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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.906.46438" ID-GBIF-Dataset="ba71138c-80e8-4bc0-a9e8-25927a0b3818" ID-PMC="PMC6989621" ID-Pensoft-Pub="1313-2970-906-41" ID-Pensoft-UUID="F711F888C51E5B7696DD6DFAC353BE28" ID-PubMed="32021556" ID-ZooBank="35E0E0266C394DF78C393EA9DD1F9440" ModsDocID="1313-2970-906-41" checkinTime="1579803474772" checkinUser="pensoft" docAuthor="Ma, Yunlong, Yang, Xingke &amp; Liu, Xingyue" docDate="2020" docId="73095F4DA45B5F21B35551606C3164D8" docLanguage="en" docName="ZooKeys 906: 41-71" docOrigin="ZooKeys 906" docSource="http://dx.doi.org/10.3897/zookeys.906.46438" docTitle="Ankylopteryx yangi Ma &amp; Yang &amp; Liu 2020, sp. nov." docType="treatment" docUuid="5E1748C9-4C93-4DAC-9C9D-66C63FAA8EC8" docUuidSource="ZooBank" docVersion="4" id="F711F888C51E5B7696DD6DFAC353BE28" lastPageNumber="41" masterDocId="F711F888C51E5B7696DD6DFAC353BE28" masterDocTitle="Notes on the green lacewing subgenus Ankylopteryx Brauer, 1864 (s. str.) (Neuroptera, Chrysopidae) from China, with description of a new species" masterLastPageNumber="71" masterPageNumber="41" pageNumber="41" updateTime="1668168355858" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Notes on the green lacewing subgenus Ankylopteryx Brauer, 1864 (s. str.) (Neuroptera, Chrysopidae) from China, with description of a new species</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ma, Yunlong</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yang, Xingke</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liu, Xingyue</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>906</mods:number>
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<mods:start>41</mods:start>
<mods:end>71</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.906.46438</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.906.46438</mods:identifier>
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<treatment ID-GBIF-Taxon="161838105" LSID="urn:lsid:zoobank.org:act:5E1748C9-4C93-4DAC-9C9D-66C63FAA8EC8" httpUri="http://treatment.plazi.org/id/73095F4DA45B5F21B35551606C3164D8" lastPageNumber="41" pageId="0" pageNumber="41">
<subSubSection pageId="0" pageNumber="41" type="nomenclature">
<paragraph pageId="0" pageNumber="41">
<taxonomicName LSID="73095F4D-A45B-5F21-B355-51606C3164D8" authority="Ma &amp; Yang &amp; Liu, 2020" authorityName="Ma &amp; Yang &amp; Liu" authorityYear="2020" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ankylopteryx yangi" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov.">Ankylopteryx (A.) yangi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 19" captionStartId="F1" captionText="Figures 1 - 9. Habitus photos of the Ankylopteryx species. 1 Ankylopteryx (A.) delicatula Banks, 1937 (Yunnan, Jinghong, sex unknown, CAU) 2 Ankylopteryx (A.) doleschalii Brauer, 1864, paratype (Indonesia, Ambonia, paratype, male, provided by NHMV) 3 Ankylopteryx (A.) ferruginea Tsukaguchi, 1995 (Guangxi, Longzhou, female, CAU) 4 Ankylopteryx (A.) gracilis Nakahara, 1955 (Guangxi, Ningming, female, CAU) 5 Ankylopteryx (A.) lii Yang, 1987 (Xizang, Zaeyu, paratype, male, CAU) 6 Ankylopteryx (A.) magnimaculata Yang, 1987 (Xizang, Zaeyu, holotype, male, CAU) 7 Ankylopteryx (A.) octopunctata candida (Fabricius, 1798) (Guangxi, Ningming, female, CAU) 8 Ankylopteryx (A.) yangi sp. nov. (Taiwan, Pingtung, paratype, female, CAU) 9 Ankylopteryx (A.) tibetana Yang, 1987 (Xizang, Zaeyu, holotype, male, CAU). Scale bars: 5.0 mm." figureDoi="10.3897/zookeys.906.46438.figures1-9" httpUri="https://binary.pensoft.net/fig/374369" pageId="0" pageNumber="41">Figs 9</figureCitation>
<figureCitation captionStart="Figures 8690" captionStartId="F17" captionText="Figures 86 - 90. Ankylopteryx (A.) yangi sp. nov. (Guizhou, Libo, holotype, male, CAU). 86 head, frontal 87 head, lateral 88 head and thorax, dorsal 89 protibia and mesotibia 90 forewing and hind wing." figureDoi="10.3897/zookeys.906.46438.figures86-90" httpUri="https://binary.pensoft.net/fig/374385" pageId="0" pageNumber="41">, 86-90</figureCitation>
<figureCitation captionStart="Figures 9198" captionStartId="F18" captionText="Figures 91 - 98. 91 - 94 Ankylopteryx (A.) yangi sp. nov., male abdomen (Guizhou, Libo, holotype, male, CAU). 91 segment A 7 - terminus, lateral 92 gonarcal complex, dorsal 93 gonarcal complex, lateral 94 gonarcus. 95 - 98 Ankylopteryx (A.) yangi sp. nov., female abdomen (Guizhou, Libo, paratype, female, CAU). 95 segment A 7 - terminus, lateral 96 terminalia, ventral 97 callus cerci 98 spermatheca. cc callus cerci ent entoprocessus g. l. gonaphophyses lateralis gsac gonosaccus gon gonarcus pp pseudopenis S 7 seventh sternum S 8 + 9 fused eighth + ninth sternum sg. d. dorsal lobe of subgenitale sg. v. ventral lobe of subgenitale sm spermatheca sm. d. spermathecal duct t. s. transverse sclerite T 8 eighth tergum T 9 + e ninth tergum + ectoproct." figureDoi="10.3897/zookeys.906.46438.figures91-98" httpUri="https://binary.pensoft.net/fig/374386" pageId="0" pageNumber="41">, 91-94</figureCitation>
<figureCitation captionStart="Figures 9198" captionStartId="F18" captionText="Figures 91 - 98. 91 - 94 Ankylopteryx (A.) yangi sp. nov., male abdomen (Guizhou, Libo, holotype, male, CAU). 91 segment A 7 - terminus, lateral 92 gonarcal complex, dorsal 93 gonarcal complex, lateral 94 gonarcus. 95 - 98 Ankylopteryx (A.) yangi sp. nov., female abdomen (Guizhou, Libo, paratype, female, CAU). 95 segment A 7 - terminus, lateral 96 terminalia, ventral 97 callus cerci 98 spermatheca. cc callus cerci ent entoprocessus g. l. gonaphophyses lateralis gsac gonosaccus gon gonarcus pp pseudopenis S 7 seventh sternum S 8 + 9 fused eighth + ninth sternum sg. d. dorsal lobe of subgenitale sg. v. ventral lobe of subgenitale sm spermatheca sm. d. spermathecal duct t. s. transverse sclerite T 8 eighth tergum T 9 + e ninth tergum + ectoproct." figureDoi="10.3897/zookeys.906.46438.figures91-98" httpUri="https://binary.pensoft.net/fig/374386" pageId="0" pageNumber="41">, 95-98</figureCitation>
<figureCitation captionStart="Figures 99107" captionStartId="F19" captionText="Figures 99 - 107. Gonarcal complex. 99 Ankylopteryx (A.) lii Yang, dorsal 100 Ankylopteryx (A.) octopunctata candida (Fabricius), dorsal 101 Ankylopteryx (A.) tibetana Yang, dorsal 102 Ankylopteryx (A.) magnimaculatus Yang, dorsal 103 Ankylopteryx (A.) yangi sp. nov., dorsal 104 Ankylopteryx (A.) octopunctata candida (Fabricius), lateral 105 Ankylopteryx (A.) yangi sp. nov., lateral 106 Ankylopteryx (A.) ferruginea Tsukaguchi, dorsal 107 Ankylopteryx (A.) gracilis Nakahara, dorsal." figureDoi="10.3897/zookeys.906.46438.figures99-107" httpUri="https://binary.pensoft.net/fig/374387" pageId="0" pageNumber="41">, 103</figureCitation>
<figureCitation captionStart="Figures 99107" captionStartId="F19" captionText="Figures 99 - 107. Gonarcal complex. 99 Ankylopteryx (A.) lii Yang, dorsal 100 Ankylopteryx (A.) octopunctata candida (Fabricius), dorsal 101 Ankylopteryx (A.) tibetana Yang, dorsal 102 Ankylopteryx (A.) magnimaculatus Yang, dorsal 103 Ankylopteryx (A.) yangi sp. nov., dorsal 104 Ankylopteryx (A.) octopunctata candida (Fabricius), lateral 105 Ankylopteryx (A.) yangi sp. nov., lateral 106 Ankylopteryx (A.) ferruginea Tsukaguchi, dorsal 107 Ankylopteryx (A.) gracilis Nakahara, dorsal." figureDoi="10.3897/zookeys.906.46438.figures99-107" httpUri="https://binary.pensoft.net/fig/374387" pageId="0" pageNumber="41">, 105</figureCitation>
<figureCitation captionStart="Figure 112" captionStartId="F21" captionText="Figure 112. Known distribution of Ankylopteryx (s. str.) species from China and neighboring countries. Ankylopteryx (A.) delicatula Banks (white triangle); Ankylopteryx (A.) doleschalii Brauer (black triangle); Ankylopteryx (A.) ferruginea Tsukaguchi (circle); Ankylopteryx (A.) magnimaculata Yang (white square); Ankylopteryx (A.) yangi sp. n. (black square)." figureDoi="10.3897/zookeys.906.46438.figure112" httpUri="https://binary.pensoft.net/fig/374389" pageId="0" pageNumber="41">, 112</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
<paragraph pageId="0" pageNumber="41">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="41">Holotype ♂, China, Guizhou, Libo, Maolan, Limingguan, 2013.X.14, Liu Xingyue (CAU). Paratypes 1 ♀, China, Guizhou, Libo, Maolan Limingguan, 2013.X.14, Liang Feiyang (CAU); 1 ♀, Taiwan, Pingtung, Lilungshan, 2013.VI.18, Liang Feiyang (CAU).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="diagnosis">
<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="41">Frons with three brownish small spots between antennae, and anteriorly with two arcuate markings, which are more or less connected with each other at posterior ends; gena with long brownish stripe; clypeal markings contiguous with frontal markings posteriorly and labial markings anteriorly; maxillary palp and labial palp pale green. Thorax with mesonotum entirely brown; protibia and mesotibia with median markings. Forewing with intramedian cell short and wide. Abdomen with brownish markings on terga 2-8.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Description.</paragraph>
<paragraph pageId="0" pageNumber="41">Measurements: Head width 0.7-1.0 mm; ratio of head width/eye width 1.72-1.74; prothorax 0.7-0.9 mm long and 0.85-1.0 mm wide. Forewing 11.5-13.9 mm long, 5.0-6.0 mm wide; length of first intramedian cell 0.98 mm; 12 radial cells; 4-5 Banksian cells (b cells), 4-5 lower Banksian cells (b' cells); 7-8 inner gradates, 7-10 outer gradates. Hind wing 10.0-13.5 mm long, 3.2-4.0 mm wide; 11 radial cells; 4-6 Banksian cells (b cell), 5-6 lower Banksian cell (b' cells); 4-6 inner gradates, 5-7 outer gradates.</paragraph>
<paragraph pageId="0" pageNumber="41">Male. Head with vertex creamy yellow, immaculate; frons creamy yellow, with three small spots between antennae, brownish stripes below toruli, and a pair of median arcuate markings anteriorly, more or less connected with each other posteriorly; gena with long brownish stripe extending along inner ocular margin to posterolateral part of clypeus; tentorial pits with brownish margins; scape with brownish stripe; clypeus with brownish arcuate markings contiguous with frontal markings anteriorly and labial markings posteriorly; maxillary palp and labial palp pale green.</paragraph>
<paragraph pageId="0" pageNumber="41">Prothorax almost pale green, with wide, brownish lateral stripe, and with white long setae. Mesothorax entirely brown dorsally, with sparse white long setae. Metathorax pale green, with sparse white long setae. Legs pale green, tarsomere 5 and pretarsal claws dark brown; protibia with a brownish median marking; mesotibia with a smaller median brownish marking.</paragraph>
<paragraph pageId="0" pageNumber="41">Forewing broad, slightly tapering apically, hyaline; pterostigma brownish, extending over four crossveins; setae almost whitish; veins mostly pale green; costal crossveins at junctions with wing margin, radial crossveins at junctions with R and dcc brown; Radical sector (Rs) sinuated; transverse veins pale green; gradate series of crossveins almost brown; dcc closed. Hind wing narrow, more acutely tapering apically than forewing, hyaline; pterostigma faint, extending over three crossveins; transverse veins pale green; gradate series of crossveins almost brown.</paragraph>
<paragraph pageId="0" pageNumber="41">Abdomen pale green, with brownish markings on terga 2-8, and tergal markings slightly wider than that on anteriorly neighbouring terga. Abdominal setae white, microsetae dense, and long setae sparse.</paragraph>
<paragraph pageId="0" pageNumber="41">T9+ectoproct about half as long as tergum 8, with dorsal invagination shallow; ectoproct with rounded dorsal and posterior margins; callus cerci rounded, trichobothria densely ranged. S8+9 fused, as long as wide, with line of fusion not demarcated; lateral margin almost straight, posterior margin rounded. Only gonarcus, entoprocessus, and pseudopenis present. Gonarcus broadened at apex of lateral arms. Gonarcus with the general width normally broad; entoprocessus attached at about basal fourth of gonarcus, about half as long as gonarcus, medially fused forming an arch over pseudopenis; pseudopenis about twice as long as entoprocessus, broadened subapically, long and distinctly curved, and pointed apically; gonosaccus with sparse setae.</paragraph>
<paragraph pageId="0" pageNumber="41">Female. External characters same as male. Sternum 7 distinctly convex posteromedially, setose posteriorly. Subgenitale and spermatheca with spermaduct present; subgenitale bilobed apically; spermatheca round, as wide as long; spermaduct coiled, much longer than spermatheca.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="distribution">
<paragraph pageId="0" pageNumber="41">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="41">China (Guizhou, Taiwan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="etymology">
<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="41">
This new species is dedicated to Professor Yang Chikun, who made tremendous contributions to the taxonomy of
<taxonomicName class="Insecta" family="Chrysopidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="family">Chrysopidae</taxonomicName>
from China.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="remarks">
<paragraph pageId="0" pageNumber="41">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="41">
This new species appears to be closely related to
<taxonomicName authority="candida" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. octopunctata" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="octopunctata">
<emphasis italics="true" pageId="0" pageNumber="41">A. octopunctata candida</emphasis>
</taxonomicName>
in having similar frontal markings, but it differs from the latter species by the presence of median arcuate frontal markings (absent in
<taxonomicName authority="candida" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. octopunctata" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="octopunctata">
<emphasis italics="true" pageId="0" pageNumber="41">A. octopunctata candida</emphasis>
</taxonomicName>
), the ratio of gonarcus/entoprocessus (2.0 in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. yangi" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="41">A. yangi</emphasis>
</taxonomicName>
versus 1.0 in
<taxonomicName authority="candida" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. octopunctata" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="octopunctata">
<emphasis italics="true" pageId="0" pageNumber="41">A. octopunctata candida</emphasis>
</taxonomicName>
), and the distinctly curved pseudopenis (moderately curved in
<taxonomicName authority="candida" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. octopunctata" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="octopunctata">
<emphasis italics="true" pageId="0" pageNumber="41">A. octopunctata candida</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="key to chinese species of ankylopteryx (s. str.) (revised after yang et al. 2005)">
<paragraph pageId="0" pageNumber="41">
Key to Chinese species of
<taxonomicName class="Insecta" family="Chrysopidae" genus="Ankylopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ankylopteryx" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="41">Ankylopteryx</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="0" pageNumber="41">s. str.</emphasis>
) (revised after
<bibRefCitation author="Yang, XK" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="41" refId="B41" refString="Yang, XK, Yang, CK, Li, WZ, 2005. Fauna Sinica: Insecta, Volume 39: Neuroptera: Chrysopidae. Science Press, Beijing" title="Fauna Sinica: Insecta, Volume 39: Neuroptera: Chrysopidae." year="2005">Yang et al. 2005</bibRefCitation>
)1
</paragraph>
<paragraph pageId="0" pageNumber="41">
<table pageId="0" pageNumber="41">
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">First intramedian cell very long and narrow</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">First intramedian cell short and wide</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Frontal area between antennae with three black spots (Fig.
<figureCitation captionStart="Figures 3640" captionStartId="F7" captionText="Figures 36 - 40. Ankylopteryx (A.) gracilis Nakahara (Japan, Okinawa, Iriomote-jima, male, CAU). 36 head, frontal 37 head, lateral 38 head and thorax, dorsal 39 protibia 40 forewing and hind wing." figureDoi="10.3897/zookeys.906.46438.figures36-40" httpUri="https://binary.pensoft.net/fig/374375" pageId="0" pageNumber="41">36</figureCitation>
); scape and pedicel with brownish stripes (Fig.
<figureCitation captionStart="Figures 3640" captionStartId="F7" captionText="Figures 36 - 40. Ankylopteryx (A.) gracilis Nakahara (Japan, Okinawa, Iriomote-jima, male, CAU). 36 head, frontal 37 head, lateral 38 head and thorax, dorsal 39 protibia 40 forewing and hind wing." figureDoi="10.3897/zookeys.906.46438.figures36-40" httpUri="https://binary.pensoft.net/fig/374375" pageId="0" pageNumber="41">37</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. gracilis" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="41">A. gracilis</emphasis>
</taxonomicName>
Nakahara, 1955
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Frontal area between antennae immaculate; scape and pedicel immaculate</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Both wings with yellowish marking patterns (Figs
<figureCitation captionStart="Figures 2329" captionStartId="F5" captionText="Figures 23 - 29. Ankylopteryx (A.) ferruginea Tsukaguchi. 23 head, frontal 24 head, lateral 25 thorax, dorsal 26 protibia 27 mesotibia and metatibia 28 forewing 29 hind wing (23 - 27: Yunnan, Lancang, male, CAU; 28, 29: Hainan, Yinggeling, female, CAU)." figureDoi="10.3897/zookeys.906.46438.figures23-29" httpUri="https://binary.pensoft.net/fig/374373" pageId="0" pageNumber="41">28</figureCitation>
,
<figureCitation captionStart="Figures 2329" captionStartId="F5" captionText="Figures 23 - 29. Ankylopteryx (A.) ferruginea Tsukaguchi. 23 head, frontal 24 head, lateral 25 thorax, dorsal 26 protibia 27 mesotibia and metatibia 28 forewing 29 hind wing (23 - 27: Yunnan, Lancang, male, CAU; 28, 29: Hainan, Yinggeling, female, CAU)." figureDoi="10.3897/zookeys.906.46438.figures23-29" httpUri="https://binary.pensoft.net/fig/374373" pageId="0" pageNumber="41">29</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. ferruginea" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis italics="true" pageId="0" pageNumber="41">A. ferruginea</emphasis>
</taxonomicName>
Tsukaguchi, 1995
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Both wings with brownish marking patterns (Figs
<figureCitation captionStart="Figures 1822" captionStartId="F4" captionText="Figures 18 - 22. Ankylopteryx (A.) doleschalii Brauer (Indonesia, Ambonia, holotype, ex, provided by NHMV). 18 head, frontal 19 head and thorax, dorsal 20 protibia and mesotibia 21 forewing 22 hind wing." figureDoi="10.3897/zookeys.906.46438.figures18-22" httpUri="https://binary.pensoft.net/fig/374372" pageId="0" pageNumber="41">21</figureCitation>
,
<figureCitation captionStart="Figures 1822" captionStartId="F4" captionText="Figures 18 - 22. Ankylopteryx (A.) doleschalii Brauer (Indonesia, Ambonia, holotype, ex, provided by NHMV). 18 head, frontal 19 head and thorax, dorsal 20 protibia and mesotibia 21 forewing 22 hind wing." figureDoi="10.3897/zookeys.906.46438.figures18-22" httpUri="https://binary.pensoft.net/fig/374372" pageId="0" pageNumber="41">22</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. doleschalii" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="doleschalii">
<emphasis italics="true" pageId="0" pageNumber="41">A. doleschalii</emphasis>
</taxonomicName>
Brauer, 1864
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Frontal area between antennae with brownish spots</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">Frontal area between antennae immaculate</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Three spots present (Fig.
<figureCitation captionStart="Figures 8690" captionStartId="F17" captionText="Figures 86 - 90. Ankylopteryx (A.) yangi sp. nov. (Guizhou, Libo, holotype, male, CAU). 86 head, frontal 87 head, lateral 88 head and thorax, dorsal 89 protibia and mesotibia 90 forewing and hind wing." figureDoi="10.3897/zookeys.906.46438.figures86-90" httpUri="https://binary.pensoft.net/fig/374385" pageId="0" pageNumber="41">86</figureCitation>
); both wings without large brownish vittae along posterior margins (Fig.
<figureCitation captionStart="Figures 8690" captionStartId="F17" captionText="Figures 86 - 90. Ankylopteryx (A.) yangi sp. nov. (Guizhou, Libo, holotype, male, CAU). 86 head, frontal 87 head, lateral 88 head and thorax, dorsal 89 protibia and mesotibia 90 forewing and hind wing." figureDoi="10.3897/zookeys.906.46438.figures86-90" httpUri="https://binary.pensoft.net/fig/374385" pageId="0" pageNumber="41">90</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. yangi" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="41">A. yangi</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Two spots present (Fig.
<figureCitation captionStart="Figures 1014" captionStartId="F2" captionText="Figures 10 - 14. Ankylopteryx (A.) delicatula Banks (Yunnan, Jinghong, sex unknown, CAU). 10 head, frontal 11 head, dorsal 12 thorax, dorsal 13 protibia and mesotibia 14 forewing and hind wing. Veins (black lettering): C Costa Sc Subcosta R Radius Rs Radial sector Psc Pseudocubitus Psm Pseudomedia. Cells (red lettering): b 1, b 4 first, fourth upper Banksian cells b' 1, b' 4 first, fourth lower Banksian cells r 1, r 9, r 11 first, ninth, eleventh radial cell." figureDoi="10.3897/zookeys.906.46438.figures10-14" httpUri="https://binary.pensoft.net/fig/374370" pageId="0" pageNumber="41">10</figureCitation>
); both wings with large brownish vittae along posterior margins near base (Fig.
<figureCitation captionStart="Figures 1014" captionStartId="F2" captionText="Figures 10 - 14. Ankylopteryx (A.) delicatula Banks (Yunnan, Jinghong, sex unknown, CAU). 10 head, frontal 11 head, dorsal 12 thorax, dorsal 13 protibia and mesotibia 14 forewing and hind wing. Veins (black lettering): C Costa Sc Subcosta R Radius Rs Radial sector Psc Pseudocubitus Psm Pseudomedia. Cells (red lettering): b 1, b 4 first, fourth upper Banksian cells b' 1, b' 4 first, fourth lower Banksian cells r 1, r 9, r 11 first, ninth, eleventh radial cell." figureDoi="10.3897/zookeys.906.46438.figures10-14" httpUri="https://binary.pensoft.net/fig/374370" pageId="0" pageNumber="41">14</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. delicatula" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="delicatula">
<emphasis italics="true" pageId="0" pageNumber="41">A. delicatula</emphasis>
</taxonomicName>
Banks, 1937
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Both wings with brownish vittae along posterior margins (Figs
<figureCitation captionStart="Figures 5863" captionStartId="F11" captionText="Figures 58 - 63. Ankylopteryx (A.) magnimaculatus Yang (Xizang, Zaeyu, holotype, male, CAU). 58 head, frontal 59 head, lateral 60 head and thorax, dorsal 61 protibia and mesotibia 62 forewing 63 hind wing." figureDoi="10.3897/zookeys.906.46438.figures58-63" httpUri="https://binary.pensoft.net/fig/374379" pageId="0" pageNumber="41">62</figureCitation>
,
<figureCitation captionStart="Figures 5863" captionStartId="F11" captionText="Figures 58 - 63. Ankylopteryx (A.) magnimaculatus Yang (Xizang, Zaeyu, holotype, male, CAU). 58 head, frontal 59 head, lateral 60 head and thorax, dorsal 61 protibia and mesotibia 62 forewing 63 hind wing." figureDoi="10.3897/zookeys.906.46438.figures58-63" httpUri="https://binary.pensoft.net/fig/374379" pageId="0" pageNumber="41">63</figureCitation>
); general width of gonarcus narrow (Figs
<figureCitation captionStart="Figures 6466" captionStartId="F12" captionText="Figures 64 - 66. Ankylopteryx (A.) lii Yang, male abdomen (Xizang, Zaeyu, holotype, male, CAU). 64 segment A 7 - terminus, lateral 65 gonarcal complex, dorsal 66 gonarcal complex, dorsolateral. cc callus cerci ent entoprocessus gsac gonosaccus gon gonarcus pp pseudopenis S 8 + 9 fused eighth + ninth sternum T 8 eighth tergum T 9 + e ninth tergum + ectoproct." figureDoi="10.3897/zookeys.906.46438.figures64-66" httpUri="https://binary.pensoft.net/fig/374380" pageId="0" pageNumber="41">65</figureCitation>
,
<figureCitation captionStart="Figures 6466" captionStartId="F12" captionText="Figures 64 - 66. Ankylopteryx (A.) lii Yang, male abdomen (Xizang, Zaeyu, holotype, male, CAU). 64 segment A 7 - terminus, lateral 65 gonarcal complex, dorsal 66 gonarcal complex, dorsolateral. cc callus cerci ent entoprocessus gsac gonosaccus gon gonarcus pp pseudopenis S 8 + 9 fused eighth + ninth sternum T 8 eighth tergum T 9 + e ninth tergum + ectoproct." figureDoi="10.3897/zookeys.906.46438.figures64-66" httpUri="https://binary.pensoft.net/fig/374380" pageId="0" pageNumber="41">66</figureCitation>
,
<figureCitation captionStart="Figures 99107" captionStartId="F19" captionText="Figures 99 - 107. Gonarcal complex. 99 Ankylopteryx (A.) lii Yang, dorsal 100 Ankylopteryx (A.) octopunctata candida (Fabricius), dorsal 101 Ankylopteryx (A.) tibetana Yang, dorsal 102 Ankylopteryx (A.) magnimaculatus Yang, dorsal 103 Ankylopteryx (A.) yangi sp. nov., dorsal 104 Ankylopteryx (A.) octopunctata candida (Fabricius), lateral 105 Ankylopteryx (A.) yangi sp. nov., lateral 106 Ankylopteryx (A.) ferruginea Tsukaguchi, dorsal 107 Ankylopteryx (A.) gracilis Nakahara, dorsal." figureDoi="10.3897/zookeys.906.46438.figures99-107" httpUri="https://binary.pensoft.net/fig/374387" pageId="0" pageNumber="41">102</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. magnimaculata" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="magnimaculata">
<emphasis italics="true" pageId="0" pageNumber="41">A. magnimaculata</emphasis>
</taxonomicName>
Yang, 1987
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="41">
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
Both wings without brownish vittae along posterior margins (Figs
<figureCitation captionStart="Figures 6772" captionStartId="F13" captionText="Figures 67 - 72. Ankylopteryx (A.) octopunctata candida (Fabricis) (Guangxi, Ningming, male, CAU). 67 head, frontal 68 head, frontolateral 69 head, thorax and protibia, dorsal 70 mesotibia 71 forewing 72 hind wing." figureDoi="10.3897/zookeys.906.46438.figures67-72" httpUri="https://binary.pensoft.net/fig/374381" pageId="0" pageNumber="41">71</figureCitation>
,
<figureCitation captionStart="Figures 6772" captionStartId="F13" captionText="Figures 67 - 72. Ankylopteryx (A.) octopunctata candida (Fabricis) (Guangxi, Ningming, male, CAU). 67 head, frontal 68 head, frontolateral 69 head, thorax and protibia, dorsal 70 mesotibia 71 forewing 72 hind wing." figureDoi="10.3897/zookeys.906.46438.figures67-72" httpUri="https://binary.pensoft.net/fig/374381" pageId="0" pageNumber="41">72</figureCitation>
); general width of gonarcus normal (Figs
<figureCitation captionStart="Figures 7376" captionStartId="F14" captionText="Figures 73 - 76. Ankylopteryx (A.) octopunctata candida Fabricius, male abdomen (Laos, Luang Namtha, male, CAU). 73 segment A 7 - terminus, lateral 74 callus cerci 75 gonarcal complex, dorsal 76 gonarcal complex, lateral. cc callus cerci ent entoprocessus gsac gonosaccus gon gonarcus pp pseudopenis S 8 + 9 fused eighth + ninth sternum T 8 eighth tergum T 9 + e ninth tergum + ectoproct." figureDoi="10.3897/zookeys.906.46438.figures73-76" httpUri="https://binary.pensoft.net/fig/374382" pageId="0" pageNumber="41">75</figureCitation>
,
<figureCitation captionStart="Figures 7376" captionStartId="F14" captionText="Figures 73 - 76. Ankylopteryx (A.) octopunctata candida Fabricius, male abdomen (Laos, Luang Namtha, male, CAU). 73 segment A 7 - terminus, lateral 74 callus cerci 75 gonarcal complex, dorsal 76 gonarcal complex, lateral. cc callus cerci ent entoprocessus gsac gonosaccus gon gonarcus pp pseudopenis S 8 + 9 fused eighth + ninth sternum T 8 eighth tergum T 9 + e ninth tergum + ectoproct." figureDoi="10.3897/zookeys.906.46438.figures73-76" httpUri="https://binary.pensoft.net/fig/374382" pageId="0" pageNumber="41">76</figureCitation>
,
<figureCitation captionStart="Figures 99107" captionStartId="F19" captionText="Figures 99 - 107. Gonarcal complex. 99 Ankylopteryx (A.) lii Yang, dorsal 100 Ankylopteryx (A.) octopunctata candida (Fabricius), dorsal 101 Ankylopteryx (A.) tibetana Yang, dorsal 102 Ankylopteryx (A.) magnimaculatus Yang, dorsal 103 Ankylopteryx (A.) yangi sp. nov., dorsal 104 Ankylopteryx (A.) octopunctata candida (Fabricius), lateral 105 Ankylopteryx (A.) yangi sp. nov., lateral 106 Ankylopteryx (A.) ferruginea Tsukaguchi, dorsal 107 Ankylopteryx (A.) gracilis Nakahara, dorsal." figureDoi="10.3897/zookeys.906.46438.figures99-107" httpUri="https://binary.pensoft.net/fig/374387" pageId="0" pageNumber="41">100</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="41" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="41">
<taxonomicName authority="candida" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="A. octopunctata" order="Neuroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="octopunctata">
<emphasis italics="true" pageId="0" pageNumber="41">A. octopunctata candida</emphasis>
</taxonomicName>
(Fabricius, 1798)
</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>