208 lines
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208 lines
25 KiB
XML
<document ID-CLB-Dataset="6286" ID-DOI="http://dx.doi.org/10.3897/zookeys.1042.66576" ID-GBIF-Dataset="598dbded-7da1-48b8-b8fa-2e648b22a198" ID-PMC="PMC8195948" ID-Pensoft-Pub="1313-2970-1042-1" ID-Pensoft-UUID="0930CA7A3EDB571AA10D026EF39B1739" ID-PubMed="34163286" ID-ZooBank="B0B0F76E89504A0AB45D701730486B56" ModsDocID="1313-2970-1042-1" checkinTime="1622838178301" checkinUser="pensoft" docAuthor="Zhang, Wen-Xuan, Hu, Fang-Shuo & Yin, Zi-Wei" docDate="2021" docId="BC7F7BA6A02657A0AB24449D96A75047" docLanguage="en" docName="ZooKeys 1042: 1-22" docOrigin="ZooKeys 1042" docPubDate="2021-06-04" docSource="http://dx.doi.org/10.3897/zookeys.1042.66576" docTitle="Horniella sabahensis Zhang & Yin 2021, sp. nov." docType="treatment" docUuid="EC5C6E94-0D2A-461D-8EA7-1C750F093066" docUuidSource="ZooBank" docVersion="6" id="0930CA7A3EDB571AA10D026EF39B1739" lastPageNumber="1" masterDocId="0930CA7A3EDB571AA10D026EF39B1739" masterDocTitle="Six new species of Horniella Raffray from the Oriental region (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="1" updateTime="1720468530034" updateUser="ExternalLinkService">
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<mods:mods id="885DD3E57521B9985F32940634E3A910" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1949E3C2F2E9F91F5767BC98B4EE69B6">
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<mods:title id="46A0165D088604E47FA9CA28A339AFDA">Six new species of Horniella Raffray from the Oriental region (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="DB936D6AD74AC87C2623DCD4AC06E3E2" type="personal">
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<mods:roleTerm id="3F6E087E6AF7EB6A4C0082B5796EE499">Author</mods:roleTerm>
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<mods:namePart id="029A31C37A19B1E3562017B90A7306CE">Zhang, Wen-Xuan</mods:namePart>
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<mods:affiliation id="132288F9E13C68925ECEC344A2B6B2BB">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
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</mods:name>
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<mods:name id="CCBE009C9EA647E1596A9C8AE1720A94" type="personal">
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<mods:roleTerm id="6173251E5AF8F4264F895FDECD173420">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="37DA0FC57CBC43088DA81961975A6B64">Hu, Fang-Shuo</mods:namePart>
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<mods:nameIdentifier id="028A65FBA88BF06DCBB8F6062039FFA7" type="ORCID">https://orcid.org/0000-0003-1269-8904</mods:nameIdentifier>
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<mods:affiliation id="4E510E230BB8F8FCE609C03611EF4F50">Department of Entomology, National Chung Hsing University 145 Xingda Rd., South District, Taichung City 402, Taiwan, China</mods:affiliation>
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</mods:name>
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<mods:name id="D0783CEB7E733C7585344567D8A7001B" type="personal">
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<mods:roleTerm id="E12182287EC4D1BD9489E331F7D9FD46">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E42C9994057866D02690AFB0FAF6D135">Yin, Zi-Wei</mods:namePart>
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<mods:nameIdentifier id="F588302F10EF731CE8F84897A3D007FD" type="ORCID">https://orcid.org/0000-0001-6659-9448</mods:nameIdentifier>
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<mods:affiliation id="9F818581309F337B7E2432C8CE190F73">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
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<mods:nameIdentifier id="2B4AD8C70E17D94222F40991852B558D" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="621E26BD7782B631EC1B2D84998B013A">text</mods:typeOfResource>
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<mods:titleInfo id="B94E5EEF4174093C766ECA1618550104">
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<mods:title id="7404F42355DE8107C9AE4EEFE0A0A2CB">ZooKeys</mods:title>
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<mods:part id="423AA0A9BB201270A77BF37F5A6C1A7C">
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<mods:date id="983001BC39607A2B669E76EAC4B9719B">2021</mods:date>
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<mods:detail id="F532C1BE510D00A918AAB079DE5A88BC" type="pubDate">
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<mods:number id="95E21277DFF21A561200C9C0EACFDA42">2021-06-04</mods:number>
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<mods:detail id="074047551A9D62AD249E5066AEA38029" type="volume">
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<mods:number id="1C021A1406407C0C4F138157E80A82CC">1042</mods:number>
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<mods:start id="DF60E4477592115713C6D29D25FEE7EA">1</mods:start>
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<mods:end id="DA3D2640796C9FB727C735A0CC564F78">22</mods:end>
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<mods:url id="DFCA8017A0D6FD38CC56099F8AD83E9E">http://dx.doi.org/10.3897/zookeys.1042.66576</mods:url>
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<mods:classification id="7441DBAE9150A9B65267B6B51CAD5333">journal article</mods:classification>
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<mods:identifier id="FD213BFE5957C2F22FCD05F09C8E1AB6" type="DOI">http://dx.doi.org/10.3897/zookeys.1042.66576</mods:identifier>
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<mods:identifier id="9FAF685C7BFFA8833E331B41883694EC" type="Pensoft-Pub">1313-2970-1042-1</mods:identifier>
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<treatment id="BC7F7BA6A02657A0AB24449D96A75047" ID-GBIF-Taxon="182494861" LSID="urn:lsid:zoobank.org:act:EC5C6E94-0D2A-461D-8EA7-1C750F093066" httpUri="http://treatment.plazi.org/id/BC7F7BA6A02657A0AB24449D96A75047" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="A5BB79E0CFA9397AC9D0FC9AE9E493DC" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="DF968F1E474F255533AB1A9765BF1A24" pageId="0" pageNumber="1">
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<taxonomicName id="4E192E238DC4E95A3A7998A0210B00DF" ID-CoL="8Y4MG" LSID="http://zoobank.org/EC5C6E94-0D2A-461D-8EA7-1C750F093066" authority="Zhang & Yin" authorityName="Zhang & Yin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Horniella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Horniella sabahensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">Horniella sabahensis Zhang & Yin</taxonomicName>
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<taxonomicNameLabel id="F8BFA1B0E3A04C760DC2817B1FD18A9D" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="8F6B7BB9B0E16C0728445934E6ACD762" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dorsal habitus of Horniella species A H. sabahensis sp. nov. B H. taiwanensis sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1048.66576.figure7" httpUri="https://binary.pensoft.net/fig/551740" pageId="0" pageNumber="1">Figures 7A</figureCitation>
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<figureCitation id="CAA14668A70070E2E9F03C70D9C8157B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">, 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DC78EB537479FB32959EBFBD99C4E8D4" pageId="0" pageNumber="1" type="type material">
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<paragraph id="8FECF4D09D1D84F8EBA86E5B807EB158" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="3864DA173F17C7667898A4ADB4E9CCC1" pageId="0" pageNumber="1">
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<emphasis id="F38512988212052AC62F0262FD5EC30E" bold="true" pageId="0" pageNumber="1">
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<emphasis id="A612D6F63C39D2DC61BDEAAAC3121F67" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: East Malaysia
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</emphasis>
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: ♂, 'Borneo: Sabah, Batu Punggul Resort, primary forest, 24.vi.-1.vii.96, Kodada lgt. / vegetation debris and forest floor litter accumulated around large trees near river.' (MHNG).
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</paragraph>
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<caption id="88FC523774323626EFDF6D4AE9F0F299" doi="10.3897/zookeys.1048.66576.figure7" httpUri="https://binary.pensoft.net/fig/551740" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
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<paragraph id="B2DCEFA978C331FD64D755DB9DA1667A" pageId="0" pageNumber="1">
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<emphasis id="CCEC978C5288F759CFF0075263C37D2A" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
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Dorsal habitus of
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<taxonomicName id="B1610A254D967D417020CE20F81109C5" class="Insecta" family="Staphylinidae" genus="Horniella" kingdom="Animalia" lsidName="Horniella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="0E2586E04C14479A5C1E1EC075CCC427" italics="true" pageId="0" pageNumber="1">Horniella</emphasis>
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</taxonomicName>
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species
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<emphasis id="C0DE58C1832EA70E111BA794012A6D20" bold="true" pageId="0" pageNumber="1">A</emphasis>
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<taxonomicName id="AF6A64797E8D061144BEE6B9ED8A5A0E" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. sabahensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="2E1304CAE6EC101E704A18B5BF923B7F" italics="true" pageId="0" pageNumber="1">H. sabahensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="59F59B15EF88693790DE213AF8B4AA6C" bold="true" pageId="0" pageNumber="1">B</emphasis>
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<taxonomicName id="9229E40F3A467D9A463A62E6BB95977C" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. taiwanensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="taiwanensis">
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<emphasis id="E0434E40DC9D4EF9C7AEC75869DAD386" italics="true" pageId="0" pageNumber="1">H. taiwanensis</emphasis>
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</taxonomicName>
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sp. nov. Scale bars: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="8AD66292F9D2486A521B338A191AFB06" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="1551DC0C86F0DFCF53A93301E0DAFC06" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="7100B9A3A948C54123F60219E0F317F7" pageId="0" pageNumber="1">
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<emphasis id="988713AB068A085E76F89880D46B0B11" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
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Head longer than wide, anterolateral genal projections weakly developed, anterior margin of projection oblique; scape lacking expansion at lateral margin, antennomeres 9-11 moderately enlarged, forming distinct club. Pronotum rounded at lateral margins. Profemur with two tiny ventral spines near base; metatibia with preapical triangular denticle. Tergite 1 (IV) with median carina extending posteriorly for approximately 3/4 of tergal length, lacking discal carinae, tergite 2 (V) with short median carina. Aedeagus with slightly asymmetric median lobe, apical portion of median lobe narrowed, apex truncate in dorso-ventral view; endophallus lacking sclerite, composed of elongate membranous structure with many small denticles.
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</paragraph>
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</subSubSection>
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<subSubSection id="56EFB9226D5B786750B873749ADC1655" pageId="0" pageNumber="1" type="description">
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<paragraph id="D95136D792B2641602F5B324F944109D" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="A2053D1D9415BB5FDF656E392F01BE3B" pageId="0" pageNumber="1">
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<emphasis id="FC00C8A27059DA48A75F0FD8D799BEF6" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
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Body reddish-brown, length 3.41 mm. Head (Fig.
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<figureCitation id="637BAC5588A03F303C25218A365CD176" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8A</figureCitation>
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) longer than wide, HL 0.68 mm, HW 0.59 mm; anterolateral genal projection weakly developed, anterior margin of projection oblique; antenna 1.96 mm long, scape lacking expansion at lateral margin, antennomeres 2-8 slightly elongate or moniliform, distinct club formed by apical three enlarged antennomeres, antennomere 11 largest, slightly shorter than antennomeres 9 and 10 combined; indistinct lateral postantennal pits present; eyes prominent, each composed of approximately 40 large facets, usual area of ocular canthus only slightly prominent (Fig.
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<figureCitation id="371A3B5FE3B7924B9529D7C2F4F1425C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8B</figureCitation>
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).
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</paragraph>
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<caption id="E6013A062961D44F9F335CE53C3717E2" doi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
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<paragraph id="6B0DDB965E22AF218BE3150592611C9E" pageId="0" pageNumber="1">
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<emphasis id="A5CFB45EFD52B8E86A545D4A8FB650F0" bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
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Diagnostic characters of
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<taxonomicName id="1A261BE02DDBBF4D735D42175CE23BD4" authorityName="Zhang & Yin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Horniella" kingdom="Animalia" lsidName="Horniella sabahensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="C8374B6926A60A252DDD49D0CD518F7C" italics="true" pageId="0" pageNumber="1">Horniella sabahensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis id="95557CF455EAED1C27DCFBBC8F9A65DF" bold="true" pageId="0" pageNumber="1">A</emphasis>
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left half of head, in dorsal view
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<emphasis id="43866AA4FEF2176E8601A78A26D9BCE6" bold="true" pageId="0" pageNumber="1">B</emphasis>
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head, in lateral view
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<emphasis id="324EA0718B2580753C3FACFAF2F1564D" bold="true" pageId="0" pageNumber="1">C</emphasis>
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protrochanter and profemur
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<emphasis id="0B9D2E58169B762B108D885522E4F25F" bold="true" pageId="0" pageNumber="1">D</emphasis>
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protibia
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<emphasis id="ADC1E88CF209855162C5C36A88368E9D" bold="true" pageId="0" pageNumber="1">E</emphasis>
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mesotrochanter and mesofemur
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<emphasis id="074E19F5D2EDC9FCFA227D3436D6EBFA" bold="true" pageId="0" pageNumber="1">F</emphasis>
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mesotibia
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<emphasis id="7156ED6A754215D4505B89897F00021A" bold="true" pageId="0" pageNumber="1">G</emphasis>
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metatibia
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<emphasis id="A107FF9904A52271C1759E5AE2065292" bold="true" pageId="0" pageNumber="1">H-J</emphasis>
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aedeagus, in dorsal (
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<emphasis id="A7ED2CA9CAF6A97E09AC792E0F38A687" bold="true" pageId="0" pageNumber="1">H</emphasis>
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), lateral (
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<emphasis id="604935F092BD07878519C4E0AA56001E" bold="true" pageId="0" pageNumber="1">I</emphasis>
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), and ventral (
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<emphasis id="11614D63E3FD01A6B6F5CCCC9947709C" bold="true" pageId="0" pageNumber="1">J</emphasis>
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) view. Scale bars: 0.2 mm (
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<emphasis id="0D9FED84C2286172B89FB7F3DB00132F" bold="true" pageId="0" pageNumber="1">A, B, H-J</emphasis>
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); 0.3 mm (
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<emphasis id="8AEDA603DA605DDC52BD1BC375CEA3B6" bold="true" pageId="0" pageNumber="1">C-G</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph id="FBB2AE56214D32E3AD26338ADF750811" pageId="0" pageNumber="1">Pronotum distinctly longer than wide, PL 0.71 mm, PW 0.6 mm; widest at middle; lateral margins rounded; disc moderately convex, finely punctate, with distinct median antebasal and lateral antebasal foveae connected by complete transverse antebasal sulcus.</paragraph>
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<paragraph id="BD3920EBB815DD4BB470EEC25DDCA9B6" pageId="0" pageNumber="1">Elytra much wider than long, EL 0.77 mm, EW 1.2 mm; each elytron with two large, setose basal foveae; discal striae extending from outer basal foveae to apical 2/3 of elytral length.</paragraph>
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<paragraph id="068CBAD49050AAC08E4355B94330ACAA" pageId="0" pageNumber="1">
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Legs elongate; protrochanter (Fig.
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<figureCitation id="A101DCF3AF08320CB7DD794794B1E817" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8C</figureCitation>
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) simple, profemur (Fig.
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<figureCitation id="19F255201F13524FB4553F959F9F040C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8C</figureCitation>
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) with two tiny ventral spines at base, protibia (Fig.
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<figureCitation id="CB7026BA9B600B64829BE5273BFA473C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8D</figureCitation>
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) simple; mesotrochanter, mesofemur (Fig.
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<figureCitation id="4FEA23AABA025CD26D38F6AA00FE6EB7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8E</figureCitation>
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) and mesotibia (Fig.
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<figureCitation id="B30FA7C9E910DC04284DDD357B9A60ED" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8F</figureCitation>
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) simple; metatibia (Fig.
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<figureCitation id="F865321FDEA55DA09C5ED4098B44AC90" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8G</figureCitation>
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) with distinct apical triangular denticle.
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</paragraph>
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<paragraph id="5E63E88D35A1E1352ED243F465F28B07" pageId="0" pageNumber="1">Abdomen slightly longer than broad, broadest at lateral margins of tergite 1 (IV), AL 1.25 mm, AW 1.16 mm; tergite 1 (IV) largest, slightly shorter than tergites 2 and 3 (V and VI) combined, with median carina extending to near basal 3/4 of tergal length, lacking discal carinae, tergite 2 (V) with median carina extending to near basal 1/4 of tergal length, tergites 2-4 (V-VII) each with small basolateral foveae. Sternite 2 (IV) with broad basal sulcus, lacking mediobasal foveae, basolateral foveae developed as large cuticular pockets, with two pairs of antebasal nodules, sternites 3-5 (V-VII) with basolateral foveae, one median and two lateral nodules, sternite 7 (IX) nearly oval, with well-sclerotized apical half and less sclerotized basal half.</paragraph>
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<paragraph id="1B074BADD0EBE9A7D34F0DE5680D8D93" pageId="0" pageNumber="1">
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Aedeagus (Fig.
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<figureCitation id="2D298224474E8FF8B34D4360C1B03364" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Diagnostic characters of Horniella sabahensis sp. nov. A left half of head, in dorsal view B head, in lateral view C protrochanter and profemur D protibia E mesotrochanter and mesofemur F mesotibia G metatibia H-J aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.2 mm (A, B, H-J); 0.3 mm (C-G)." figureDoi="10.3897/zookeys.1048.66576.figure8" httpUri="https://binary.pensoft.net/fig/551741" pageId="0" pageNumber="1">8H-J</figureCitation>
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) 0.57 mm long, with slightly asymmetric median lobe, apical part of median lobe narrowed, apex broadly truncate in dorso-ventral view; endophallus lacking strongly sclerotized structures, composed of broad, elongate membrane with numerous small denticles.
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</paragraph>
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<paragraph id="441AAB960A48E96CC38628CB4689662C" pageId="0" pageNumber="1">
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<emphasis id="AD3B9CFE3256CD1B311F22D3222041F7" bold="true" pageId="0" pageNumber="1">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="ADA34120DB8EA0752AB95D64F66F9C52" pageId="0" pageNumber="1" type="comparisons">
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<paragraph id="AC109951C59E95442BBA8FEBF6FF7615" pageId="0" pageNumber="1">Comparative notes.</paragraph>
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<paragraph id="159803D07A86AF329A823FD7E6F7E249" pageId="0" pageNumber="1">
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<taxonomicName id="D23A14FEF45C9450BC6935B492D4F425" authorityName="Zhang & Yin" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Horniella" kingdom="Animalia" lsidName="Horniella sabahensis" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="F5EEFE62978EF9EC3C5D479D7CD9F400" italics="true" pageId="0" pageNumber="1">Horniella sabahensis</emphasis>
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</taxonomicName>
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sp. nov. is placed as a member of the
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<taxonomicName id="913C0811D9C41FD67B741AB2CBC088A5" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. hirtella" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hirtella">
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<emphasis id="984596A0EDB7EF65C75B15D4DB62E135" italics="true" pageId="0" pageNumber="1">H. hirtella</emphasis>
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</taxonomicName>
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group. Males of this species share with
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<taxonomicName id="B3564D0FEF4285D829764D4097B8A91F" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. prolixo" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="prolixo">
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<emphasis id="D9DA000AB874AF94CEF9C31AEFDE4BD0" italics="true" pageId="0" pageNumber="1">H. prolixo</emphasis>
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</taxonomicName>
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Yin & Li from Thailand the weakly developed anterolateral genal projections, lack of an expansion at the lateral margin of the scape, and a moderately expanded preapical portion of the metatibia. They can be best separated by the larger body size (3.41 mm vs 2.95-3.02 mm), tergite V with a short median carina (lacking in
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<taxonomicName id="8CB3DFB52F772F0553F0BA07167A2EDB" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. prolixo" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="prolixo">
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<emphasis id="D31B9B3BC06BD144515150AE8E418F3F" italics="true" pageId="0" pageNumber="1">H. prolixo</emphasis>
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</taxonomicName>
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), as well as the much narrower apex of the aedeagus of the new species.
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</paragraph>
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</subSubSection>
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<subSubSection id="054C39ECAE1C176001D4C9E4E242842A" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="913BF0DC8F350491AAACCD28D3299868" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="F42AF22E1B65C75A8D10EC29AD3F6E2E" pageId="0" pageNumber="1">East Malaysia: Sabah.</paragraph>
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</subSubSection>
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<subSubSection id="0BBA9132BF82B4EC959999D5BA470699" pageId="0" pageNumber="1" type="etymology">
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<paragraph id="198C3ED275AC0567B66F47A1864EB17F" pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph id="31D86798519467E42F22AAFD078391E5" pageId="0" pageNumber="1">The new species is named after its type locality, Sabah, East Malaysia.</paragraph>
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</subSubSection>
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</treatment>
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</document> |