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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62253" ID-GBIF-Dataset="0a292fd0-be2d-427e-b277-c19af79971db" ID-PMC="PMC8222342" ID-Pensoft-Pub="1313-2970-1044-153" ID-Pensoft-UUID="944578388D005036B42D68229F152A89" ID-PubMed="34183876" ID-ZooBank="A2219253DB2B4C5992D677595A6C1438" ModsDocID="1313-2970-1044-153" checkinTime="1623875446139" checkinUser="pensoft" docAuthor="Maddison, David R. &amp; Porch, Nick" docDate="2021" docId="BC7D7FFF74C25896A285A134C22A8F60" docLanguage="en" docName="ZooKeys 1044: 153-196" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62253" docTitle="Tasmanitachoides erwini Maddison &amp; Porch 2021, sp. nov." docType="treatment" docUuid="5FF236BF-1E87-4480-8798-230E71470E66" docUuidSource="ZooBank" docVersion="5" id="944578388D005036B42D68229F152A89" lastPageNumber="153" masterDocId="944578388D005036B42D68229F152A89" masterDocTitle="A preliminary phylogeny and review of the genus Tasmanitachoides, with descriptions of two new species (Coleoptera, Carabidae, Bembidarenini)" masterLastPageNumber="196" masterPageNumber="153" pageNumber="153" updateTime="1732566981577" updateUser="ExternalLinkService">
<mods:mods id="3E0335E820B83D3A1C016C7D3239B388" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="509A40F87BF76FAFF1ECED2884E9DA41">
<mods:title id="127E24828E073EC85375E87F3E3B0A98">A preliminary phylogeny and review of the genus Tasmanitachoides, with descriptions of two new species (Coleoptera, Carabidae, Bembidarenini)</mods:title>
</mods:titleInfo>
<mods:name id="128AEFCB40788498650DFF6A5A942107" type="personal">
<mods:role id="8F2EDB120AA54886A053E6F5108E8275">
<mods:roleTerm id="069378C7E10EE0DAFC8A1B28E8A39532">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA8594B2B5B612A62DD67667B4A1C0AD">Maddison, David R.</mods:namePart>
<mods:nameIdentifier id="BC24DB1F9BAD421E0C4C1E161B59CDB0" type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
<mods:affiliation id="9F86D956E65F2ECE8427E067D7133E07">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
<mods:nameIdentifier id="8BBF07E46B5A082788AFF27437CA57D8" type="email">david.maddison@science.oregonstate.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="180856846393E05E63B4F91AF42E2E27" type="personal">
<mods:role id="8AE220B3476CA4A063956A34D6EBE534">
<mods:roleTerm id="BC8789346701CB025A459F305FC8CEC0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="91F7FA14EE6ED7BF97BF650E0283BF5F">Porch, Nick</mods:namePart>
<mods:affiliation id="9275EDF07C2BB4A20AD77155478C2B5E">School of Life and Environmental Sciences, Deakin University, Geelong 3216, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="DD4836916DA296B8FB68E687B5042EF5">text</mods:typeOfResource>
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<mods:title id="1CB5F183AA42825D2A72F334B1ABBABE">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="AFFB2060D0E7C9885E1BBA624720A320">
<mods:date id="3CCE72FB68F5013ECAE31174EA580FE9">2021</mods:date>
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<mods:number id="D821C765D8150FC80B37E9DB791A6D84">2021-06-16</mods:number>
</mods:detail>
<mods:detail id="9CEF03B36C086B11903080955E1DECA6" type="volume">
<mods:number id="BDBE91BA6D534E40350BA963B06C5B3D">1044</mods:number>
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<mods:url id="273A91E623030D05E20E4FD209DC204F">http://dx.doi.org/10.3897/zookeys.1044.62253</mods:url>
</mods:location>
<mods:classification id="189F9A0FEFCEC4C8F1E17640BFA76B8C">journal article</mods:classification>
<mods:identifier id="42386B69EA0E5BC6504315276BF2583F" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62253</mods:identifier>
<mods:identifier id="971B05D5220CED2DC7D66A0F7B50F9D1" type="Pensoft-Pub">1313-2970-1044-153</mods:identifier>
<mods:identifier id="47275FF108AADB1F85303C0B944E5F23" type="ZooBank">A2219253DB2B4C5992D677595A6C1438</mods:identifier>
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</mods:mods>
<treatment id="BC7D7FFF74C25896A285A134C22A8F60" ID-GBIF-Taxon="182754124" LSID="urn:lsid:zoobank.org:act:5FF236BF-1E87-4480-8798-230E71470E66" httpUri="http://treatment.plazi.org/id/BC7D7FFF74C25896A285A134C22A8F60" lastPageNumber="153" pageId="0" pageNumber="153" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Bembidarenini">
<subSubSection id="B538B13F5FF0226CA8FBE64CE7B8B4EE" pageId="0" pageNumber="153" type="nomenclature">
<paragraph id="3A10D0AD7A4F254614FB984882FE52A4" pageId="0" pageNumber="153">
<taxonomicName id="801FC2E2B4A2E1292D3B40D55ED222DC" ID-CoL="8M9PZ" LSID="http://zoobank.org/5FF236BF-1E87-4480-8798-230E71470E66" authority="Maddison &amp; Porch, 2021" authorityName="Maddison &amp; Porch" authorityYear="2021" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides erwini" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="erwini" status="sp. nov.">Tasmanitachoides erwini</taxonomicName>
<taxonomicNameLabel id="ECCD54E8407ADEF0E8C737056B36FCE8" pageId="0" pageNumber="153">sp. nov.</taxonomicNameLabel>
<figureCitation id="784FA0D846043885AAC82909DE18AB00" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tasmanitachoides erwini sp. nov., adult male, from the type locality. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1044.62253.figure1" httpUri="https://binary.pensoft.net/fig/555123" pageId="0" pageNumber="153">Figures 1</figureCitation>
<figureCitation id="B94DE61EC518FE46A0DC4827D0C04ED5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitats of Tasmanitachoides A Australia: ACT: Murrumbidgee River at Angle Crossing, 35.5825 ° S, 149.1100 ° E, 598 m. This and similar areas a few meters upstream are habitat of T. murrumbidgensis, T. sp. &quot; Tambo R &quot;, T. cf. gerdi, T. sp. &quot; Angle Crossing 1 &quot;, T. wilsoni, T. maior, T. sp. &quot; Angle Crossing 2 &quot;, and T. rufescens B Australia: Victoria: Flat Rock Creek at highway B 23, 37.2835 ° S, 149.2223 ° E, 256 m. Habitat of T. lutus, T. leai, and T. angulicollis C Australia: Tasmania: mouth of Machinery Creek into the River Forth at C 136, 41.4712 ° S, 146.1366 ° E, 126 m. Habitat of T. leai, T. kingi, T. erwini, and T. sp. &quot; River Forth &quot; D Australia: Tasmania: Lake St Clair, 42.1121 ° S, 146.2051 ° E, 741 m. Habitat of T. hobarti." figureDoi="10.3897/zookeys.1044.62253.figure2" httpUri="https://binary.pensoft.net/fig/555124" pageId="0" pageNumber="153">, 2C</figureCitation>
<figureCitation id="2225DB27CEC471A283A3F3B3585C039D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Adults of the Tasmanitachoides wattsensis group and other Tasmanitachoides A T. wattsensis, voucher DNA 5758; NSW: Leatherbarrel Creek u / s Alpine Way B T. sp. &quot; Lerderderg R &quot;, voucher V 101049; VIC: Lerderderg River, 6.8 km N. Bacchus Marsh C T. erwini, voucher V 101469 D T. leai, voucher V 101467 E T. hobarti, voucher V 101463 F T. glabellus, paratype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.62253.figure12" httpUri="https://binary.pensoft.net/fig/555134" pageId="0" pageNumber="153">, 12C</figureCitation>
<figureCitation id="D970BF2F8FD193E62F45912D5E6E8363" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Left elytron of Tasmanitachoides A T. sp. &quot; Lerderderg R &quot;, voucher V 101049 B T. erwini, voucher V 101469 C T. hobarti, voucher V 101463 D T. leai, voucher V 101467 E T. rufescens, voucher V 101478 F T. sp. &quot; River Forth &quot;, voucher DNA 5555 G T. sp. &quot; Angle Crossing 2 &quot;, voucher DNA 5678 H T. flindersianus, paratype J T. bicolor, voucher V 101472 K T. fitzroyi, voucher V 101471 L T. maior, voucher DNA 5508 M T. arnhemensis, voucher V 101476. N T. murrumbidgensis, voucher V 101464 P T. minor, voucher V 101474 Q T. sp. &quot; Tambo R &quot;, voucher V 101465." figureDoi="10.3897/zookeys.1044.62253.figure16" httpUri="https://binary.pensoft.net/fig/555138" pageId="0" pageNumber="153">, 16B</figureCitation>
<figureCitation id="C340B82CD2D346FAB54D23835B1F7526" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dorsal view of head of Tasmanitachoides adults A T. wilsoni, voucher V 101470 B T. baehri, voucher V 101479 C T. lutus, voucher V 101462 D T. cf. gerdi, voucher DNA 5676 E T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 F T. obliquiceps, voucher V 101477 G T. erwini, voucher V 101469 H T. hobarti, voucher V 101463 J T. leai, voucher V 101467 K T. rufescens, voucher V 101478 L T. fitzroyi, voucher V 101471 M T. murrumbidgensis, voucher V 101464." figureDoi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153">, 17G</figureCitation>
<figureCitation id="B59DC2E39BC0FE2986E7ABCEAF65A84D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Elytral microsculpture of Tasmanitachoides, dorsal view A T. baehri, voucher V 101484 B T. erwini, voucher V 101483. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153">, 18B</figureCitation>
<figureCitation id="AD42B2FF4EC921699ACA94EB1BF3DC89" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Aedeagus of Tasmanitachoides, ventral view A T. baehri, voucher DNA 5569 (holotype) B T. wilsoni, voucher DNA 5514 C T. erwini, voucher V 101481 D T. sp. &quot; Lerderderg R &quot;, voucher DNA 2029. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153">, 19C</figureCitation>
<figureCitation id="B5DAE2437C8E68C7F40A60BEBE2ACC67" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagus of Tasmanitachoides, left lateral view A T. baehri, voucher DNA 2032 B T. erwini, voucher DNA 5583 C T. erwini, voucher DNA 5509 (holotype). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure20" httpUri="https://binary.pensoft.net/fig/555142" pageId="0" pageNumber="153">, 20B, C</figureCitation>
<figureCitation id="9F3D373CDE04C0E4798F2B113072F020" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Known geographic distributions of Tasmanitachoides baehri and T. erwini in southeastern Australia. Circles: T. baehri; stars: T. erwini." figureDoi="10.3897/zookeys.1044.62253.figure21" httpUri="https://binary.pensoft.net/fig/555143" pageId="0" pageNumber="153">, 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87761CF74292BCADA00B5B46E7B5D571" pageId="0" pageNumber="153" type="materials_examined">
<paragraph id="B788DEB1334433AF02860B5BABF0F882" pageId="0" pageNumber="153">Material examined.</paragraph>
<paragraph id="4F499B7ED6256FC6A7F63FFA1601C127" pageId="0" pageNumber="153">
<materialsCitation id="B4FD4FA63D20C9C641D66DA9103731A6" accessionNumber="MW291170, MW291262, MW291215, MW291234, MW291305" collectingDate="2019-01-14" collectorName="D. R. Maddison, N. A. Porch, David R. Maddison" country="Australia" elevation="126" latitude="-41.4712" location="Male" longLatPrecision="6" longitude="146.1366" specimenCount="1" stateProvince="Tasmania" typeStatus="Holotype">
<emphasis id="C2B6AF5C2D0337F95E750A885133A70C" bold="true" pageId="0" pageNumber="153">
<emphasis id="39A40035234919BA46FE137CFB2B8685" italics="true" pageId="0" pageNumber="153">
<typeStatus id="FE65466FBF53BD61B113E3C6A5E1603F">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<location id="DAA4938815AB4ED64F1CF511FA3CF758" LSID="urn:lsid:plazi:treatment:BC7D7FFF74C25896A285A134C22A8F60:DAA4938815AB4ED64F1CF511FA3CF758" country="Australia" latitude="-41.4712" longLatPrecision="6" longitude="146.1366" name="Male" stateProvince="Tasmania">Male</location>
(ANIC), labeled: &quot;
<collectingCountry id="99CB0FA14D12DD6EBD5AD0EE7BEFB501" name="Australia">Australia</collectingCountry>
:
<collectingRegion id="46A519FEE6003421D38291F4801B3031" country="Australia" name="Tasmania">Tasmania</collectingRegion>
: River Forth at C136,
<geoCoordinate id="9053774EA1372B4FB42E029106315AC0" degrees="41.4712" direction="south" orientation="latitude" precision="5" value="-41.4712">41.4712°S</geoCoordinate>
<geoCoordinate id="DAAF534BE8D64610E4BEAAC9B704674C" degrees="146.1366" direction="east" orientation="longitude" precision="5" value="146.1366">146.1366°E</geoCoordinate>
,
<quantity id="459186082EA519AB274304DDCC445B39" metricMagnitude="2" metricUnit="m" metricValue="1.26" unit="m" value="126.0">
<elevation id="9E69FF58BBB0564C0FFCBBE7DE4924DA" metricMagnitude="2" metricUnit="m" metricValue="1.26" unit="m" value="126.0">126 m</elevation>
</quantity>
,
<collectingDate id="F50AC8A8AF48F838438CA78D12BC80C7" value="2019-01-14">14.i.2019</collectingDate>
. DRM 19.012.
<collectorName id="70FCBBB6094DA046DCFB7AE8131A8171">D.R. Maddison</collectorName>
&amp;
<collectorName id="47685ADA37D793AB6E03CDA8584F5E17">N.A. Porch</collectorName>
&quot;, &quot;
<collectorName id="764EC788F7CC04DC1708B8B94E62A98B">David R. Maddison</collectorName>
DNA5509 DNA Voucher&quot; [pale green paper], &quot;
<typeStatus id="AFD0CB04F9327357A1FF747231D036F0">HOLOTYPE</typeStatus>
<taxonomicName id="18DA4C32AE8DF29BD5DF1E59860D0C3C" authorityName="Maddison &amp; Porch" authorityYear="2021" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides erwini" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="erwini">Tasmanitachoides erwini</taxonomicName>
Maddison &amp; Porch&quot; [partly handwritten, on red paper]. Genitalia mounted in Euparal on coverslip pinned with specimen; extracted DNA stored separately. GenBank accession numbers for DNA sequences of the
<typeStatus id="E98C78EF3A1688E1D38D6E026B2AD08D">holotype</typeStatus>
are
<accessionNumber id="3ED22F5580EC62FB4233BDE497EA6384" isEnumeration="true">MW291170, MW291262, MW291215, MW291234</accessionNumber>
, and
<accessionNumber id="C97C9EA74954EF100B748E6BC6AFBC46" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW291305">MW291305</accessionNumber>
.
</materialsCitation>
</paragraph>
<paragraph id="17269E5C2F7505BD2979CA9BE79CB5E0" pageId="0" pageNumber="153">
<materialsCitation id="C5614BADD6D3C31E40AE0CD56825E083" accessionNumber="MW291170, MW291262, MW291215, MW291234, MW291305" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="L. StClaire-Queenstown Jan., Mersey R Vy. Mar." country="Australia" county="Queenstown" elevation="126" latitude="-41.532" location="Same" longLatPrecision="69" longitude="146.426" municipality="King River" specimenCount="1" stateProvince="Tasmania" typeStatus="Paratypes">
<emphasis id="230D07FEF29BF969D02CBA8C4E5DE534" bold="true" italics="true" pageId="0" pageNumber="153">
<typeStatus id="FFCA715AF9244F1D5A44A7B39686909A">Paratypes</typeStatus>
</emphasis>
(23).
<location id="05D5F14D7879742CFFE4AC9913F7F1E2" LSID="urn:lsid:plazi:treatment:BC7D7FFF74C25896A285A134C22A8F60:05D5F14D7879742CFFE4AC9913F7F1E2" country="Australia" county="Queenstown" latitude="-41.532" longLatPrecision="69" longitude="146.426" municipality="King River" name="Same" stateProvince="Tasmania">Same</location>
label data as holotype (20; ANIC, OSAC, NPC, ZSM, NMV, QVMAG, NHMUK, TMAG, USNM).
<location id="BE1A6F4573C2175E1EF1A2EE68AD872B" LSID="urn:lsid:plazi:treatment:BC7D7FFF74C25896A285A134C22A8F60:BE1A6F4573C2175E1EF1A2EE68AD872B" country="Australia" county="Queenstown" latitude="-41.532" longLatPrecision="69" longitude="146.426" municipality="King River" name="In" stateProvince="Tasmania">In</location>
addition to these, we have seen three
<typeStatus id="7E5451583A48F24D5C5FA1858E09DC69" isOtherTypeStatus="true">additional specimens</typeStatus>
, all in the MCZ, which we have designated as
<typeStatus id="16B1D4B4F7CFE307F240F2ABE58DA491">paratypes</typeStatus>
.
<location id="8E2D9921B4C85AFD5D63E24AB3235B94" LSID="urn:lsid:plazi:treatment:BC7D7FFF74C25896A285A134C22A8F60:8E2D9921B4C85AFD5D63E24AB3235B94" country="Australia" county="Queenstown" latitude="-41.532" longLatPrecision="69" longitude="146.426" municipality="King River" name="Two" stateProvince="Tasmania">Two</location>
are a labeled &quot;
<collectorName id="3AD50F8E8C197F559CCBA1D53DDF41E1">L. StClaire-Queenstown Jan.</collectorName>
'57 Tas Darlingtons&quot; &quot;
<taxonomicName id="24BB78FF60FC9D30F3824E4ACBF46B61" class="Insecta" family="Carabidae" genus="Tachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachys" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">Tachys</taxonomicName>
<taxonomicName id="FC43F82D934F8EA8D77CA172FB6B6918" baseAuthorityName="Blackburn" baseAuthorityYear="1901" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides hobarti" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="hobarti">Tasmanitachoides hobarti</taxonomicName>
(Sl.) det Darl.
<normalizedToken id="5F0FF77EB971DD90A4A80D4521A6FD08" originalValue="‘61”">'61&quot;</normalizedToken>
; according to
<bibRefCitation id="155150B9B034AC8C2185CAAE48A6A382" DOI="https://doi.org/10.1155/1962/71454" author="Darlington, Jr PJ" journalOrPublisher="Psyche" pageId="0" pageNumber="153" pagination="117 - 128" refId="B11" refString="Darlington, Jr PJ, 1962. Australian carabid beetles XI. some Tachys. Psyche 69: 117 - 128, DOI: https://doi.org/10.1155/1962/71454" title="Australian carabid beetles XI. some Tachys." url="https://doi.org/10.1155/1962/71454" volume="69" year="1962">Darlington (1962</bibRefCitation>
:117) these two specimens are from the crossing of the
<collectingMunicipality id="D2F37D33536F41A0D9CE60DAF0D969BB">King River</collectingMunicipality>
by the
<collectingCounty id="0C807961C2F30E27E93BB618D106CA51">Queenstown</collectingCounty>
road, which at the time (before the Crotty Dam) would have been approximately
<geoCoordinate id="F0126CD1AEEEBFBB96ECE9813BDDB7D9" degrees="42.074" direction="south" orientation="latitude" precision="55" value="-42.074">42.074°S</geoCoordinate>
<geoCoordinate id="BE8D24083D9AE90798B023437F515D7C" degrees="145.652" direction="east" orientation="longitude" precision="55" value="145.652">145.652°E</geoCoordinate>
. The third is labeled &quot;
<collectorName id="9B4B539D01C89790A967E17A00B4598E">Mersey R Vy. Mar.</collectorName>
'57 Tas Darlingtons&quot; &quot;
<taxonomicName id="0FC9CB25D4E42AA8723F09D234C4C69F" class="Insecta" family="Carabidae" genus="Tachys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachys" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">Tachys</taxonomicName>
<taxonomicName id="C4706BA2C0D43E640417C083F8D4BF25" baseAuthorityName="Blackburn" baseAuthorityYear="1901" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides hobarti" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="hobarti">Tasmanitachoides hobarti</taxonomicName>
(Sl.) det Darl.
<normalizedToken id="CA4978BDD9FD5C0BFFC028E361E98611" originalValue="‘61”">'61&quot;</normalizedToken>
. According to the map in
<bibRefCitation id="1B386027A98106C555EBAA67FC470360" DOI="https://doi.org/10.1155/1960/81583" author="Darlington, Jr PJ" journalOrPublisher="Psyche" pageId="0" pageNumber="153" pagination="111 - 126" refId="B10" refString="Darlington, Jr PJ, 1960. Australian carabid beetles IV. List of localities, 1956-1958. Psyche 67: 111 - 126, DOI: https://doi.org/10.1155/1960/81583" title="Australian carabid beetles IV. List of localities, 1956 - 1958." url="https://doi.org/10.1155/1960/81583" volume="67" year="1960">Darlington (1960)</bibRefCitation>
, this locality is at approximately
<geoCoordinate id="E92693CCAB12DE5C73085C4E282C6078" degrees="41.532" direction="south" orientation="latitude" precision="55" value="-41.532">41.532°S</geoCoordinate>
,
<geoCoordinate id="DF36D65D4B93E0CED25994E997F5DA04" degrees="146.426" direction="east" orientation="longitude" precision="55" value="146.426">146.426°E</geoCoordinate>
. These specimens formed
<normalizedToken id="CAE80B2E94595C7B478384A4A520F68F" originalValue="Darlington’s">Darlington's</normalizedToken>
concept of
<taxonomicName id="30C59A0FC01998664FE0EABB78B7D29E" baseAuthorityName="Blackburn" baseAuthorityYear="1901" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides hobarti" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="hobarti">
<emphasis id="38D1D74289966EF21E11223D97333A5B" italics="true" pageId="0" pageNumber="153">Tasmanitachoides hobarti</emphasis>
</taxonomicName>
. They also are specimens studied and figured by
<bibRefCitation id="5D79146EB88276563BEAC47097AA6CF9" author="Erwin, TL" journalOrPublisher="Breviora" pageId="0" pageNumber="153" pagination="1 - 19" publicationUrl="https://www.biodiversitylibrary.org/page/3664881page/87/mode/1up" refId="B12" refString="Erwin, TL, 1972. Two new genera of bembidiine carabid beetles from Australia and South America with notes on their phylogenetic and zoogeographic significance (Coleoptera). Breviora 383: 1 - 19, https://www.biodiversitylibrary.org/page/3664881page/87/mode/1up" title="Two new genera of bembidiine carabid beetles from Australia and South America with notes on their phylogenetic and zoogeographic significance (Coleoptera)." url="https://www.biodiversitylibrary.org/page/3664881page/87/mode/1up" volume="383" year="1972">Erwin (1972)</bibRefCitation>
as
<taxonomicName id="E3082EF28E889FFC01749074DFB7CD26" lsidName="T. hobarti" pageId="0" pageNumber="153" rank="species" species="hobarti">
<emphasis id="1C6E9F78F0A3858E6E5892DACCB93812" italics="true" pageId="0" pageNumber="153">T. hobarti</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C555C5FEE6D513EDCD184FC2EFE0CD60" pageId="0" pageNumber="153" type="type locality">
<paragraph id="CEA1B24B49015BAA68D1875067B73A20" pageId="0" pageNumber="153">Type locality.</paragraph>
<paragraph id="20E02BDCB1F5C8BBA3EB8806EAC6B801" pageId="0" pageNumber="153">
Australia: Tasmania: at the mouth of Machinery Creek into the River Forth at road C136,
<geoCoordinate id="014B7E8C05E86EED425ADB303932EAF7" degrees="41.4712" direction="south" orientation="latitude" precision="5" value="-41.4712">41.4712°S</geoCoordinate>
,
<geoCoordinate id="261D32CF75EAFB47F61B3B6C2BE521D0" degrees="146.1366" direction="east" orientation="longitude" precision="5" value="146.1366">146.1366°E</geoCoordinate>
, 126 m.
</paragraph>
</subSubSection>
<subSubSection id="615A4ED682C915C52BE270BC57E378BC" pageId="0" pageNumber="153" type="derivation of specific epithet">
<paragraph id="914CACA3D6E446FE01F7EE287EBA8E16" pageId="0" pageNumber="153">Derivation of specific epithet.</paragraph>
<paragraph id="85F9FAEFD0642CC9C9C1E6E18D85E10F" pageId="0" pageNumber="153">
We are honored to name this species after the late Terry Lee Erwin, for his many contributions to carabidology and systematics in general, and to our knowledge of
<taxonomicName id="D315392F3FC0B75AEE6B9C5DC3085F88" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="B4952E17884F4DC28706ABB6D268331C" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
and other bembidarenines in particular.
</paragraph>
</subSubSection>
<subSubSection id="9C1C1F7AAE38F70ED979D14905E9791B" pageId="0" pageNumber="153" type="diagnosis">
<paragraph id="4DEAC147C2792B40DA8E83A59A30104B" pageId="0" pageNumber="153">Diagnosis and description.</paragraph>
<paragraph id="A09B9ED2FDB47E157D3F1CBEC8CF6E1C" pageId="0" pageNumber="153">
Length 2.25-2.75 mm (n = 7); most specimens less than 2.6 mm. One of the darker species of
<taxonomicName id="0D768CDB9BAE8548A52E524195A45FF5" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="461BC97227E85ACAC9C32C60ECFB3435" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="733E65BCFBE924B1BFA1D0B9ABA88C91" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tasmanitachoides erwini sp. nov., adult male, from the type locality. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1044.62253.figure1" httpUri="https://binary.pensoft.net/fig/555123" pageId="0" pageNumber="153">1</figureCitation>
): body piceous to black; appendages piceous, including basal antennomeres, with the exception of the tarsi, which are slightly paler. Body relatively flat and parallel-sided; elytra narrowing posteriorly, and thus more pointed than other species. Head without tubercles at anterior corners of clypeus, and without concave region in anterior half. Frontal furrows (Fig.
<figureCitation id="7355C7636BCAC3F81E87C4EAD230B261" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dorsal view of head of Tasmanitachoides adults A T. wilsoni, voucher V 101470 B T. baehri, voucher V 101479 C T. lutus, voucher V 101462 D T. cf. gerdi, voucher DNA 5676 E T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 F T. obliquiceps, voucher V 101477 G T. erwini, voucher V 101469 H T. hobarti, voucher V 101463 J T. leai, voucher V 101467 K T. rufescens, voucher V 101478 L T. fitzroyi, voucher V 101471 M T. murrumbidgensis, voucher V 101464." figureDoi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153">17G</figureCitation>
) more or less straight, reaching backward to approximately the center of the eye, parallel or slightly diverging posteriorly; bottom of furrows rugose. Pronotum relatively narrow (Fig.
<figureCitation id="42B083517FF7F37CEC06C806928F7474" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Adults of the Tasmanitachoides wattsensis group and other Tasmanitachoides A T. wattsensis, voucher DNA 5758; NSW: Leatherbarrel Creek u / s Alpine Way B T. sp. &quot; Lerderderg R &quot;, voucher V 101049; VIC: Lerderderg River, 6.8 km N. Bacchus Marsh C T. erwini, voucher V 101469 D T. leai, voucher V 101467 E T. hobarti, voucher V 101463 F T. glabellus, paratype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.62253.figure12" httpUri="https://binary.pensoft.net/fig/555134" pageId="0" pageNumber="153">12C</figureCitation>
), slightly sinuate laterally in front of the right or slightly acute hind angle. First stria abruptly sinuate, very close to the suture in the anterior fifth or fourth, at which point it abruptly bends away from the suture. Striae 3 and 4 very weak, almost absent in some specimens; the striae 3 and 4 are joined at the anterior discal seta (ed3; Fig.
<figureCitation id="F679BE4F84CEB6FD8752DE8C422641CD" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Left elytron of Tasmanitachoides A T. sp. &quot; Lerderderg R &quot;, voucher V 101049 B T. erwini, voucher V 101469 C T. hobarti, voucher V 101463 D T. leai, voucher V 101467 E T. rufescens, voucher V 101478 F T. sp. &quot; River Forth &quot;, voucher DNA 5555 G T. sp. &quot; Angle Crossing 2 &quot;, voucher DNA 5678 H T. flindersianus, paratype J T. bicolor, voucher V 101472 K T. fitzroyi, voucher V 101471 L T. maior, voucher DNA 5508 M T. arnhemensis, voucher V 101476. N T. murrumbidgensis, voucher V 101464 P T. minor, voucher V 101474 Q T. sp. &quot; Tambo R &quot;, voucher V 101465." figureDoi="10.3897/zookeys.1044.62253.figure16" httpUri="https://binary.pensoft.net/fig/555138" pageId="0" pageNumber="153">16B</figureCitation>
), and in most specimens are merged in front of that point. Stria 5 distinctly engraved throughout the entire anterior half; in posterior half it gradually weakens toward the rear. Stria 6 consisting of a few isolated punctures; stria 7 absent. Discal setae ed6 in stria 3. Microsculpture weak, sculpticells weakly engraved, and thus the surface is shiny; sculpticells isodiametric on head and pronotum, slightly longitudinally stretched on elytra (Fig.
<figureCitation id="A051C6B38480D4672E704220EBA2008E" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Elytral microsculpture of Tasmanitachoides, dorsal view A T. baehri, voucher V 101484 B T. erwini, voucher V 101483. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153">18B</figureCitation>
). Aedeagus (Figs
<figureCitation id="02D17DFC4B39CA0BD29251B86F38C4A7" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Aedeagus of Tasmanitachoides, ventral view A T. baehri, voucher DNA 5569 (holotype) B T. wilsoni, voucher DNA 5514 C T. erwini, voucher V 101481 D T. sp. &quot; Lerderderg R &quot;, voucher DNA 2029. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153">19C</figureCitation>
,
<figureCitation id="A036F1128D49F3A0E98FFCCE09DEFCBF" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagus of Tasmanitachoides, left lateral view A T. baehri, voucher DNA 2032 B T. erwini, voucher DNA 5583 C T. erwini, voucher DNA 5509 (holotype). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure20" httpUri="https://binary.pensoft.net/fig/555142" pageId="0" pageNumber="153">20B, C</figureCitation>
) with internal sac sclerites elongated and relatively straight, very similar to those of
<taxonomicName id="3AE828E5C9073B16000560F71A217C0E" genus="T." lsidName="T." pageId="0" pageNumber="153" rank="genus">
<emphasis id="1D91F930685856CE58C2CB41BBB52EAF" italics="true" pageId="0" pageNumber="153">T.</emphasis>
</taxonomicName>
sp. &quot;Lerderderg R&quot; (Fig.
<figureCitation id="8C75686965F0C2453E26247EE450FB97" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Aedeagus of Tasmanitachoides, ventral view A T. baehri, voucher DNA 5569 (holotype) B T. wilsoni, voucher DNA 5514 C T. erwini, voucher V 101481 D T. sp. &quot; Lerderderg R &quot;, voucher DNA 2029. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153">19D</figureCitation>
). Ventral surface of the aedeagus quite straight (Fig.
<figureCitation id="B16ACE8E1F840A87EE4823D19E87C442" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagus of Tasmanitachoides, left lateral view A T. baehri, voucher DNA 2032 B T. erwini, voucher DNA 5583 C T. erwini, voucher DNA 5509 (holotype). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure20" httpUri="https://binary.pensoft.net/fig/555142" pageId="0" pageNumber="153">20B, C</figureCitation>
).
</paragraph>
<caption id="FA51768D990DD056B1C721CB4985ACEF" doi="10.3897/zookeys.1044.62253.figure20" httpUri="https://binary.pensoft.net/fig/555142" pageId="0" pageNumber="153" start="Figure 20" startId="F20">
<paragraph id="2E9CD27E700FFE6DED1863B36E8C592A" pageId="0" pageNumber="153">
<emphasis id="BF212ACD3262B842B0573F640B61F2A1" bold="true" pageId="0" pageNumber="153">Figure 20.</emphasis>
Aedeagus of
<taxonomicName id="E34AEE4753B90C130EA24C6E9C0282E7" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="9228054F38B50A296BD2F3584BE6DAF7" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
, left lateral view
<emphasis id="5973C8FB2854F95D474061F50F7F91B3" bold="true" pageId="0" pageNumber="153">A</emphasis>
<taxonomicName id="E9D8663BF14D46D754BC9323680A8AFC" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="F5EFA5ED01F4CBD7A15E91CDE8C0DD46" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
, voucher DNA2032
<emphasis id="F06478D131475ABE489E6DFEF91A633E" bold="true" pageId="0" pageNumber="153">B</emphasis>
<taxonomicName id="DE6EECF6B1989BDC72778E01835CCD58" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="31F6132309C710D3168674EA82476C9D" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
, voucher DNA5583
<emphasis id="48E1C132F36D329B3B04682E1D267C48" bold="true" pageId="0" pageNumber="153">C</emphasis>
<taxonomicName id="0AE385055309490DDDF5E09174873F2B" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="FE602AB11B5F407C0CC7FCF3A8BFEC6B" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
, voucher DNA5509 (holotype). Scale bar: 100
<normalizedToken id="1A4BE50DCC3F0D41688199FC5FE18715" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="636EB35ED4A82CA990480C9BAB11982E" pageId="0" pageNumber="153" type="comparisons">
<paragraph id="17A1E6EA400630456D9A67978CA6C764" pageId="0" pageNumber="153">Comparison with related species.</paragraph>
<paragraph id="C08B5BAF0233CF7225613CF2F5FFD57D" pageId="0" pageNumber="153">
As with other members of the
<taxonomicName id="BA9346640558C95B73EDD96E72B683C1" baseAuthorityName="Blackburn" baseAuthorityYear="1901" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides wattsensis" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="wattsensis">
<emphasis id="E351012E3E2C96333BFA37E707B60FCB" italics="true" pageId="0" pageNumber="153">Tasmanitachoides wattsensis</emphasis>
</taxonomicName>
group, this species has a relatively unmodified clypeus, without anterior lateral tubercles, and with the third and fourth elytral striae nearly effaced. Its darker color (including the entirely piceous antenna) and flatness distinguish it from other members of the group. It is the only known species of the group from Tasmania. From the other two large and dark
<taxonomicName id="8B72038CAD1F76961F15A97D430BF4A0" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="47DBBBF2C25D9FFC83EC3BE93685DF1A" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
from Tasmania,
<taxonomicName id="F9C0A5BBD80FEAD986498BF71C9B8DF6" lsidName="T. hobarti" pageId="0" pageNumber="153" rank="species" species="hobarti">
<emphasis id="BDD615A53439453718AD402FB0312FB5" italics="true" pageId="0" pageNumber="153">T. hobarti</emphasis>
</taxonomicName>
and
<taxonomicName id="8254AFCABA7F822D49D59B8ECFD36316" lsidName="T. leai" pageId="0" pageNumber="153" rank="species" species="leai">
<emphasis id="5F37DF8F0959FBF036CFF92C14E796B2" italics="true" pageId="0" pageNumber="153">T. leai</emphasis>
</taxonomicName>
,
<taxonomicName id="3E045363A8A0E3CEE781D9DD52034E34" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="F39B445FCAE7AA33E554D6CD03165908" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
is distinguished by having a darker antennomere 1 and flatter body. From
<taxonomicName id="DA1869690DAD7231E25528FC70AB12DA" lsidName="T. hobarti" pageId="0" pageNumber="153" rank="species" species="hobarti">
<emphasis id="C0F8A8A41240D4728E53BDAD4B7B72C4" italics="true" pageId="0" pageNumber="153">T. hobarti</emphasis>
</taxonomicName>
it is further distinguished by the much weaker striae 3 and 4; from
<taxonomicName id="DB72D6F65AB6439318507A2B65A9A027" lsidName="T. leai" pageId="0" pageNumber="153" rank="species" species="leai">
<emphasis id="A35BF305489BDB67F5492406E1EE4E5E" italics="true" pageId="0" pageNumber="153">T. leai</emphasis>
</taxonomicName>
by the longer stria 5.
</paragraph>
</subSubSection>
<subSubSection id="2E659C9F795F34FBAEBA8DAB42D16247" pageId="0" pageNumber="153" type="distribution">
<paragraph id="AE538EF71139CE24CC480F09B8E8159C" pageId="0" pageNumber="153">Geographic distribution.</paragraph>
<paragraph id="78BDD96FF1B91238399F5918DD59AFA6" pageId="0" pageNumber="153">
Only known from northwestern Tasmania (Fig.
<figureCitation id="1286A335B6F300D2AD5A5EAA1EB6CEFE" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Known geographic distributions of Tasmanitachoides baehri and T. erwini in southeastern Australia. Circles: T. baehri; stars: T. erwini." figureDoi="10.3897/zookeys.1044.62253.figure21" httpUri="https://binary.pensoft.net/fig/555143" pageId="0" pageNumber="153">21</figureCitation>
).
</paragraph>
<caption id="BF1C68C37EADF12F48CBDFCCE314F4E7" doi="10.3897/zookeys.1044.62253.figure21" httpUri="https://binary.pensoft.net/fig/555143" pageId="0" pageNumber="153" start="Figure 21" startId="F21">
<paragraph id="B7CFF659B572623010D58D0379805B70" pageId="0" pageNumber="153">
<emphasis id="5339A9640338B13881591A1CE423D106" bold="true" pageId="0" pageNumber="153">Figure 21.</emphasis>
Known geographic distributions of
<taxonomicName id="CAE92345878A220C1D14C8FF47C854D5" authorityName="Maddison &amp; Porch" authorityYear="2021" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides baehri" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="baehri">
<emphasis id="4FCE9A8313663574479540F41EB17991" italics="true" pageId="0" pageNumber="153">Tasmanitachoides baehri</emphasis>
</taxonomicName>
and
<taxonomicName id="852347F5F9866184E2DF881AEE559C06" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="CA98510CF9DF960008056F03336228B6" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
in southeastern Australia. Circles:
<taxonomicName id="5B866ED6AAB76862DB2C404D162582B3" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="19F6D0E6DEFD11F0843A129D44B0F3AA" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
; stars:
<taxonomicName id="C93CA18D7736C335BD25AC0F5AFCBD35" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="8AA09D13BDBBB64D80728D49CC95FC78" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1038B6939F80C95972FF75108E4C5C45" pageId="0" pageNumber="153" type="habitat">
<paragraph id="6711B74B30396A364BC6E447F6D0CE18" pageId="0" pageNumber="153">Habitat.</paragraph>
<paragraph id="1F0AE5243C92A144736AFE2507C082C3" pageId="0" pageNumber="153">
At the type locality, members of this species were found during daylight hours in fine gravel on the banks of Mineral Creek at its mouth into the River Forth (Fig.
<figureCitation id="72E9623015D3EBF09A17B043E42EA712" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitats of Tasmanitachoides A Australia: ACT: Murrumbidgee River at Angle Crossing, 35.5825 ° S, 149.1100 ° E, 598 m. This and similar areas a few meters upstream are habitat of T. murrumbidgensis, T. sp. &quot; Tambo R &quot;, T. cf. gerdi, T. sp. &quot; Angle Crossing 1 &quot;, T. wilsoni, T. maior, T. sp. &quot; Angle Crossing 2 &quot;, and T. rufescens B Australia: Victoria: Flat Rock Creek at highway B 23, 37.2835 ° S, 149.2223 ° E, 256 m. Habitat of T. lutus, T. leai, and T. angulicollis C Australia: Tasmania: mouth of Machinery Creek into the River Forth at C 136, 41.4712 ° S, 146.1366 ° E, 126 m. Habitat of T. leai, T. kingi, T. erwini, and T. sp. &quot; River Forth &quot; D Australia: Tasmania: Lake St Clair, 42.1121 ° S, 146.2051 ° E, 741 m. Habitat of T. hobarti." figureDoi="10.3897/zookeys.1044.62253.figure2" httpUri="https://binary.pensoft.net/fig/555124" pageId="0" pageNumber="153">2C</figureCitation>
); specimens were found after splashing the gravel with water. The banks had no visible vegetation. Present in the same habitat were
<taxonomicName id="EBF4582EE55FA5329C81FF500E45FEA5" baseAuthorityName="Sloane" baseAuthorityYear="1896" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides leai" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="leai">
<emphasis id="FF8936819535033960AA7631CE84FC96" italics="true" pageId="0" pageNumber="153">Tasmanitachoides leai</emphasis>
</taxonomicName>
,
<taxonomicName id="9678B5963F072C7956E1B4EC1E16646A" lsidName="T. kingi" pageId="0" pageNumber="153" rank="species" species="kingi">
<emphasis id="11CBFD2EA15163A1921199DE8C8FDD9F" italics="true" pageId="0" pageNumber="153">T. kingi</emphasis>
</taxonomicName>
, and
<taxonomicName id="7F5B777F699F06C36D28873D098A73C9" genus="T." lsidName="T." pageId="0" pageNumber="153" rank="genus">
<emphasis id="30E7F63A12F32479A20C71F53C9B498A" italics="true" pageId="0" pageNumber="153">T.</emphasis>
</taxonomicName>
sp. &quot;River Forth&quot;.
</paragraph>
</subSubSection>
<subSubSection id="A3BE8041407259632C7229933521DC0D" pageId="0" pageNumber="153" type="phylogenetic relationships">
<paragraph id="8CB471AED1EA498D650DA9BB82DC74BF" pageId="0" pageNumber="153">Phylogenetic relationships.</paragraph>
<paragraph id="55005EE011F6A41CCDF6F0C749E69F7E" pageId="0" pageNumber="153">
This species belongs to the
<taxonomicName id="8CECA7D711A494F19F9447748945DDA1" baseAuthorityName="Blackburn" baseAuthorityYear="1901" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides wattsensis" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="wattsensis">
<emphasis id="DA347200A280A4660B0A40EEDA207CDE" italics="true" pageId="0" pageNumber="153">Tasmanitachoides wattsensis</emphasis>
</taxonomicName>
species group, and appears to be the sister to
<taxonomicName id="2E24B837A8CF1099EC31EB5DB4F6E20A" genus="T." lsidName="T." pageId="0" pageNumber="153" rank="genus">
<emphasis id="C20FC8C6BA2A6BFBF529065E756F311F" italics="true" pageId="0" pageNumber="153">T.</emphasis>
</taxonomicName>
sp. &quot;Lerderderg R&quot; among the sampled species (Figs
<figureCitation id="B081DFCF71FC434710388433A01726D7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Maximum likelihood tree for the concatenated matrix of all gene fragments (main figure) and for the concatenated matrix of all genes except COI (inset; this shows only part of the tree). Branch length is shown proportional to relative divergence, as estimated by IQ-TREE; scale bar indicates 0.01 units. Outgroups (Bembidarenas, Argentinatochoides, and Andinodontis; Table 1) not depicted." figureDoi="10.3897/zookeys.1044.62253.figure5" httpUri="https://binary.pensoft.net/fig/555127" pageId="0" pageNumber="153">5</figureCitation>
-
<figureCitation id="F1EE830D9A2EDF7A937E276B14A34581" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Maximum likelihood trees for wg, CAD 4, and CAD 2. Branch length is shown proportional to relative divergence, as estimated by IQ-TREE; scale bars indicate 0.01 units. Outgroups not depicted." figureDoi="10.3897/zookeys.1044.62253.figure9" httpUri="https://binary.pensoft.net/fig/555131" pageId="0" pageNumber="153">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="345CE1DB7E30E8CA6600CDAD6B9B2633" pageId="0" pageNumber="153" type="notes">
<paragraph id="BE44901C792B014343DAA8178CAC4A1E" pageId="0" pageNumber="153">Notes.</paragraph>
<paragraph id="D5091348D2177C0D996760B10F70B193" pageId="0" pageNumber="153">
This is the species illustrated by
<bibRefCitation id="AECCEC2D7257524D3A306D28F7AE7D46" author="Erwin, TL" journalOrPublisher="Breviora" pageId="0" pageNumber="153" pagination="1 - 19" publicationUrl="https://www.biodiversitylibrary.org/page/3664881page/87/mode/1up" refId="B12" refString="Erwin, TL, 1972. Two new genera of bembidiine carabid beetles from Australia and South America with notes on their phylogenetic and zoogeographic significance (Coleoptera). Breviora 383: 1 - 19, https://www.biodiversitylibrary.org/page/3664881page/87/mode/1up" title="Two new genera of bembidiine carabid beetles from Australia and South America with notes on their phylogenetic and zoogeographic significance (Coleoptera)." url="https://www.biodiversitylibrary.org/page/3664881page/87/mode/1up" volume="383" year="1972">Erwin (1972)</bibRefCitation>
as
<taxonomicName id="4226D44C88569F0BB9FAECB426788DC9" lsidName="T. hobarti" pageId="0" pageNumber="153" rank="species" species="hobarti">
<emphasis id="C5592FD03B6319D7EBADFF6C58835224" italics="true" pageId="0" pageNumber="153">T. hobarti</emphasis>
</taxonomicName>
. This is evident both by the localities of the specimens he examined (as the localities match
<normalizedToken id="6622FBAFB9082A7E5E31A4DD984C03D1" originalValue="Darlington’s">Darlington's</normalizedToken>
), and because of the figures themselves, including the features of the genitalia, which match those of this species rather than
<taxonomicName id="E917F9C7AB0A1A7661DDCDC439F008B5" lsidName="T. hobarti" pageId="0" pageNumber="153" rank="species" species="hobarti">
<emphasis id="C3B3488FB4381D6328F89EC392E1430D" italics="true" pageId="0" pageNumber="153">T. hobarti</emphasis>
</taxonomicName>
. The male genitalia of &quot;
<taxonomicName id="3D00922D52D55388DCA386341E9D6AE2" baseAuthorityName="Blackburn" baseAuthorityYear="1901" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides hobarti" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="hobarti">
<emphasis id="3BD6A3D8B08684CE91D95075F2B7A252" italics="true" pageId="0" pageNumber="153">Tasmanitachoides hobarti</emphasis>
</taxonomicName>
&quot; figured in
<bibRefCitation id="7A06B58B06F7109C969FFE5EC6CAC990" DOI="https://doi.org/10.1071/IT9900867" author="Baehr, M" journalOrPublisher="Invertebrate Taxonomy" pageId="0" pageNumber="153" pagination="867 - 894" refId="B1" refString="Baehr, M, 1990. Revision of the Australian ground-beetle genus Tasmanitachoides Erwin (Insecta: Coleoptera: Carabidae: Bembidiinae), with special regard to the tropical species. Invertebrate Taxonomy 4: 867 - 894, DOI: https://doi.org/10.1071/IT9900867" title="Revision of the Australian ground-beetle genus Tasmanitachoides Erwin (Insecta: Coleoptera: Carabidae: Bembidiinae), with special regard to the tropical species." url="https://doi.org/10.1071/IT9900867" volume="4" year="1990">Baehr (1990</bibRefCitation>
: Fig.
<figureCitation id="9E874F40FA780DCD14B56B6C96719C93" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Adults of the Tasmanitachoides wattsensis group and other Tasmanitachoides A T. wattsensis, voucher DNA 5758; NSW: Leatherbarrel Creek u / s Alpine Way B T. sp. &quot; Lerderderg R &quot;, voucher V 101049; VIC: Lerderderg River, 6.8 km N. Bacchus Marsh C T. erwini, voucher V 101469 D T. leai, voucher V 101467 E T. hobarti, voucher V 101463 F T. glabellus, paratype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.62253.figure12" httpUri="https://binary.pensoft.net/fig/555134" pageId="0" pageNumber="153">12</figureCitation>
) is of this species as well. Some specimens from the type series (collected 14 January 2019) are teneral.
</paragraph>
</subSubSection>
</treatment>
</document>
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