<figureCitation id="F2DC2A708E42C83067FA55F4FD2E8888" box="[694,727,914,940]" captionStart="FIGURES 7 – 12" captionStartId="6.[151,271,1895,1919]" captionTargetBox="[151,1436,194,1871]" captionTargetId="figure@6.[151,1436,194,1871]" captionTargetPageId="6" captionText="FIGURES 7 – 12. Unduloribates dianae sp. nov., scanning electron micrographs; figures 7 - 10, 12 of adult females; 7, detail of bothridium and bothridial seta; 8, detail of blocky cerotegument on notogaster; 9, detail of prodorsum and anterior of notogaster; 10, prodorsum, lateral aspect; 11, egg in female hysterosoma; 12, tarsus I and anterior of tibia I. Scale bars: 7, 8, 11, 12, 10 µm; 9, 10, 20 µm." httpUri="https://zenodo.org/record/191498/files/figure.png" pageId="2" pageNumber="49">10</figureCitation>
). Pedotectum II covering part of acetabulum II. Discidium present between acetabula II and III, merging with circumpedal carina posteriorly and with short, tapered, tuberculate custodium anteriorly (
<figureCitation id="F2DC2A708E42C830677355B9FD6188DD" box="[575,664,991,1017]" captionStart="FIGURES 1 – 3" captionStartId="4.[151,271,1925,1949]" captionTargetBox="[207,1392,160,1911]" captionTargetId="figure@4.[197,1403,139,1911]" captionTargetPageId="4" captionText="FIGURES 1 – 3. Unduloribates dianae sp. nov., adult female: 1, dorsal aspect, dorsophragmata indicated by arrow; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed). Scale bars: 100 µm." httpUri="https://zenodo.org/record/191496/files/figure.png" pageId="2" pageNumber="49">Figs. 2</figureCitation>
,
<figureCitation id="F2DC2A708E42C83067EA55B9FD4088DD" box="[678,697,991,1017]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,272,867,891]" captionTargetBox="[151,1435,151,859]" captionTargetId="figure@5.[151,1436,139,897]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Unduloribates dianae sp. nov., scanning electron micrographs of adult females: 4, ventral aspect; 5, dorsal aspect; 6, notogaster with much of cerotegument removed. Scale bars: 100 µm." httpUri="https://zenodo.org/record/191497/files/figure.png" pageId="2" pageNumber="49">4</figureCitation>
). Notogaster longer than wide; anterior margin undulating, concave posterior to bothridia, and strongly convex lateral to bothridia (
<figureCitation id="F2DC2A708E42C83061405263FB988F3B" box="[1036,1121,1029,1055]" captionStart="FIGURES 1 – 3" captionStartId="4.[151,271,1925,1949]" captionTargetBox="[207,1392,160,1911]" captionTargetId="figure@4.[197,1403,139,1911]" captionTargetPageId="4" captionText="FIGURES 1 – 3. Unduloribates dianae sp. nov., adult female: 1, dorsal aspect, dorsophragmata indicated by arrow; 2, ventral aspect (legs removed); 3, prodorsum, lateral aspect (gnathosoma, notogaster and legs removed). Scale bars: 100 µm." httpUri="https://zenodo.org/record/191496/files/figure.png" pageId="2" pageNumber="49">Figs. 1</figureCitation>
,
<figureCitation id="F2DC2A708E42C830613E5263FB7D8F3B" box="[1138,1156,1029,1055]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,272,867,891]" captionTargetBox="[151,1435,151,859]" captionTargetId="figure@5.[151,1436,139,897]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Unduloribates dianae sp. nov., scanning electron micrographs of adult females: 4, ventral aspect; 5, dorsal aspect; 6, notogaster with much of cerotegument removed. Scale bars: 100 µm." httpUri="https://zenodo.org/record/191497/files/figure.png" pageId="2" pageNumber="49">6</figureCitation>
). Pteromorphs without hinge, tapered to point anteroventrally (
<figureCitation id="F2DC2A708E42C830673E524DFD448F61" box="[626,701,1067,1093]" captionStart="FIGURES 4 – 6" captionStartId="5.[151,272,867,891]" captionTargetBox="[151,1435,151,859]" captionTargetId="figure@5.[151,1436,139,897]" captionTargetPageId="5" captionText="FIGURES 4 – 6. Unduloribates dianae sp. nov., scanning electron micrographs of adult females: 4, ventral aspect; 5, dorsal aspect; 6, notogaster with much of cerotegument removed. Scale bars: 100 µm." httpUri="https://zenodo.org/record/191497/files/figure.png" pageId="2" pageNumber="49">Fig. 4</figureCitation>
). Notogastral setation 10 pairs; octotaxic system absent. Five pairs of lyrifissures present. Epimeral setation 3-1-2-3, 3-1-3-3 or 3-2-7-7; 6 to 10 pairs genital setae, 1 pair aggenital, 3 pairs of adanal and 2 pairs of anal setae. Postanal porose area absent. Trägårdh’s organ present. Palp with normal 5 segments and tarsal lyrifissure; setal formula: 0-2-1-3-9(1); palpal eupathidium
<emphasis id="5893EAE78E42C830602652F8FA658F93" box="[1386,1436,1182,1207]" italics="true" pageId="2" pageNumber="49">acm</emphasis>
attached to solenidion at least along distal third of length (
<figureCitation id="F2DC2A708E42C830660252A5FC5F8FF9" box="[846,934,1219,1245]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,272,1296,1320]" captionTargetBox="[150,1436,741,1284]" captionTargetId="figure@7.[145,1440,741,1286]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Unduloribates dianae sp. nov., scanning electron micrographs of adult females; 13, gnathosoma, ventral aspect; 14, gnathosoma, anterior aspect; 15, pedipalp with lateral detail of gena and rutellum. Scale bars: 20 µm." httpUri="https://zenodo.org/record/191499/files/figure.png" pageId="2" pageNumber="49">Fig. 15</figureCitation>
, arrow). Axillary saccule of subcapitulum present. Chelicera chelate-dentate with 2 slender, barbed setae (
<figureCitation id="F2DC2A708E42C8306631528CFC2D8E20" box="[893,980,1258,1284]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,272,1296,1320]" captionTargetBox="[150,1436,741,1284]" captionTargetId="figure@7.[145,1440,741,1286]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Unduloribates dianae sp. nov., scanning electron micrographs of adult females; 13, gnathosoma, ventral aspect; 14, gnathosoma, anterior aspect; 15, pedipalp with lateral detail of gena and rutellum. Scale bars: 20 µm." httpUri="https://zenodo.org/record/191499/files/figure.png" pageId="2" pageNumber="49">Fig. 14</figureCitation>
). Rutellum pantelobasic (
<figureCitation id="F2DC2A708E42C83061B3528CFA768E20" box="[1279,1423,1258,1284]" captionStart="FIGURES 13 – 15" captionStartId="7.[151,272,1296,1320]" captionTargetBox="[150,1436,741,1284]" captionTargetId="figure@7.[145,1440,741,1286]" captionTargetPageId="7" captionText="FIGURES 13 – 15. Unduloribates dianae sp. nov., scanning electron micrographs of adult females; 13, gnathosoma, ventral aspect; 14, gnathosoma, anterior aspect; 15, pedipalp with lateral detail of gena and rutellum. Scale bars: 20 µm." httpUri="https://zenodo.org/record/191499/files/figure.png" pageId="2" pageNumber="49">Figs. 13–15</figureCitation>
). Legs heterotridactylous; claws with fine denticles dorsally (
<figureCitation id="F2DC2A708E42C83066025376FC5F8E0E" box="[846,934,1296,1322]" captionStart="FIGURES 7 – 12" captionStartId="6.[151,271,1895,1919]" captionTargetBox="[151,1436,194,1871]" captionTargetId="figure@6.[151,1436,194,1871]" captionTargetPageId="6" captionText="FIGURES 7 – 12. Unduloribates dianae sp. nov., scanning electron micrographs; figures 7 - 10, 12 of adult females; 7, detail of bothridium and bothridial seta; 8, detail of blocky cerotegument on notogaster; 9, detail of prodorsum and anterior of notogaster; 10, prodorsum, lateral aspect; 11, egg in female hysterosoma; 12, tarsus I and anterior of tibia I. Scale bars: 7, 8, 11, 12, 10 µm; 9, 10, 20 µm." httpUri="https://zenodo.org/record/191498/files/figure.png" pageId="2" pageNumber="49">Fig. 12</figureCitation>
). Femora I to IV and trochanters III and IV with porose areas (
<figureCitation id="F2DC2A708E42C830643F5350FDD58E74" box="[371,556,1334,1360]" captionStart="FIGURES 16 – 19" captionStartId="7.[151,269,1945,1969]" captionText="FIGURES 16 – 19. Unduloribates dianae sp. nov., legs of adult female, all abaxial aspect: 16, leg I (trochanter removed); 17, leg II (trochanter and femur removed); 18, leg III; 19, leg IV. Scale bars: 50 µm." pageId="2" pageNumber="49">Figs. 16, 18, 19</figureCitation>
). Famulus
<emphasis id="5893EAE78E42C83067E25350FD368E6B" box="[686,719,1334,1359]" italics="true" pageId="2" pageNumber="49">(e)</emphasis>
rod-like, rounded distally (
<figureCitation id="F2DC2A708E42C83061415350FB9C8E74" box="[1037,1125,1334,1360]" captionStart="FIGURES 7 – 12" captionStartId="6.[151,271,1895,1919]" captionTargetBox="[151,1436,194,1871]" captionTargetId="figure@6.[151,1436,194,1871]" captionTargetPageId="6" captionText="FIGURES 7 – 12. Unduloribates dianae sp. nov., scanning electron micrographs; figures 7 - 10, 12 of adult females; 7, detail of bothridium and bothridial seta; 8, detail of blocky cerotegument on notogaster; 9, detail of prodorsum and anterior of notogaster; 10, prodorsum, lateral aspect; 11, egg in female hysterosoma; 12, tarsus I and anterior of tibia I. Scale bars: 7, 8, 11, 12, 10 µm; 9, 10, 20 µm." httpUri="https://zenodo.org/record/191498/files/figure.png" pageId="2" pageNumber="49">Fig. 12</figureCitation>
). Seta
<emphasis id="5893EAE78E42C83061FF5350FB3B8E6B" box="[1203,1218,1334,1359]" italics="true" pageId="2" pageNumber="49">d</emphasis>
absent from tibiae I to IV and genua I to III. Solenidion ω2 present on tarsus II. Solenidia on tibiae and genua short, except φ
<emphasis id="5893EAE78E42C8306038530AFA858E5E" box="[1396,1404,1388,1402]" italics="true" pageId="2" pageNumber="49">1</emphasis>
of tibia I flagellate. Tibia I without tubercle projecting over base of tarsus I.
</paragraph>
<paragraph id="6A5836F58E42C830658A53CDFC0C8DDD" blockId="2.[151,1437,609,1785]" pageId="2" pageNumber="49">
<emphasis id="5893EAE78E42C830658A53CDFEB28EE0" box="[198,331,1451,1476]" italics="true" pageId="2" pageNumber="49">Immatures.</emphasis>
Apheredermous, plicate, without hysterosomal macrosclerites or excentrosclerites. Line of dehiscence extends to seta
<emphasis id="5893EAE78E42C830649D53B7FE1E8ECE" box="[465,487,1489,1514]" italics="true" pageId="2" pageNumber="49">la</emphasis>
(
<figureCitation id="F2DC2A708E42C83064B453B7FDB78ECF" box="[504,590,1489,1515]" captionStart="FIGURES 20 – 21" captionStartId="11.[151,271,1968,1992]" captionTargetBox="[186,1402,156,1967]" captionTargetId="figure@11.[176,1420,143,1972]" captionTargetPageId="11" captionText="FIGURES 20 – 21. Unduloribates dianae sp. nov., immatures; 20, tritonymph, dorsal aspect, legs removed; 21, larva, dorsal aspect, legs removed. Scale bars: 50 µm." httpUri="https://zenodo.org/record/191500/files/figure.png" pageId="2" pageNumber="49">Fig. 20</figureCitation>
, arrow). Prodorsal porose regions present (
<figureCitation id="F2DC2A708E42C830610953B7FB648ECF" box="[1093,1181,1489,1515]" captionStart="FIGURES 20 – 21" captionStartId="11.[151,271,1968,1992]" captionTargetBox="[186,1402,156,1967]" captionTargetId="figure@11.[176,1420,143,1972]" captionTargetPageId="11" captionText="FIGURES 20 – 21. Unduloribates dianae sp. nov., immatures; 20, tritonymph, dorsal aspect, legs removed; 21, larva, dorsal aspect, legs removed. Scale bars: 50 µm." httpUri="https://zenodo.org/record/191500/files/figure.png" pageId="2" pageNumber="49">Fig. 20</figureCitation>
). Gastronotal setation unideficient; larva with 11 (
<emphasis id="5893EAE78E42C8306497539EFE0B8D31" box="[475,498,1528,1557]" italics="true" pageId="2" pageNumber="49">h3</emphasis>
absent), nymphs with 15 pairs of setae (
<emphasis id="5893EAE78E42C8306684539EFC218D31" box="[968,984,1528,1557]" italics="true" pageId="2" pageNumber="49">f1</emphasis>
, absent). Gastronotal setae of
<emphasis id="5893EAE78E42C8306076539EFABE8D35" box="[1338,1351,1528,1553]" italics="true" pageId="2" pageNumber="49">c</emphasis>
,
<emphasis id="5893EAE78E42C8306019539EFA9A8D35" box="[1365,1379,1528,1553]" italics="true" pageId="2" pageNumber="49">d</emphasis>
, and
<emphasis id="5893EAE78E42C83065DB5047FF668D1E" box="[151,159,1569,1594]" italics="true" pageId="2" pageNumber="49">l</emphasis>
series monomorphic in larva and nymphs (
<figureCitation id="F2DC2A708E42C83067DA5047FCDA8D1F" box="[662,803,1569,1595]" captionStart="FIGURES 20 – 21" captionStartId="11.[151,271,1968,1992]" captionTargetBox="[186,1402,156,1967]" captionTargetId="figure@11.[176,1420,143,1972]" captionTargetPageId="11" captionText="FIGURES 20 – 21. Unduloribates dianae sp. nov., immatures; 20, tritonymph, dorsal aspect, legs removed; 21, larva, dorsal aspect, legs removed. Scale bars: 50 µm." httpUri="https://zenodo.org/record/191500/files/figure.png" pageId="2" pageNumber="49">Figs. 20, 21</figureCitation>
). Opisthonotal gland present in all instars. Coxisternal porose regions present in all nymphs. Porose regions surrounding opening of opisthonotal gland, positioned ventrolaterally, in all nymphs (
<figureCitation id="F2DC2A708E42C83064B3500BFDA18DA3" box="[511,600,1645,1671]" captionStart="FIGURES 20 – 21" captionStartId="11.[151,271,1968,1992]" captionTargetBox="[186,1402,156,1967]" captionTargetId="figure@11.[176,1420,143,1972]" captionTargetPageId="11" captionText="FIGURES 20 – 21. Unduloribates dianae sp. nov., immatures; 20, tritonymph, dorsal aspect, legs removed; 21, larva, dorsal aspect, legs removed. Scale bars: 50 µm." httpUri="https://zenodo.org/record/191500/files/figure.png" pageId="2" pageNumber="49">Fig. 20</figureCitation>
). Apodemato-acetabular tracheal system or porose homologues absent. Paraprocts atrichous in larva, protonymph and deutonymph. Cupule development normal. Bothridium, bothridial seta and seta
<emphasis id="5893EAE78E42C83064E550DFFE468DF6" box="[425,447,1721,1746]" italics="true" pageId="2" pageNumber="49">in</emphasis>
fully formed in all immatures. Bothridium cup-shaped. Seta
<emphasis id="5893EAE78E42C83061CA50DFFB6C8DF6" box="[1158,1173,1721,1746]" italics="true" pageId="2" pageNumber="49">d</emphasis>
absent from tibiae I to IV and genua I to III. Setation of protonymphal leg IV normal: 0-0-0-0-7.
</paragraph>
</subSubSection>
<subSubSection id="22FD657E8E42C83165DB514AFCF98A57" lastPageId="3" lastPageNumber="50" pageId="2" pageNumber="49" type="discussion">
<paragraph id="6A5836F58E42C83065DB514AFF078C61" blockId="2.[151,1436,1836,2014]" box="[151,254,1836,1861]" pageId="2" pageNumber="49">
<emphasis id="5893EAE78E42C83065DB514AFF078C61" box="[151,254,1836,1861]" italics="true" pageId="2" pageNumber="49">Remarks</emphasis>
</paragraph>
<paragraph id="6A5836F58E42C830658A5134FBBF8CFA" blockId="2.[151,1436,1836,2014]" pageId="2" pageNumber="49">
<bibRefCitation id="0E764B048E42C830658A5134FEA98C48" author="Piffl" box="[198,336,1874,1900]" pageId="2" pageNumber="61" refString="Piffl, E. (1972). Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
suggested that the rostral incision found in
<taxonomicName id="ADE74D768E42C83066155134FBFB8C4F" box="[857,1026,1874,1899]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="5893EAE78E42C83066155134FBFB8C4F" box="[857,1026,1874,1899]" italics="true" pageId="2" pageNumber="49">Unduloribates</emphasis>
</taxonomicName>
is distinct from the genal notch of many poronotic brachypyline taxa because it is an incision of the rostral tectum. Although the genal notch as defined by
<bibRefCitation id="0E764B048E42C830645751F8FD8D8C9C" author="Grandjean" box="[283,628,1950,1976]" pageId="2" pageNumber="61" refString="Grandjean, F. (1952) Au sujet de l'ectosquelette du podosoma chez les Oribates superieurs et de sa terminologie. Bulletin de la Societe Zoologique de France, 77, 13 - 36." type="journal article" year="1952">Grandjean (1952, his fig. 2C)</bibRefCitation>
is an incision of the rostral tectum, a genal notch is oblique, whereas the incision of
<taxonomicName id="ADE74D768E42C830640A51A2FE168CF9" box="[326,495,1988,2013]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="5893EAE78E42C830640A51A2FE168CF9" box="[326,495,1988,2013]" italics="true" pageId="2" pageNumber="49">Unduloribates</emphasis>
</taxonomicName>
is vertical; like
<bibRefCitation id="0E764B048E42C83067E051A2FCCD8CFA" author="Piffl" box="[684,820,1988,2014]" pageId="2" pageNumber="61" refString="Piffl, E. (1972). Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
we consider it distinct.
</paragraph>
<paragraph id="6A5836F58E43C831658A56FEFCF98A57" blockId="3.[151,1437,152,375]" pageId="3" pageNumber="50">
<bibRefCitation id="0E764B048E43C831658A56FEFEAD8B96" author="Piffl" box="[198,340,152,178]" pageId="3" pageNumber="61" refString="Piffl, E. (1972). Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
did not illustrate or describe humerosejugal porose areas Am and Ah for
<emphasis id="5893EAE78E43C83161BF56FEFF048BFC" italics="true" pageId="3" pageNumber="50">
<taxonomicName id="ADE74D768E43C83161BF56FEFF018BFC" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="medusa">Unduloribates medusa</taxonomicName>
,
</emphasis>
and they have not been described for other species. However, these porose areas are visible in dissected lateral mounts or in young, weakly sclerotized adults of the species described below, and could have been easily overlooked in other species.
<bibRefCitation id="0E764B048E43C83167CE576AFCE98A02" author="Piffl" box="[642,784,268,294]" pageId="3" pageNumber="61" refString="Piffl, E. (1972). Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
beautifully illustrated components of the gnathosoma including the axillary saccule at the base of the palps.
<bibRefCitation id="0E764B048E43C831665C5754FC608A68" author="Piffl" box="[784,921,306,332]" pageId="3" pageNumber="61" refString="Piffl, E. (1972). Zur Systematik der Oribatiden (Acari) (Neue Oribatiden aus Nepal, Costa Rica und Brasilien ergeben eine neue Familie der Unduloribatidae und erweitern die Polypterozetidae um die Gattungen Podopterotegaeus, Nodocepheus, Eremaeozetes und Tumerozetes). Khumbu Himal, 4, 269 - 314." type="journal article" year="1972">Piffl (1972)</bibRefCitation>
also illustrated the dorsal and ventral of the larva of
<taxonomicName id="ADE74D768E43C83165B4573FFE8C8A56" box="[248,373,345,370]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="medusa">
<emphasis id="5893EAE78E43C83165B4573FFE8C8A56" box="[248,373,345,370]" italics="true" pageId="3" pageNumber="50">U. medusa</emphasis>
</taxonomicName>
, and noted the absence of seta
<emphasis id="5893EAE78E43C83167AE573FFD008A53" box="[738,761,345,375]" italics="true" pageId="3" pageNumber="50">h3</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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