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<document id="38D6AE858D6705AA034F0601BCFFAE31" ID-DOI="10.5281/zenodo.5351908" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351908" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630207225511" checkinUser="tatiana" docAuthor="Polgar, Gianluca, Jaafar, Zeehan &amp; Konstantinidis, Peter" docDate="2013" docId="C35E87E72D472E4AB6E4FD08ABE2F83C" docLanguage="en" docName="61rbz311-321.pdf" docOrigin="Raffles Bulletin of Zoology 61 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Boleophthalmus poti Polgar &amp; Jaafar &amp; Konstantinidis 2013, new species" docType="treatment" docUuid="A7F636FD-495E-4AF2-A3DB-4D1416247748" docUuidSource="ZooBank" docVersion="5" lastPageNumber="316" masterDocId="3F67FF9F2D452E4FB521FFF4AB13FFEE" masterDocTitle="A New Species Of Mudskipper, Boleophthalmus Poti (Teleostei: Gobiidae: Oxudercinae) From The Gulf Of Papua, Papua New Guinea, And A Key To The Genus" masterLastPageNumber="321" masterPageNumber="311" pageNumber="313" updateTime="1699185193955" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0D864ABF10C2A48B9FDD31274957D707" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B51E406B36CF8ECE1931A72BD523CBCB">
<mods:title id="468F29B0D4A0C9A7B4F023E1A322C926">A New Species Of Mudskipper, Boleophthalmus Poti (Teleostei: Gobiidae: Oxudercinae) From The Gulf Of Papua, Papua New Guinea, And A Key To The Genus</mods:title>
</mods:titleInfo>
<mods:name id="B321C0027F3669270E9741BB8B4DE068" type="personal">
<mods:role id="9672A1F2447D5AAFC8F353C263DB79E7">
<mods:roleTerm id="7F07C94FBA80A3CC92BB18A669EA19AF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8066219C4C297F7E28877DDBFC173C27">Polgar, Gianluca</mods:namePart>
</mods:name>
<mods:name id="BEAAC855B51E2D98C9B3E155E7365896" type="personal">
<mods:role id="E8CE7CBA85893E53EF0EC0B6A08C31AC">
<mods:roleTerm id="4429090215CEA23F536E0D000C34BE65">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00D0FB3E897FCD2687876A16D99D1FF3">Jaafar, Zeehan</mods:namePart>
</mods:name>
<mods:name id="0BE5D848600CC8985B9779AF134F4B92" type="personal">
<mods:role id="79F90F7D9911292562C4F1763A504958">
<mods:roleTerm id="F1115A38A12A778DB552A47F28EB2B7E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EF6DE2A13BEC556DFEFD85303B22BA8">Konstantinidis, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource id="E5D55E23DD22D2AA9FD1362DB2E8AE1E">text</mods:typeOfResource>
<mods:relatedItem id="AE6300D870076158A4646937B9C14518" type="host">
<mods:titleInfo id="BC05B9BC90180F5810B402DEC41E3079">
<mods:title id="54AC2C01AB2E594ADFE38E84EEB7409C">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="CB6EF442D99533AFA146D632A0C9751A">
<mods:date id="E204A18D652BFFDD3A0F3A09C9D42AA6">2013</mods:date>
<mods:detail id="02B84AA826737FFC7B86A49978EA2D5E" type="pubDate">
<mods:number id="B96E98E31DCED37F664816590E9D8F44">2013-02-28</mods:number>
</mods:detail>
<mods:detail id="2FD455C86E7B73E0E40AFAEBBD9EB4A5" type="volume">
<mods:number id="35DA0578481B9FAE9AD47AE4A984A2AB">61</mods:number>
</mods:detail>
<mods:detail id="8D09DB935ABDA0153DB4C544B6633E6A" type="issue">
<mods:number id="C25964E528D8DB77685EDBD70712D518">1</mods:number>
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<mods:start id="A25F7D81B553D60AAEC82EBCBA797322">311</mods:start>
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<mods:classification id="9723367FE753827FAC56FCE438A5336D">journal article</mods:classification>
<mods:identifier id="8ACC0E8FC314D98F89CF4A1FE5B353E7" type="DOI">10.5281/zenodo.5351908</mods:identifier>
<mods:identifier id="CF06F28D3B43F3B964231C91ACB6D2BA" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F233B75DBD186A468ED80A11C3FB8F99" type="Zenodo-Dep">5351908</mods:identifier>
<mods:identifier id="CC1A15FE4D95D96DD8F449E12AF71F61" type="ZooBank">0912AFAC-C74A-4D4D-BCA7-FDFA42380278</mods:identifier>
</mods:mods>
<treatment id="C35E87E72D472E4AB6E4FD08ABE2F83C" LSID="urn:lsid:zoobank.org:act:A7F636FD-495E-4AF2-A3DB-4D1416247748" httpUri="http://treatment.plazi.org/id/C35E87E72D472E4AB6E4FD08ABE2F83C" lastPageId="5" lastPageNumber="316" pageId="2" pageNumber="313">
<subSubSection id="03ED657A2D472E4DB6E4FD08AE52FCFB" box="[965,1345,764,789]" pageId="2" pageNumber="313" type="nomenclature">
<paragraph id="4B4836F12D472E4DB6E4FD08AE52FCFB" blockId="2.[965,1345,764,821]" box="[965,1345,764,789]" pageId="2" pageNumber="313">
<heading id="1000819D2D472E4DB6E4FD08AE52FCFB" bold="true" box="[965,1345,764,789]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="313" reason="2">
<emphasis id="7983EAE32D472E4DB6E4FD08AE52FCFB" bold="true" box="[965,1345,764,789]" pageId="2" pageNumber="313">
<taxonomicName id="8CF74D722D472E4DB6E4FD08AFB8FCFB" ID-CoL="5WN5Z" authority="Polgar &amp; Jaafar &amp; Konstantinidis, 2013" authorityName="Polgar &amp; Jaafar &amp; Konstantinidis" authorityYear="2013" box="[965,1195,764,789]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="313" phylum="Chordata" rank="species" species="poti" status="sp. nov.">
<emphasis id="7983EAE32D472E4DB6E4FD08AFB8FCFB" bold="true" box="[965,1195,764,789]" italics="true" pageId="2" pageNumber="313">Boleophthalmus poti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="62B057982D472E4DB19AFD09AE52FCFB" box="[1211,1345,765,789]" pageId="2" pageNumber="313" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D472E4DB6D5FCE9AE02FCDB" box="[1012,1297,797,821]" pageId="2" pageNumber="313" type="description">
<paragraph id="4B4836F12D472E4DB6D5FCE9AE02FCDB" blockId="2.[965,1345,764,821]" box="[1012,1297,797,821]" pageId="2" pageNumber="313">
(
<figureCitation id="D3CC2A742D472E4DB6DDFCE9AF5AFCDB" box="[1020,1097,797,821]" captionStart="Fig" captionStartId="2.[835,869,2002,2024]" captionTargetBox="[835,1474,1286,1979]" captionTargetId="figure-716@2.[835,1476,1285,1979]" captionTargetPageId="2" captionText="Fig.2. Boleophthalmus poti:A, holotype, MSNG 56891; B, paratype, preserved specimen, female (108.2 mm) (BMNH 2011.1.27.1); C, an individual in its natural habitat, on the mud banks of Wapi Creek, Purutu Island, Fly River delta. Scale bars =10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351912" httpUri="https://zenodo.org/record/5351912/files/figure.png" pageId="2" pageNumber="313">Figs. 2</figureCitation>
,
<figureCitation id="D3CC2A742D472E4DB176FCE9AF75FCDB" box="[1111,1126,797,821]" captionStart="Fig" captionStartId="3.[113,148,1916,1938]" captionTargetBox="[106,752,1529,1891]" captionTargetId="figure-883@3.[91,755,1516,1897]" captionTargetPageId="3" captionText="Fig. 3. Boleophthalmus poti, BMNH 2011.1.27.1. Lateral view of the suspensorium. Abbreviations: ap = autopalatine; d = dentary; ep = ectopterygoid; hm = hyomandibular; io = interopercle; m = maxilla; mp = metapterygoid; op = opercle; pm = premaxilla; po = preopercle; pp = ascending processes of the premaxilla;q =quadrate; sop = subopercle; sp = symplectic; st = symphyseal tooth. Scale bar = 1 mm. the original image was reversed horizontally." figureDoi="http://doi.org/10.5281/zenodo.5351914" httpUri="https://zenodo.org/record/5351914/files/figure.png" pageId="2" pageNumber="313">3</figureCitation>
,
<figureCitation id="D3CC2A742D472E4DB155FCE9AF90FCDB" box="[1140,1155,797,821]" captionStart="Fig" captionStartId="5.[814,853,1868,1890]" captionTargetBox="[813,1454,986,1844]" captionTargetId="figure-437@5.[813,1456,986,1844]" captionTargetPageId="5" captionText="Fig. 5. Shape and colouration of freshly dead species of Boleophthalmus with un-notched dentary teeth: A, female of B. birdsongi (95 mm; ZRC 52250); B, female of B. caeruleomaculatus (106 mm; MSNG 54689); and C, paratype, male of B. poti (83.4 mm; USNM 40566). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351918" httpUri="https://zenodo.org/record/5351918/files/figure.png" pageId="2" pageNumber="313">5</figureCitation>
; Tables 1, 2)
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D472E4CB662FCAFAA97FEE3" lastPageId="3" lastPageNumber="314" pageId="2" pageNumber="313" type="materials_examined">
<paragraph id="4B4836F12D472E4DB662FCAFAF7FFC29" blockId="2.[835,1474,859,968]" pageId="2" pageNumber="313">
<emphasis id="7983EAE32D472E4DB662FCAFAF17FC9F" bold="true" box="[835,1028,859,881]" pageId="2" pageNumber="313">
<emphasis id="7983EAE32D472E4DB662FCAFA8EDFC9F" bold="true" box="[835,1022,859,881]" italics="true" pageId="2" pageNumber="313">Material examined</emphasis>
.
</emphasis>
—
<materialsCitation id="FB9F3CAC2D472E4DB109FCA8AF7FFC29" collectingDate="2007-09-24" collectionCode="MSNG" collectorName="G. Polgar &amp; Polgar, A &amp; C. Tenakenai" country="Papua New Guinea" latitude="-8.429361" location="Sisikura Island" longLatPrecision="1" longitude="143.60667" municipality="Fly river" pageId="2" pageNumber="313" specimenCode="MSNG 56891" specimenCount="1" stateProvince="Western" typeStatus="holotype">
<typeStatus id="944C88532D472E4DB109FCA8AF96FC9C" box="[1064,1157,860,882]" pageId="2" pageNumber="313" type="holotype">Holotype</typeStatus>
:
<specimenCode id="1B519E8A2D472E4DB1ACFCA8AE05FC9C" box="[1165,1302,860,882]" collectionCode="MSNG" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="2" pageNumber="313">MSNG 56891</specimenCode>
, female, 89.9 mm SL;
<collectingCountry id="33E076612D472E4DB64AFC8DAF35FC61" box="[875,1062,889,911]" name="Papua New Guinea" pageId="2" pageNumber="313">Papua New Guinea</collectingCountry>
,
<collectingRegion id="8933F8132D472E4DB10FFC8DAFCAFC61" box="[1070,1241,889,911]" country="Papua New Guinea" name="Western" pageId="2" pageNumber="313">Western Province</collectingRegion>
,
<collectingMunicipality id="AB2CAC8B2D472E4DB1C3FC8DAE27FC61" box="[1250,1332,889,911]" pageId="2" pageNumber="313">Fly river</collectingMunicipality>
delta,
<location id="4E28602A2D472E4DB052FC8DA893FC45" LSID="urn:lsid:plazi:treatment:C35E87E72D472E4AB6E4FD08ABE2F83C:4E28602A2D472E4DB052FC8DA893FC45" country="Papua New Guinea" latitude="-8.429361" longLatPrecision="1" longitude="143.60667" municipality="Fly river" name="Sisikura Island" pageId="2" pageNumber="313" stateProvince="Western">Sisikura Island</location>
,
<geoCoordinate id="2EC350362D472E4DB6ACFC61AF11FC45" box="[909,1026,917,939]" degrees="8" direction="south" minutes="25" orientation="latitude" pageId="2" pageNumber="313" precision="1" seconds="45.7" value="-8.429361">8°25'45.7&quot;S</geoCoordinate>
,
<geoCoordinate id="2EC350362D472E4DB12EFC61AF8DFC45" box="[1039,1182,917,939]" degrees="143" direction="east" minutes="36" orientation="longitude" pageId="2" pageNumber="313" precision="1" seconds="24.0" value="143.60667">143°36'24.0&quot;E</geoCoordinate>
, coll.
<collectorName id="E60253272D472E4DB1FFFC62AE2DFC45" box="[1246,1342,917,939]" pageId="2" pageNumber="313">G. Polgar</collectorName>
, A. Sacchetti &amp;
<collectorName id="E60253272D472E4DB67FFC46A8F3FC26" box="[862,992,946,968]" pageId="2" pageNumber="313">C. Tenakenai</collectorName>
,
<date id="3F4910312D472E4DB6CAFC46AF7BFC29" box="[1003,1128,946,967]" pageId="2" pageNumber="313" value="2007-09-24">
<collectingDate id="2F0DE9D92D472E4DB6CAFC46AF7BFC29" box="[1003,1128,946,967]" pageId="2" pageNumber="313" value="2007-09-24">24 Sep.2007</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="4B4836F12D472E4DB662FC18A8B7FB25" blockId="2.[835,1475,1004,1227]" pageId="2" pageNumber="313">
<materialsCitation id="FB9F3CAC2D472E4DB662FC18A8C3FBF0" collectionCode="BMNH" pageId="2" pageNumber="313" specimenCode="BMNH 2011.1" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="944C88532D472E4DB662FC18A8B5FBEF" box="[835,934,1004,1025]" pageId="2" pageNumber="313" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="1B519E8A2D472E4DB692FC18AF5FFBEF" box="[947,1100,1004,1025]" collectionCode="BMNH" pageId="2" pageNumber="313">BMNH 2011.1</specimenCode>
.27.1,
<specimenCount id="5DF1FD782D472E4DB1A9FC18AFCCFBEC" box="[1160,1247,1004,1026]" pageId="2" pageNumber="313" type="female">1 female</specimenCount>
(cleared and stained), 108.2 mm SL
</materialsCitation>
;
<materialsCitation id="FB9F3CAC2D472E4DB6FCFBFDA89DFBD5" collectionCode="MSNG" pageId="2" pageNumber="313" specimenCode="MSNG 56892" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCode id="1B519E8A2D472E4DB6FCFBFDAF78FBF1" box="[989,1131,1033,1055]" collectionCode="MSNG" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="2" pageNumber="313">MSNG 56892</specimenCode>
,
<specimenCount id="5DF1FD782D472E4DB156FBFDAFA8FBF1" box="[1143,1211,1033,1055]" pageId="2" pageNumber="313" type="male">1 male</specimenCount>
and
<specimenCount id="5DF1FD782D472E4DB1D1FBFDAE5AFBF1" box="[1264,1353,1033,1055]" pageId="2" pageNumber="313" type="female">1 female</specimenCount>
, 89.6–104.5 mm SL
</materialsCitation>
;
<materialsCitation id="FB9F3CAC2D472E4DB6B6FBD2AE5BFBD5" box="[919,1352,1061,1083]" collectionCode="USNM" pageId="2" pageNumber="313" specimenCode="USNM 405556" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCode id="1B519E8A2D472E4DB6B6FBD2AF3FFBD5" box="[919,1068,1061,1083]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="313" type="Museum">USNM 405556</specimenCode>
,
<specimenCount id="5DF1FD782D472E4DB117FBD2AF91FBD5" box="[1078,1154,1061,1083]" pageId="2" pageNumber="313" type="male">3 males</specimenCount>
, 80.5–105.8 mm SL
</materialsCitation>
;
<materialsCitation id="FB9F3CAC2D472E4DB073FBD2AEACFBB6" accessionNumber="ZRC 52240" collectingDate="2007-09-24" collectionCode="ZRC" collectorName="G. Polgar &amp; Polgar, A &amp; C. Tenakenai" country="Papua New Guinea" latitude="-8.429361" location="Sisikura Island" longLatPrecision="1" longitude="143.60667" municipality="Fly river" pageId="2" pageNumber="313" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Western" typeStatus="paratype">
<collectionCode id="2DE6AE342D472E4DB073FBD2AE92FBD5" box="[1362,1409,1062,1083]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="313">ZRC</collectionCode>
52240
<specimenCount id="5DF1FD782D472E4DB662FBB6A89CFBB6" box="[835,911,1090,1112]" pageId="2" pageNumber="313" type="male">2 males</specimenCount>
and
<specimenCount id="5DF1FD782D472E4DB6E0FBB6AF0AFBB6" box="[961,1049,1090,1112]" pageId="2" pageNumber="313" type="female">1 female</specimenCount>
, 78.0–99.4 mm SL; same data as holotype
</materialsCitation>
.
<materialsCitation id="FB9F3CAC2D472E4DB662FBABA8B2FB25" collectingDate="2007-09-23" collectionCode="MSNG" collectorName="G. Polgar &amp; A. Sacchetti &amp; C. Tenakenai" country="Papua New Guinea" county="Fly river" latitude="-8.389167" location="Wapi Creek" longLatPrecision="1" longitude="143.52316" municipality="Purutu Island" pageId="2" pageNumber="313" specimenCode="MSNG 56893" specimenCount="1" specimenCount-male="1" stateProvince="Western" typeStatus="paratype">
<specimenCode id="1B519E8A2D472E4DB662FBABA8DEFB9B" box="[835,973,1119,1141]" collectionCode="MSNG" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="2" pageNumber="313">MSNG 56893</specimenCode>
,
<specimenCount id="5DF1FD782D472E4DB6F9FBABAF0FFB9B" box="[984,1052,1119,1141]" pageId="2" pageNumber="313" type="male">1 male</specimenCount>
, 99.5 mm SL;
<collectingCountry id="33E076612D472E4DB18CFB94AE79FB9B" box="[1197,1386,1119,1141]" name="Papua New Guinea" pageId="2" pageNumber="313">Papua New Guinea</collectingCountry>
,
<collectingRegion id="8933F8132D472E4DB055FB94A888FB7C" country="Papua New Guinea" name="Western" pageId="2" pageNumber="313">Western Province</collectingRegion>
,
<collectingCounty id="A2294E7D2D472E4DB684FB88A8EAFB7C" box="[933,1017,1148,1170]" pageId="2" pageNumber="313">Fly river</collectingCounty>
delta,
<collectingMunicipality id="AB2CAC8B2D472E4DB11DFB88AFD3FB7C" box="[1084,1216,1148,1170]" pageId="2" pageNumber="313">Purutu Island</collectingMunicipality>
,
<location id="4E28602A2D472E4DB1EBFB88AE2CFB7C" LSID="urn:lsid:plazi:treatment:C35E87E72D472E4AB6E4FD08ABE2F83C:4E28602A2D472E4DB1EBFB88AE2CFB7C" box="[1226,1343,1148,1170]" country="Papua New Guinea" county="Fly river" latitude="-8.389167" longLatPrecision="1" longitude="143.52316" municipality="Purutu Island" name="Wapi Creek" pageId="2" pageNumber="313" stateProvince="Western">Wapi Creek</location>
,
<geoCoordinate id="2EC350362D472E4DB068FB88AEADFB7F" box="[1353,1470,1148,1169]" degrees="8" direction="south" minutes="23" orientation="latitude" pageId="2" pageNumber="313" precision="1" seconds="21.0" value="-8.389167">8°23'21.0&quot;S</geoCoordinate>
,
<geoCoordinate id="2EC350362D472E4DB662FB6DA8C1FB40" box="[835,978,1177,1198]" degrees="143" direction="east" minutes="31" orientation="longitude" pageId="2" pageNumber="313" precision="1" seconds="23.4" value="143.52316">143°31'23.4&quot;E</geoCoordinate>
, coll.
<collectorName id="E60253272D472E4DB133FB6DAF61FB41" box="[1042,1138,1177,1199]" pageId="2" pageNumber="313">G. Polgar</collectorName>
,
<collectorName id="E60253272D472E4DB15CFB6DAFE5FB41" box="[1149,1270,1177,1199]" pageId="2" pageNumber="313">A. Sacchetti</collectorName>
&amp;
<collectorName id="E60253272D472E4DB03BFB6DAE8DFB41" box="[1306,1438,1177,1199]" pageId="2" pageNumber="313">C. Tenakenai</collectorName>
,
<date id="3F4910312D472E4DB08BFB6DA8B2FB25" pageId="2" pageNumber="313" value="2007-09-23">
<collectingDate id="2F0DE9D92D472E4DB08BFB6DA8B2FB25" pageId="2" pageNumber="313" value="2007-09-23">23 Sep.2007</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="1F8866792D472E4DB662F826AE29F7D0" ID-DOI="http://doi.org/10.5281/zenodo.5351912" ID-Zenodo-Dep="5351912" httpUri="https://zenodo.org/record/5351912/files/figure.png" pageId="2" pageNumber="313" startId="2.[835,869,2002,2024]" targetBox="[835,1474,1286,1979]" targetPageId="2">
<paragraph id="4B4836F12D472E4DB662F826AE29F7D0" blockId="2.[835,1476,2002,2110]" pageId="2" pageNumber="313">
Fig. 2.
<taxonomicName id="8CF74D722D472E4DB6A3F827AF50F809" authorityName="Polgar &amp; Jaafar &amp; Konstantinidis" authorityYear="2013" box="[898,1091,2003,2023]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="313" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D472E4DB6A3F827AF50F809" box="[898,1091,2003,2023]" italics="true" pageId="2" pageNumber="313">Boleophthalmus poti</emphasis>
</taxonomicName>
: A, holotype, MSNG 56891; B, paratype, preserved specimen, female (108.2 mm) (BMNH 2011.1.27.1); C, an individual in its natural habitat, on the mud banks of Wapi Creek, Purutu Island, Fly River delta. Scale bars =10 mm.
</paragraph>
</caption>
<paragraph id="4B4836F12D462E4CB550FF4AAA97FEE3" blockId="3.[113,752,189,269]" pageId="3" pageNumber="314">
<materialsCitation id="FB9F3CAC2D462E4CB550FF4AA916FF01" accessionNumber="ZRC 51367" collectionCode="ZRC" country="Papua New Guinea" location="Purutu Island" municipality="Fly River" pageId="3" pageNumber="314" specimenCount="2">
Non-type material:
<collectionCode id="2DE6AE342D462E4CB410FF49AA73FF3C" box="[305,352,189,210]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="314">ZRC</collectionCode>
51367,
<specimenCount id="5DF1FD782D462E4CB493FF49AAF6FF3D" box="[434,485,189,211]" pageId="3" pageNumber="314" type="generic">2 ex.</specimenCount>
,
<quantity id="8C0F9B142D462E4CB4D1FF49A94FFF3D" box="[496,604,189,211]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="3" pageNumber="314" unit="mm" value="42.5" valueMax="50.0" valueMin="35.0">35–50 mm</quantity>
,
<location id="4E28602A2D462E4CB746FF4AA9FFFF3D" LSID="urn:lsid:plazi:treatment:C35E87E72D472E4AB6E4FD08ABE2F83C:4E28602A2D462E4CB746FF4AA9FFFF3D" box="[615,748,189,211]" country="Papua New Guinea" municipality="Fly River" name="Purutu Island" pageId="3" pageNumber="314">Purutu Island</location>
,
<collectingMunicipality id="AB2CAC8B2D462E4CB550FF2EABDFFF1E" box="[113,204,218,240]" pageId="3" pageNumber="314">Fly River</collectingMunicipality>
delta,
<collectingCountry id="33E076612D462E4CB42CFF2FAADAFF1E" box="[269,457,218,240]" name="Papua New Guinea" pageId="3" pageNumber="314">Papua New Guinea</collectingCountry>
, 2007
</materialsCitation>
;
<materialsCitation id="FB9F3CAC2D462E4CB72EFF2EAA93FEE3" accessionNumber="ZRC 52249" collectingDate="2007-09-24" collectionCode="ZRC" collectorName="G. Polgar &amp; Polgar, A &amp; C. Tenakenai" country="Papua New Guinea" latitude="-8.429361" location="Sisikura Island" longLatPrecision="1" longitude="143.60667" municipality="Fly river" pageId="3" pageNumber="314" specimenCount="1" stateProvince="Western">
<collectionCode id="2DE6AE342D462E4CB72EFF2EA92DFF01" box="[527,574,218,239]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="314">ZRC</collectionCode>
52249,
<specimenCount id="5DF1FD782D462E4CB7ADFF2EA9AEFF1E" box="[652,701,218,240]" pageId="3" pageNumber="314" type="generic">1 ex.</specimenCount>
, 80.4 mm, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D462E4CB550FEC1ABE7FCE3" pageId="3" pageNumber="314" type="diagnosis">
<paragraph id="4B4836F12D462E4CB550FEC1ABE7FCE3" blockId="3.[113,753,309,781]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB550FEC1ABF5FEA3" bold="true" box="[113,230,309,333]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB550FEC1ABCCFEA3" bold="true" box="[113,223,309,333]" italics="true" pageId="3" pageNumber="314">Diagnosis</emphasis>
.
</emphasis>
—
<taxonomicName id="8CF74D722D462E4CB42AFEC2AAF4FEA3" authorityName="Polgar &amp; Jaafar &amp; Konstantinidis" authorityYear="2013" box="[267,487,310,333]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D462E4CB42AFEC2AAF4FEA3" box="[267,487,310,333]" italics="true" pageId="3" pageNumber="314">Boleophthalmus poti</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="8CF74D722D462E4CB74FFEC2A9E3FEA3" authorityName="Murdy" authorityYear="1989" box="[622,752,310,333]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="birdsongi">
<emphasis id="7983EAE32D462E4CB74FFEC2A9E3FEA3" box="[622,752,310,333]" italics="true" pageId="3" pageNumber="314">B. birdsongi</emphasis>
</taxonomicName>
and
<taxonomicName id="8CF74D722D462E4CB583FEA2AA9CFE83" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[162,399,342,365]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D462E4CB583FEA2AA9CFE83" box="[162,399,342,365]" italics="true" pageId="3" pageNumber="314">B. caeruleomaculatus</emphasis>
</taxonomicName>
, sharing with these species unnotched, flattened and horizontally disposed dentary teeth. It is distinguished from these species (and all remaining congeners) by having elongate D1 fin spines, protruding from the fin margin, in both sexes and the following unique features of dorsal-fin colouration in live individuals: D1 with clearly demarcated sky-blue to whitish marginal band; interradial membranes of D1 greenish-brown, spotted with whitish to yellowish speckles, sometimes fused into broken, narrow and wavy lines; interradial membranes of D2 grey to pale brown, darker posteriorly, pale grey distally, with a very thin white margin and columns of 0–5 whitish spots between adjacent rays; D2 rays darker than interradial membranes.
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D462E4CB550FCC1AFA2FAFB" pageId="3" pageNumber="314" type="description">
<paragraph id="4B4836F12D462E4CB550FCC1AE40FED8" blockId="3.[113,753,821,1453]" lastBlockId="3.[812,1453,189,310]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB550FCC1ABECFCA3" bold="true" box="[113,255,821,845]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB550FCC1ABEBFCA3" bold="true" box="[113,248,821,845]" italics="true" pageId="3" pageNumber="314">Description</emphasis>
.
</emphasis>
— General body shape as in
<figureCitation id="D3CC2A742D462E4CB747FCC1A9AAFCA3" box="[614,697,821,845]" captionStart="Fig" captionStartId="2.[835,869,2002,2024]" captionTargetBox="[835,1474,1286,1979]" captionTargetId="figure-716@2.[835,1476,1285,1979]" captionTargetPageId="2" captionText="Fig.2. Boleophthalmus poti:A, holotype, MSNG 56891; B, paratype, preserved specimen, female (108.2 mm) (BMNH 2011.1.27.1); C, an individual in its natural habitat, on the mud banks of Wapi Creek, Purutu Island, Fly River delta. Scale bars =10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351912" httpUri="https://zenodo.org/record/5351912/files/figure.png" pageId="3" pageNumber="314">Figs. 2</figureCitation>
,
<figureCitation id="D3CC2A742D462E4CB7E8FCC1A9FFFCA3" box="[713,748,821,845]" captionStart="Fig" captionStartId="5.[814,853,1868,1890]" captionTargetBox="[813,1454,986,1844]" captionTargetId="figure-437@5.[813,1456,986,1844]" captionTargetPageId="5" captionText="Fig. 5. Shape and colouration of freshly dead species of Boleophthalmus with un-notched dentary teeth: A, female of B. birdsongi (95 mm; ZRC 52250); B, female of B. caeruleomaculatus (106 mm; MSNG 54689); and C, paratype, male of B. poti (83.4 mm; USNM 40566). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351918" httpUri="https://zenodo.org/record/5351918/files/figure.png" pageId="3" pageNumber="314">5C</figureCitation>
. Meristic and morphometric data are presented in Tables 1, 2. Head subcylindrical, about as wide as deep; body subcylindrical in thoracic region, becoming slightly compressed caudally; mouth subterminal, with jaw extending posteriorly to vertical point immediately posterior to posterior margin of eye. Lower jaw containing 2–8 caninoid teeth and 14–37 distally flattened, anteriorly directed and horizontally displaced teeth on each side. One recurved, canine tooth on both sides of mandibular symphysis, internal to the anterior margin of mandible (
<figureCitation id="D3CC2A742D462E4CB558FB61ABADFB43" box="[121,190,1173,1197]" captionStart="Fig" captionStartId="3.[113,148,1916,1938]" captionTargetBox="[106,752,1529,1891]" captionTargetId="figure-883@3.[91,755,1516,1897]" captionTargetPageId="3" captionText="Fig. 3. Boleophthalmus poti, BMNH 2011.1.27.1. Lateral view of the suspensorium. Abbreviations: ap = autopalatine; d = dentary; ep = ectopterygoid; hm = hyomandibular; io = interopercle; m = maxilla; mp = metapterygoid; op = opercle; pm = premaxilla; po = preopercle; pp = ascending processes of the premaxilla;q =quadrate; sop = subopercle; sp = symplectic; st = symphyseal tooth. Scale bar = 1 mm. the original image was reversed horizontally." figureDoi="http://doi.org/10.5281/zenodo.5351914" httpUri="https://zenodo.org/record/5351914/files/figure.png" pageId="3" pageNumber="314">Fig. 3</figureCitation>
). Posterior interorbital pore present. Anterior nare positioned at tip of triangular pendulous flap overhanging upper lip; posterior nare as slit immediately in front of anterior margin of eye. Anterior oculoscapular canal pores anterior to posterior nares, at ~1/3 the distance from eye to upper lip. Gill opening extending for ~4/5 of height of pectoral-fin base. All first-dorsal fin spines protruding from fin margin: elements III–V elongate, element IV longest in both sexes (
<figureCitation id="D3CC2A742D462E4CB5D6FA61AA4BFA43" box="[247,344,1429,1453]" captionStart="Fig" captionStartId="2.[835,869,2002,2024]" captionTargetBox="[835,1474,1286,1979]" captionTargetId="figure-716@2.[835,1476,1285,1979]" captionTargetPageId="2" captionText="Fig.2. Boleophthalmus poti:A, holotype, MSNG 56891; B, paratype, preserved specimen, female (108.2 mm) (BMNH 2011.1.27.1); C, an individual in its natural habitat, on the mud banks of Wapi Creek, Purutu Island, Fly River delta. Scale bars =10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351912" httpUri="https://zenodo.org/record/5351912/files/figure.png" pageId="3" pageNumber="314">Figs. 2B</figureCitation>
,
<figureCitation id="D3CC2A742D462E4CB447FA61AA9AFA43" box="[358,393,1429,1453]" captionStart="Fig" captionStartId="5.[814,853,1868,1890]" captionTargetBox="[813,1454,986,1844]" captionTargetId="figure-437@5.[813,1456,986,1844]" captionTargetPageId="5" captionText="Fig. 5. Shape and colouration of freshly dead species of Boleophthalmus with un-notched dentary teeth: A, female of B. birdsongi (95 mm; ZRC 52250); B, female of B. caeruleomaculatus (106 mm; MSNG 54689); and C, paratype, male of B. poti (83.4 mm; USNM 40566). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351918" httpUri="https://zenodo.org/record/5351918/files/figure.png" pageId="3" pageNumber="314">5C</figureCitation>
). In larger specimens, adpressed pectoral fin reaching a vertical point anterior to terminus of D1. Caudal fin ovate; pelvic fins completely fused into round disc with large pelvic frenum connecting pelvic spines, and basal membrane connecting innermost pelvic rays.
</paragraph>
<paragraph id="4B4836F12D462E4CB60DFEA9AF87FBFB" blockId="3.[812,1453,348,1046]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFEA9A8F4FE9B" bold="true" box="[812,999,348,373]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFEA9A8F3FE9B" bold="true" box="[812,992,348,373]" italics="true" pageId="3" pageNumber="314">Live colouration</emphasis>
.
</emphasis>
— Background colouration of dorsolateral portion of body and head yellowish-brown to greyish (
<figureCitation id="D3CC2A742D462E4CB615FE69A886FE5B" box="[820,917,413,437]" captionStart="Fig" captionStartId="2.[835,869,2002,2024]" captionTargetBox="[835,1474,1286,1979]" captionTargetId="figure-716@2.[835,1476,1285,1979]" captionTargetPageId="2" captionText="Fig.2. Boleophthalmus poti:A, holotype, MSNG 56891; B, paratype, preserved specimen, female (108.2 mm) (BMNH 2011.1.27.1); C, an individual in its natural habitat, on the mud banks of Wapi Creek, Purutu Island, Fly River delta. Scale bars =10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351912" httpUri="https://zenodo.org/record/5351912/files/figure.png" pageId="3" pageNumber="314">Figs. 2C</figureCitation>
,
<figureCitation id="D3CC2A742D462E4CB682FE69A8D6FE5B" box="[931,965,413,437]" captionStart="Fig" captionStartId="5.[814,853,1868,1890]" captionTargetBox="[813,1454,986,1844]" captionTargetId="figure-437@5.[813,1456,986,1844]" captionTargetPageId="5" captionText="Fig. 5. Shape and colouration of freshly dead species of Boleophthalmus with un-notched dentary teeth: A, female of B. birdsongi (95 mm; ZRC 52250); B, female of B. caeruleomaculatus (106 mm; MSNG 54689); and C, paratype, male of B. poti (83.4 mm; USNM 40566). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351918" httpUri="https://zenodo.org/record/5351918/files/figure.png" pageId="3" pageNumber="314">5C</figureCitation>
); ventral portion greyish and paler; in some individuals, isthmus, throat and thoracic region dark grey; scattered tiny whitish speckles covering head, extending anteriorly and dorsally from snout to insertion of D1; 8–10 subvertical, saddle-like, bilaterally asymmetrical and irregular dark brown bars on dorsum (2–3 before D
<quantity id="8C0F9B142D462E4CB0A4FDC9A852FD9B" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="3" pageNumber="314" unit="in" value="1.2">1, 2 in</quantity>
front and behind D1, and 4–5 below D2), saddle-like bars never extending below lateral midline; one horizontal, darker, irregular brownish band 1/3 width of body depth may be present on sides above lateral midline and below saddlelike bars, coursing from pectoral fins to caudal peduncle; sparse whitish speckles present on flanks; several scales on caudal peduncle with whitish margins. Caudal fin with grey interradial membranes and black rays. Anal fin with blackish rays and darkened inter-radial membrane, posterior portion more densely pigmented. Pectoral fins with translucent interradial membranes and pale brown rays, muscular base with similar colouration to body. Pelvic fins with translucent interradial membranes and pale brown rays, proximally dark grey, both dorsally and ventrally.
</paragraph>
<paragraph id="4B4836F12D462E4CB60DFBC8AFA2FAFB" blockId="3.[812,1453,1084,1301]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFBC8AF3FFBBB" bold="true" box="[812,1068,1084,1109]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFBC8AF36FBBB" bold="true" box="[812,1061,1084,1109]" italics="true" pageId="3" pageNumber="314">Preserved colouration</emphasis>
.
</emphasis>
— Body colouration dorsally and laterally greyish, ventrally paler (
<figureCitation id="D3CC2A742D462E4CB1ABFBA9AFEFFB9B" box="[1162,1276,1117,1141]" captionStart="Fig" captionStartId="2.[835,869,2002,2024]" captionTargetBox="[835,1474,1286,1979]" captionTargetId="figure-716@2.[835,1476,1285,1979]" captionTargetPageId="2" captionText="Fig.2. Boleophthalmus poti:A, holotype, MSNG 56891; B, paratype, preserved specimen, female (108.2 mm) (BMNH 2011.1.27.1); C, an individual in its natural habitat, on the mud banks of Wapi Creek, Purutu Island, Fly River delta. Scale bars =10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351912" httpUri="https://zenodo.org/record/5351912/files/figure.png" pageId="3" pageNumber="314">Fig. 2A, B</figureCitation>
); isthmus, throat and thoracic regions grey, dark grey in some individuals. Small white speckles and vertical dark bars visible in preservation; on fins, live brown colour becoming grey and yellow whitish; live whitish pigmentation remains in preservation; other traits as in vivo.
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D462E4CB60DFAC8A896FA7B" pageId="3" pageNumber="314" type="distribution">
<paragraph id="4B4836F12D462E4CB60DFAC8A896FA7B" blockId="3.[812,1452,1340,1429]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFAC8A8A8FABB" bold="true" box="[812,955,1340,1365]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFAC8A8A7FABB" bold="true" box="[812,948,1340,1365]" italics="true" pageId="3" pageNumber="314">Distribution</emphasis>
.
</emphasis>
—
<taxonomicName id="8CF74D722D462E4CB6CBFACBAFDDFAB8" authorityName="Polgar &amp; Jaafar &amp; Konstantinidis" authorityYear="2013" box="[1002,1230,1343,1366]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D462E4CB6CBFACBAFDDFAB8" box="[1002,1230,1343,1366]" italics="true" pageId="3" pageNumber="314">Boleophthalmus poti</emphasis>
</taxonomicName>
is presently known only from the delta of the Fly River, in the
<collectingRegion id="8933F8132D462E4CB033FAA9AE50FA9B" box="[1298,1347,1373,1397]" country="Papua New Guinea" name="Gulf" pageId="3" pageNumber="314">Gulf</collectingRegion>
of
<collectingRegion id="8933F8132D462E4CB04AFAAAAEBFFA9B" box="[1387,1452,1374,1397]" country="Indonesia" name="Papua Barat" pageId="3" pageNumber="314">Papua</collectingRegion>
(
<figureCitation id="D3CC2A742D462E4CB615FA89A86BFA7B" box="[820,888,1405,1429]" captionStart="Fig" captionStartId="4.[137,172,1948,1970]" captionTargetBox="[186,1417,1094,1926]" captionTargetId="figure-191@4.[188,1416,1094,1924]" captionTargetPageId="4" captionText="Fig. 4. Geographical distribution of the species of Boleophthalmus: point data; bird = Boleophthalmus birdsongi; bodd = Boleophthalmus boddarti; caer = Boleophthalmus caeruleomaculatus; duss = Boleophthalmus dussumieri; pect = Boleophthalmus pectinirostris; poti = Boleophthalmus poti. Data from Murdy, 1989; Swennen et al., 1995; Rainboth, 1996; Takita et al., 1999; Nakabo, 2002; Nanami &amp; Takegaki, 2005; Chen et al., 2007; Froese &amp; Pauly, 2007; Ocean Biogeographic Information System, 2007; Baeck et al., 2008; Global Biodiversity Information Facility, 2008; Larson et al., 2008; Polgar &amp; Crosa, 2009. Data in online databases were verified by comparison with museum material and published references." figureDoi="http://doi.org/10.5281/zenodo.5351916" httpUri="https://zenodo.org/record/5351916/files/figure.png" pageId="3" pageNumber="314">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D462E4CB60DFA48A892F9BB" pageId="3" pageNumber="314" type="etymology">
<paragraph id="4B4836F12D462E4CB60DFA48A892F9BB" blockId="3.[812,1453,1468,1621]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFA48A8BBFA3B" bold="true" box="[812,936,1468,1493]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DFA48A8B2FA3B" bold="true" box="[812,929,1468,1493]" italics="true" pageId="3" pageNumber="314">Etymology</emphasis>
.
</emphasis>
— The indigenous people of the Wapi villages of Purutu Island, the
<typeStatus id="944C88532D462E4CB13AFA2AAF5AFA18" box="[1051,1097,1502,1526]" pageId="3" pageNumber="314">type</typeStatus>
locality, call this species ‘poti’, that in their language means ‘spotted’ and refers to the numerous tiny whitish spots present on the head of this species.
</paragraph>
</subSubSection>
<subSubSection id="03ED657A2D462E4AB60DF988ABE2F83C" lastPageId="5" lastPageNumber="316" pageId="3" pageNumber="314" type="discussion">
<paragraph id="4B4836F12D462E4CB60DF988AEB4F89B" blockId="3.[812,1453,1660,1909]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DF988A886F97B" bold="true" box="[812,917,1660,1685]" pageId="3" pageNumber="314">
<emphasis id="7983EAE32D462E4CB60DF988A89DF97B" bold="true" box="[812,910,1660,1685]" italics="true" pageId="3" pageNumber="314">Remarks</emphasis>
.
</emphasis>
— Freshly dead individuals of
<taxonomicName id="8CF74D722D462E4CB1D8F98BAE50F978" authorityName="Polgar &amp; Jaafar &amp; Konstantinidis" authorityYear="2013" box="[1273,1347,1663,1686]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D462E4CB1D8F98BAE50F978" box="[1273,1347,1663,1686]" italics="true" pageId="3" pageNumber="314">B. poti</emphasis>
</taxonomicName>
less than
<quantity id="8C0F9B142D462E4CB60DF969A876F95B" box="[812,869,1693,1717]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="314" unit="cm" value="5.0">5 cm</quantity>
SL (e.g., ZRC 51367) have a transparent, indistinct margin of D1, a much less densely pigmented D2 with columns of brownish spots on a transparent background, hyaline pelvic fins, scattered dark brown spots on the head and flanks on the anterior third of the body, and some irregular brown blotches on flanks, roughly corresponding to the dorsal saddle-like bands observed in larger specimens.
</paragraph>
<caption id="1F8866792D462E4CB550F888A9BDF7D1" ID-DOI="http://doi.org/10.5281/zenodo.5351914" ID-Zenodo-Dep="5351914" httpUri="https://zenodo.org/record/5351914/files/figure.png" pageId="3" pageNumber="314" startId="3.[113,148,1916,1938]" targetBox="[106,752,1529,1891]" targetPageId="3">
<paragraph id="4B4836F12D462E4CB550F888A9BDF7D1" blockId="3.[113,751,1916,2111]" pageId="3" pageNumber="314">
Fig. 3.
<taxonomicName id="8CF74D722D462E4CB596F88AAAC6F87C" authority=", BMNH" authorityName="BMNH" box="[183,469,1917,1938]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D462E4CB596F88AAA6CF87C" box="[183,383,1918,1938]" italics="true" pageId="3" pageNumber="314">Boleophthalmus poti</emphasis>
, BMNH
</taxonomicName>
2011.1.27.1. Lateral view of the suspensorium. Abbreviations: ap = autopalatine; d = dentary; ep = ectopterygoid; hm = hyomandibular; io = interopercle; m = maxilla; mp = metapterygoid; op = opercle; pm = premaxilla; po = preopercle; pp = ascending processes of the premaxilla; q = quadrate; sop = subopercle; sp = symplectic; st = symphyseal tooth. Scale bar = 1 mm. the original image was reversed horizontally.
</paragraph>
</caption>
<paragraph id="4B4836F12D462E4CB60DF86BA86FF7DB" blockId="3.[812,1453,1949,2102]" pageId="3" pageNumber="314">
<taxonomicName id="8CF74D722D462E4CB60DF86BAF5BF858" authorityName="Murdy" authorityYear="1989" box="[812,1096,1951,1974]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="birdsongi">
<emphasis id="7983EAE32D462E4CB60DF86BAF5BF858" box="[812,1096,1951,1974]" italics="true" pageId="3" pageNumber="314">Boleophthalmus birdsongi</emphasis>
</taxonomicName>
was the only congener reported previously to possess flattened dentary teeth, lacking notches (
<bibRefCitation id="2F664B002D462E4CB615F829A8D6F81B" author="Murdy, E" box="[820,965,2013,2037]" pageId="3" pageNumber="314" pagination="1 - 93" refId="ref9074" refString="Murdy, E. O., 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum, Supplement, 11: 1 - 93." type="journal article" year="1989">Murdy, 1989</bibRefCitation>
) but examination of all congeners revealed this tooth
<typeStatus id="944C88532D462E4CB6BAF80AA8DAF7F8" box="[923,969,2046,2070]" pageId="3" pageNumber="314">type</typeStatus>
is also present in
<taxonomicName id="8CF74D722D462E4CB1B2F80BAE6EF7F8" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[1171,1405,2047,2070]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D462E4CB1B2F80BAE6EF7F8" box="[1171,1405,2047,2070]" italics="true" pageId="3" pageNumber="314">B. caeruleomaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8CF74D722D462E4CB60DF7EBA866F7D8" box="[812,885,2079,2102]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="314" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D462E4CB60DF7EBA866F7D8" box="[812,885,2079,2102]" italics="true" pageId="3" pageNumber="314">B. poti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="4B4836F12D412E4BB5A9FC62AFCCFBE3" blockId="4.[136,776,917,1037]" lastBlockId="4.[835,1474,917,1037]" pageId="4" pageNumber="315">
<taxonomicName id="8CF74D722D412E4BB5A9FC62AA7DFC43" box="[136,366,918,941]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D412E4BB5A9FC62AA7DFC43" box="[136,366,918,941]" italics="true" pageId="4" pageNumber="315">Boleophthalmus poti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="8CF74D722D412E4BB72AFC62A987FC43" authorityName="Murdy" authorityYear="1989" box="[523,660,918,941]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="birdsongi">
<emphasis id="7983EAE32D412E4BB72AFC62A987FC43" box="[523,660,918,941]" italics="true" pageId="4" pageNumber="315">B. birdsongi</emphasis>
</taxonomicName>
(
<figureCitation id="D3CC2A742D412E4BB786FC61A9ECFC43" box="[679,767,917,941]" captionStart="Fig" captionStartId="5.[814,853,1868,1890]" captionTargetBox="[813,1454,986,1844]" captionTargetId="figure-437@5.[813,1456,986,1844]" captionTargetPageId="5" captionText="Fig. 5. Shape and colouration of freshly dead species of Boleophthalmus with un-notched dentary teeth: A, female of B. birdsongi (95 mm; ZRC 52250); B, female of B. caeruleomaculatus (106 mm; MSNG 54689); and C, paratype, male of B. poti (83.4 mm; USNM 40566). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351918" httpUri="https://zenodo.org/record/5351918/files/figure.png" pageId="4" pageNumber="315">Fig. 5A</figureCitation>
) in having fewer interdorsal scale rows (3–6 vs. 6–9, respectively; Tables 1, 3), a longer D1 base (13–15% SL vs. 10–13% SL, respectively;
<bibRefCitation id="2F664B002D412E4BB48FFC01A92CFBE3" author="Murdy, E" box="[430,575,1013,1037]" pageId="4" pageNumber="315" pagination="1 - 93" refId="ref9074" refString="Murdy, E. O., 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum, Supplement, 11: 1 - 93." type="journal article" year="1989">Murdy, 1989</bibRefCitation>
; Tables 2, 4), and in the colouration of the unpaired fins: in the latter species, D1 is proximally black and distally blue; D2 has a narrow whitish basal portion, proximally and medially black, and a yellowish to whitish marginal band.
</paragraph>
<caption id="1F8866792D412E4BB5A8F868A977F7AC" ID-DOI="http://doi.org/10.5281/zenodo.5351916" ID-Zenodo-Dep="5351916" httpUri="https://zenodo.org/record/5351916/files/figure.png" pageId="4" pageNumber="315" startId="4.[137,172,1948,1970]" targetBox="[186,1417,1094,1926]" targetPageId="4">
<paragraph id="4B4836F12D412E4BB5A8F868A977F7AC" blockId="4.[137,1475,1948,2114]" pageId="4" pageNumber="315">
Fig. 4. Geographical distribution of the species of
<taxonomicName id="8CF74D722D412E4BB755F869A803F85F" authorityName="Valenciennes" authorityYear="1837" box="[628,784,1949,1969]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="genus">
<emphasis id="7983EAE32D412E4BB755F869A803F85F" box="[628,784,1949,1969]" italics="true" pageId="4" pageNumber="315">Boleophthalmus</emphasis>
</taxonomicName>
: point data; bird =
<taxonomicName id="8CF74D722D412E4BB6EFF869AFDEF85F" authorityName="Murdy" authorityYear="1989" box="[974,1229,1949,1969]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="birdsongi">
<emphasis id="7983EAE32D412E4BB6EFF869AFDEF85F" box="[974,1229,1949,1969]" italics="true" pageId="4" pageNumber="315">Boleophthalmus birdsongi</emphasis>
</taxonomicName>
; bodd =
<taxonomicName id="8CF74D722D412E4BB007F869ABCFF820" baseAuthorityName="Pallas" baseAuthorityYear="1770" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="boddarti">
<emphasis id="7983EAE32D412E4BB007F869ABCFF820" italics="true" pageId="4" pageNumber="315">Boleophthalmus boddarti</emphasis>
</taxonomicName>
; caer =
<taxonomicName id="8CF74D722D412E4BB412F84EA983F820" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[307,656,1978,1998]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D412E4BB412F84EA983F820" box="[307,656,1978,1998]" italics="true" pageId="4" pageNumber="315">Boleophthalmus caeruleomaculatus</emphasis>
</taxonomicName>
; duss =
<taxonomicName id="8CF74D722D412E4BB7CBF84EA8E9F820" authorityName="Valenciennes" authorityYear="1837" box="[746,1018,1978,1998]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="dussumieri">
<emphasis id="7983EAE32D412E4BB7CBF84EA8E9F820" box="[746,1018,1978,1998]" italics="true" pageId="4" pageNumber="315">Boleophthalmus dussumieri</emphasis>
</taxonomicName>
; pect =
<taxonomicName id="8CF74D722D412E4BB170F84EAE64F820" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1105,1399,1978,1998]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="pectinirostris">
<emphasis id="7983EAE32D412E4BB170F84EAE64F820" box="[1105,1399,1978,1998]" italics="true" pageId="4" pageNumber="315">Boleophthalmus pectinirostris</emphasis>
</taxonomicName>
; poti =
<taxonomicName id="8CF74D722D412E4BB5A8F823AA44F805" box="[137,343,2007,2027]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="315" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D412E4BB5A8F823AA44F805" box="[137,343,2007,2027]" italics="true" pageId="4" pageNumber="315">Boleophthalmus poti</emphasis>
</taxonomicName>
. Data from
<bibRefCitation id="2F664B002D412E4BB4F7F822A949F805" author="Murdy, E" box="[470,602,2006,2028]" pageId="4" pageNumber="315" pagination="1 - 93" refId="ref9074" refString="Murdy, E. O., 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum, Supplement, 11: 1 - 93." type="journal article" year="1989">Murdy, 1989</bibRefCitation>
;
<bibRefCitation id="2F664B002D412E4BB746F822A82DF805" author="Swennen, C &amp; Ruttanadakul, M &amp; Haver, S &amp; Piummongkol, S &amp; Prasertsongscum, I &amp; Intanai, W &amp; Chaipakdi, P &amp; Yeesin, P" box="[615,830,2005,2028]" pageId="4" pageNumber="315" pagination="109 - 129" refId="ref9670" refString="Swennen, C., N. Ruttanadakul, M. Haver, S. Piummongkol, S. Prasertsongscum, I. Intanai, W. Chaipakdi, P. Yeesin, P. Horpet &amp; S. Detsathit, 1995. The five sympatric mudskippers (Teleostei: Gobioidea) of Pattani area, southern Thailand. Natural History Bulletin of the Siam Society, 42: 109 - 129." type="journal article" year="1995">Swennen et al., 1995</bibRefCitation>
;
<bibRefCitation id="2F664B002D412E4BB66DF822A8FAF802" author="Rainboth, W" box="[844,1001,2006,2028]" pageId="4" pageNumber="315" refId="ref9478" refString="Rainboth, W., 1996. Fishes of the Cambodian Mekong. FAO Species Identification Field Guide for Fishery Purposes. FAO, Rome. 310 pp." type="book" year="1996">Rainboth, 1996</bibRefCitation>
;
<bibRefCitation id="2F664B002D412E4BB6D4F822AFA2F805" author="Takita, T" box="[1013,1201,2006,2028]" pageId="4" pageNumber="315" pagination="131 - 138" refId="ref9743" refString="Takita, T., Agusnimar &amp; B. Ali Ahyaudin, 1999. Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca. Ichthyological Research, 46: 131 - 138." type="journal article" year="1999">Takita et al., 1999</bibRefCitation>
; Nakabo, 2002; Nanami &amp; Takegaki, 2005;
<bibRefCitation id="2F664B002D412E4BB40EF806AACEF7E6" author="Chen, S &amp; Hong, Q" box="[303,477,2034,2056]" pageId="4" pageNumber="315" pagination="615 - 619" refId="ref8241" refString="Chen, S., W. Hong, Q. Zhang &amp; Y. Su, 2007. Why does the mudskipper Boleophthalmus pectinirostris form territories in farming ponds? Journal of the Marine Biological Association of the United Kingdom, 87: 615 - 619." type="journal article" year="2007">Chen et al., 2007</bibRefCitation>
; Froese &amp; Pauly, 2007;
<bibRefCitation id="2F664B002D412E4BB7F0F807AFA3F7E6" author="Ocean Biogeographic Information System" box="[721,1200,2034,2056]" pageId="4" pageNumber="315" refId="ref9185" refString="Ocean Biogeographic Information System (OBIS), World Wide Web electronic publication. www. iobis. org. (Accessed Dec. 2007)." type="book" year="2007">Ocean Biogeographic Information System, 2007</bibRefCitation>
;
<bibRefCitation id="2F664B002D412E4BB19CF806AE66F7E6" author="Baeck, G" box="[1213,1397,2034,2056]" pageId="4" pageNumber="315" pagination="43 - 52" refId="ref8113" refString="Baeck, G. W., T. Takita &amp; Y. H. Yoon, 2008. Lifestyle of Korean mudskipper Periophthalmus magnuspinnatus with reference to a congeneric species Periophthalmus modestus. Ichthyological Research, 55: 43 - 52." type="journal article" year="2008">Baeck et al., 2008</bibRefCitation>
;
<bibRefCitation id="2F664B002D412E4BB0A0F806A91BF7CA" author="Global Biodiversity Information Facility" pageId="4" pageNumber="315" refId="ref8467" refString="Global Biodiversity Information Facility (GBIF) Data Portal, 2008. World Wide Web electronic publication. www. gbif. net. (Accessed Apr. 2008)." type="book" year="2008">Global Biodiversity Information Facility, 2008</bibRefCitation>
;
<bibRefCitation id="2F664B002D412E4BB732F7E4A9DEF7CA" author="Larson, H" box="[531,717,2063,2085]" pageId="4" pageNumber="315" pagination="135 - 155" refId="ref8851" refString="Larson, H. K., Z. Jaafar &amp; K. K. P. Lim, 2008. An annotated checklist of the gobioid fishes of Singapore. Raffles Bulletin of Zoology, 56: 135 - 155." type="journal article" year="2008">Larson et al., 2008</bibRefCitation>
; Polgar &amp; Crosa, 2009. Data in online databases were verified by comparison with museum material and published references.
</paragraph>
</caption>
<paragraph id="4B4836F12D402E4AB550FC28AA1CFABD" blockId="5.[113,752,986,1363]" pageId="5" pageNumber="316">
<taxonomicName id="8CF74D722D402E4AB550FC28AA48FC1D" box="[113,347,988,1011]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D402E4AB550FC28AA48FC1D" box="[113,347,988,1011]" italics="true" pageId="5" pageNumber="316">Boleophthalmus poti</emphasis>
</taxonomicName>
differs from
<taxonomicName id="8CF74D722D402E4AB4DBFC28A9FCFC1D" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[506,751,988,1011]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D402E4AB4DBFC28A9FCFC1D" box="[506,751,988,1011]" italics="true" pageId="5" pageNumber="316">B. caeruleomaculatus</emphasis>
</taxonomicName>
(
<figureCitation id="D3CC2A742D402E4AB55BFC0EABC5FBFC" box="[122,214,1018,1042]" captionStart="Fig" captionStartId="5.[814,853,1868,1890]" captionTargetBox="[813,1454,986,1844]" captionTargetId="figure-437@5.[813,1456,986,1844]" captionTargetPageId="5" captionText="Fig. 5. Shape and colouration of freshly dead species of Boleophthalmus with un-notched dentary teeth: A, female of B. birdsongi (95 mm; ZRC 52250); B, female of B. caeruleomaculatus (106 mm; MSNG 54689); and C, paratype, male of B. poti (83.4 mm; USNM 40566). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351918" httpUri="https://zenodo.org/record/5351918/files/figure.png" pageId="5" pageNumber="316">Fig. 5B</figureCitation>
) in the smaller number of lateral-scale rows (97–119 vs. 130–151, respectively; Tables 1, 3), a shorter D2 base (40–43% SL vs. 43–48% SL, respectively) and a shorter pelvic disc (11–12%SL vs. 16–18%SL, respectively; Tables 2, 4;
<bibRefCitation id="2F664B002D402E4AB5D0FB8EAA6DFB7C" author="Murdy, E" box="[241,382,1146,1170]" pageId="5" pageNumber="316" pagination="1 - 93" refId="ref9074" refString="Murdy, E. O., 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum, Supplement, 11: 1 - 93." type="journal article" year="1989">Murdy, 1989</bibRefCitation>
), in body colouration, which in the latter species includes sky-blue speckles on head, dorsum and flanks, and a sky-blue patch of skin below the orbits, and in the colouration of the unpaired fins:
<taxonomicName id="8CF74D722D402E4AB729FB28A9E3FB1D" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[520,752,1244,1267]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D402E4AB729FB28A9E3FB1D" box="[520,752,1244,1267]" italics="true" pageId="5" pageNumber="316">B. caeruleomaculatus</emphasis>
</taxonomicName>
lacks distinct marginal bands and has sky-blue spots on the interradial membranes, arranged in series along the whole length of rays.
</paragraph>
<paragraph id="4B4836F12D402E4AB550FA88A9A1F9DC" blockId="5.[113,752,1402,1586]" pageId="5" pageNumber="316">
<taxonomicName id="8CF74D722D402E4AB550FA88AAFCFA7D" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[113,495,1404,1427]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D402E4AB550FA88AAFCFA7D" box="[113,495,1404,1427]" italics="true" pageId="5" pageNumber="316">Boleophthalmus caeruleomaculatus</emphasis>
</taxonomicName>
also attains a larger size than the other two species (maximum recorded size =
<quantity id="8C0F9B142D402E4AB7E9FA6EAB8EFA3C" metricMagnitude="-1" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="316" unit="mm" value="165.0">165 mm</quantity>
, vs.
<quantity id="8C0F9B142D402E4AB5F0FA4EAA3AFA3C" box="[209,297,1466,1490]" metricMagnitude="-1" metricUnit="m" metricValue="1.1099999999999999" pageId="5" pageNumber="316" unit="mm" value="111.0">111 mm</quantity>
and
<quantity id="8C0F9B142D402E4AB440FA4EAAA8FA3C" box="[353,443,1466,1490]" metricMagnitude="-1" metricUnit="m" metricValue="1.06" pageId="5" pageNumber="316" unit="mm" value="106.0">106 mm</quantity>
of
<taxonomicName id="8CF74D722D402E4AB4C2FA48A97BFA3D" authorityName="Murdy" authorityYear="1989" box="[483,616,1468,1491]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="birdsongi">
<emphasis id="7983EAE32D402E4AB4C2FA48A97BFA3D" box="[483,616,1468,1491]" italics="true" pageId="5" pageNumber="316">B. birdsongi</emphasis>
</taxonomicName>
and
<taxonomicName id="8CF74D722D402E4AB781FA48A9FAFA3D" box="[672,745,1468,1491]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D402E4AB781FA48A9FAFA3D" box="[672,745,1468,1491]" italics="true" pageId="5" pageNumber="316">B. poti</emphasis>
</taxonomicName>
, respectively;
<bibRefCitation id="2F664B002D402E4AB422FA2EAA81FA1C" author="Murdy, E" box="[259,402,1498,1522]" pageId="5" pageNumber="316" pagination="1 - 93" refId="ref9074" refString="Murdy, E. O., 1989. A taxonomic revision and cladistic analysis of the oxudercine gobies (Gobiidae: Oxudercinae). Records of the Australian Museum, Supplement, 11: 1 - 93." type="journal article" year="1989">Murdy, 1989</bibRefCitation>
;
<bibRefCitation id="2F664B002D402E4AB4BEFA2EA97AFA1C" author="Takita, T" box="[415,617,1498,1522]" pageId="5" pageNumber="316" pagination="189 - 204" refId="ref9784" refString="Takita, T., H. Larson &amp; A. Ishimatsu, 2011. The natural history of mudskippers in northern Australia, with field identification characters. The Beagle. Records of the Museums and Art Galleries of the Northern Territory, 27: 189 - 204." type="journal article" year="2011">Takita et al., 2011</bibRefCitation>
; this study), and exhibits sexual dimorphism in the D1: females have elongated spines III–V, the IV being the longest one.
</paragraph>
<paragraph id="4B4836F12D402E4AB550F9AEA98BF8FC" blockId="5.[113,752,1626,1810]" pageId="5" pageNumber="316">
Multivariate analyses of meristic and morphometric datasets (n = 25;
<tableCitation id="0675034A2D402E4AB40AF98EAA94F97C" box="[299,391,1658,1682]" captionStart="Table 3" captionStartId="6.[111,164,680,702]" captionText="Table 3. Meristic dataset of the PCO analysis for B. birdsongi, B. caeruleomaculatus, and B. poti. The cleared paratype (BMNH 2011.1.27.1) is not included." pageId="5" pageNumber="316">Tables 3</tableCitation>
,
<tableCitation id="0675034A2D402E4AB4B5F98EAAB7F97C" box="[404,420,1658,1682]" captionStart="Table 4" captionStartId="7.[111,164,623,645]" captionText="Table 4. Morphometric dataset of the PCA for B. birdsongi, B. caeruleomaculatus, and B. poti; measurements are in mm. HW was not included in this analysis, since it varies with the degree of expansion of the opercles; SL and TL were also not included, since they do not convey information about shape. The cleared paratype (BMNH 2011.1.27.1) is not included;? = missing values (damaged caudal fins)." pageId="5" pageNumber="316">4</tableCitation>
;
<bibRefCitation id="2F664B002D402E4AB491F98FA99CF97C" author="Hammer, O" box="[432,655,1658,1682]" pageId="5" pageNumber="316" refId="ref8533" refString="Hammer, O., D. A. T. Harper &amp; P. D. Ryan, 2001. PAST: Paleontological statistics software package for education and data analysis. Palaeontologia Electronica, 4: art. 4, 9 pp. http: // palaeo-electronica. org / 2001 _ 1 / past / issue 1 _ 01. htm." type="book" year="2001">Hammer et al., 2001</bibRefCitation>
) showed that
<taxonomicName id="8CF74D722D402E4AB584F968AA3CF95D" authorityName="Murdy" authorityYear="1989" box="[165,303,1692,1715]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="birdsongi">
<emphasis id="7983EAE32D402E4AB584F968AA3CF95D" box="[165,303,1692,1715]" italics="true" pageId="5" pageNumber="316">B. birdsongi</emphasis>
</taxonomicName>
,
<taxonomicName id="8CF74D722D402E4AB460F968A927F95D" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[321,564,1692,1715]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D402E4AB460F968A927F95D" box="[321,564,1692,1715]" italics="true" pageId="5" pageNumber="316">B. caeruleomaculatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="8CF74D722D402E4AB756F968A9D7F95D" box="[631,708,1692,1715]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D402E4AB756F968A9D7F95D" box="[631,708,1692,1715]" italics="true" pageId="5" pageNumber="316">B. poti</emphasis>
</taxonomicName>
are morphologically distinct (
<figureCitation id="D3CC2A742D402E4AB4AAF94EA916F93C" box="[395,517,1722,1746]" captionStart="Fig" captionStartId="8.[136,171,1861,1883]" captionTargetBox="[135,1476,1238,1831]" captionTargetId="figure-8@8.[104,1495,1197,1837]" captionTargetPageId="8" captionText="Fig. 6. Multivariate morphological analyses of the three species of Boleophthalmus without notched dentary teeth, supporting the separation of the examined specimens in 3 morphologically distinct groups.A: PCO scatter plot of meristic characters, after logarithmic transformation. The two coordinate axes Coord 1 and Coord 2 explained the highest percentage of total variance (in parentheses); closed lined (convex hulls) delimit clusters of specimens of the same species. B: PCA distance biplot of morphometric characters after Darroch &amp; Mosimann transformation (1985). The two principal component axes PC1 and PC2 explained the highest percentage of total variance (in parentheses); closed lines (convex hulls) delimit clusters of specimens of the same species; the variables are projected onto the scattergram as vectors, graphically visualising their PCA loadings, or the degree they enter into the principal components. PLVL separates caer from bird and poti, while D1BL separates bird from caer and poti. bird = B. birdsongi (n = 11); caer = B. caeruleomaculatus (n = 4); poti = B. poti (n = 10). See Tables 3, 4 for the dataset." figureDoi="http://doi.org/10.5281/zenodo.5351920" httpUri="https://zenodo.org/record/5351920/files/figure.png" pageId="5" pageNumber="316">Fig. 6A, B</figureCitation>
). Consistent with the above results, the length of D1 base and the pelvic fin length had higher PC1 and PC2 loadings (&gt;0.5;
<figureCitation id="D3CC2A742D402E4AB717F90EA999F8FC" box="[566,650,1786,1810]" captionStart="Fig" captionStartId="8.[136,171,1861,1883]" captionTargetBox="[135,1476,1238,1831]" captionTargetId="figure-8@8.[104,1495,1197,1837]" captionTargetPageId="8" captionText="Fig. 6. Multivariate morphological analyses of the three species of Boleophthalmus without notched dentary teeth, supporting the separation of the examined specimens in 3 morphologically distinct groups.A: PCO scatter plot of meristic characters, after logarithmic transformation. The two coordinate axes Coord 1 and Coord 2 explained the highest percentage of total variance (in parentheses); closed lined (convex hulls) delimit clusters of specimens of the same species. B: PCA distance biplot of morphometric characters after Darroch &amp; Mosimann transformation (1985). The two principal component axes PC1 and PC2 explained the highest percentage of total variance (in parentheses); closed lines (convex hulls) delimit clusters of specimens of the same species; the variables are projected onto the scattergram as vectors, graphically visualising their PCA loadings, or the degree they enter into the principal components. PLVL separates caer from bird and poti, while D1BL separates bird from caer and poti. bird = B. birdsongi (n = 11); caer = B. caeruleomaculatus (n = 4); poti = B. poti (n = 10). See Tables 3, 4 for the dataset." figureDoi="http://doi.org/10.5281/zenodo.5351920" httpUri="https://zenodo.org/record/5351920/files/figure.png" pageId="5" pageNumber="316">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="4B4836F12D402E4AB550F8C8ABE2F83C" blockId="5.[113,752,1850,2002]" pageId="5" pageNumber="316">
<taxonomicName id="8CF74D722D402E4AB550F8C8AA4AF8BD" box="[113,345,1852,1875]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="poti">
<emphasis id="7983EAE32D402E4AB550F8C8AA4AF8BD" box="[113,345,1852,1875]" italics="true" pageId="5" pageNumber="316">Boleophthalmus poti</emphasis>
</taxonomicName>
is traditionally consumed and used as bait in the delta of the Fly River, together with the other mudskipper species
<taxonomicName id="8CF74D722D402E4AB492F888A9A9F87D" authorityName="McCulloch &amp; Waite" authorityYear="1918" box="[435,698,1916,1939]" class="Actinopterygii" family="Gobiidae" genus="Boleophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="caeruleomaculatus">
<emphasis id="7983EAE32D402E4AB492F888A9A9F87D" box="[435,698,1916,1939]" italics="true" pageId="5" pageNumber="316">B. caeruleomaculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="8CF74D722D402E4AB550F868A964F85C" authority="Quoy &amp; Gaimard" authorityName="Quoy &amp; Gaimard" box="[113,631,1946,1971]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="316" phylum="Chordata" rank="species" species="freycineti">
<emphasis id="7983EAE32D402E4AB550F868AAB9F85D" box="[113,426,1948,1971]" italics="true" pageId="5" pageNumber="316">Periophthalmodon freycineti</emphasis>
Quoy &amp; Gaimard
</taxonomicName>
(Polgar &amp; Lim, 2011).
</paragraph>
</subSubSection>
</treatment>
</document>
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