761 lines
92 KiB
XML
761 lines
92 KiB
XML
<document id="75B023C42B66D2D89471EC1758BF21E4" ID-CLB-Dataset="295785" ID-DOI="10.5281/zenodo.11148375" ID-GBIF-Dataset="87ba4acd-0a37-485d-a5e4-72131ddd82d0" ID-ISSN="1525-9153" ID-Zenodo-Dep="11148375" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714489072297" checkinUser="felipe" docAuthor="Poe, Steven, Scarpetta, Simon & Schaad, Eric W." docDate="2015" docId="2E4887F2FFD4CE57FF51F99EFD06F8E8" docLanguage="en" docName="AmphibReptileCons.9.1.e94.pdf" docOrigin="Amphibian & Reptile Conservation (e 94) 9 (1)" docStyle="DocumentStyle:D97478CDCB68366E5D4AECD282BFFB6F.1:AmphibReptileConse.2015-.journal_article.0cover" docStyleId="D97478CDCB68366E5D4AECD282BFFB6F" docStyleName="AmphibReptileConse.2015-.journal_article.0cover" docStyleVersion="1" docTitle="Anolis elcopeensis Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We 2015, sp. Nov" docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="D271FF8AFFD7CE5EFFDCFFFEFFA0FF8B" masterDocTitle="A new species of Anolis (Squamata: Iguanidae) from Panama" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1715175039523" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="ABD4DB791FF63A5E94D1DFAE4B63F2FA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A921C33D87E20D796D8E9AA1BABD9592">
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<mods:title id="D8DD085BAF7D6AE21A0B1199BAB24DC3">A new species of Anolis (Squamata: Iguanidae) from Panama</mods:title>
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<mods:name id="EBF11605267123FD57620D0F5EA625B5" type="personal">
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<mods:roleTerm id="8847FC1902A1D5E33D3661C749F09F3B">Author</mods:roleTerm>
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<mods:namePart id="ECC5F6701F782A059E44AAB8507E1259">Poe, Steven</mods:namePart>
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<mods:affiliation id="75BAAE5529C51E99263197927F816A26">Department Of Biology, Castetter Hall, University of New Mexico, Albuquerque, New Mexico 87131, USA</mods:affiliation>
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<mods:name id="505D47DBF442AE455D5CA3D40034FE4E" type="personal">
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<mods:roleTerm id="B61878D7E970063DA903A10284247C75">Author</mods:roleTerm>
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<mods:namePart id="8A8181262CE624E86457B7EFF8DC0BDD">Scarpetta, Simon</mods:namePart>
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<mods:affiliation id="4578C9BDAEDDAEA843AA724D4FFC9AB2">Field Conservation Facility, 477 Oak Road, Stanford University, Stanford, California 94305, USA</mods:affiliation>
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<mods:roleTerm id="3E9AFD03A0D08CC75A643E298755936C">Author</mods:roleTerm>
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<mods:namePart id="74650D97DE3AD9801ADC6CB5EC68A6E1">Schaad, Eric W.</mods:namePart>
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<mods:affiliation id="17D49B2A60CFCBE12D010F3385CD6D48">Department Of Biology, Castetter Hall, University of New Mexico, Albuquerque, New Mexico 87131, USA</mods:affiliation>
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<mods:typeOfResource id="2B663F3CB44F9B8E8554E83B45C90843">text</mods:typeOfResource>
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<mods:title id="CD42FBDDF1524903FE730AAFD93F335F">Amphibian & Reptile Conservation</mods:title>
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<mods:part id="675CB8E503F3D1C5271077C13188F2A6">
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<mods:date id="4BFC57FEC326E209AD5D24BFEA42820A">2015</mods:date>
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<mods:title id="18732AA5D8079A5B15D08E3E18EA2486">e 94</mods:title>
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<mods:number id="C028C3547FCC1FB1EB79AC4215F56D72">2015-04-27</mods:number>
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<mods:number id="455DD2A5F0CF1645C52EE61759EC405D">9</mods:number>
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<treatment id="2E4887F2FFD4CE57FF51F99EFD06F8E8" ID-DOI="http://doi.org/10.5281/zenodo.11148387" ID-Zenodo-Dep="11148387" LSID="urn:lsid:plazi:treatment:2E4887F2FFD4CE57FF51F99EFD06F8E8" httpUri="http://treatment.plazi.org/id/2E4887F2FFD4CE57FF51F99EFD06F8E8" lastPageId="9" lastPageNumber="10" pageId="3" pageNumber="4">
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<subSubSection id="EEFB656FFFD4CE5DFF51F99EFD49F932" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="A65E36E4FFD4CE5DFF51F99EFDA9F9F7" blockId="3.[141,521,1632,1660]" box="[141,521,1632,1660]" pageId="3" pageNumber="4">
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<heading id="FD168188FFD4CE5DFF51F99EFDA9F9F7" bold="true" box="[141,521,1632,1660]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
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<emphasis id="9495EAF6FFD4CE5DFF51F99EFDA9F9F7" bold="true" box="[141,521,1632,1660]" pageId="3" pageNumber="4">
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<taxonomicName id="61E14D67FFD4CE5DFF51F99EFE37F9F7" ID-CoL="5V5MF" authority="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We, 2015" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[141,407,1632,1660]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="elcopeensis" status="sp. nov.">
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<emphasis id="9495EAF6FFD4CE5DFF51F99EFE37F9F7" bold="true" box="[141,407,1632,1660]" italics="true" pageId="3" pageNumber="4">Anolis elcopeensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8FA6578DFFD4CE5DFE43F99EFDA9F9F7" box="[415,521,1632,1660]" pageId="3" pageNumber="4" rank="species">sp. Nov</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="A65E36E4FFD4CE5DFF51F958FD49F932" blockId="3.[141,745,1702,1721]" box="[141,745,1702,1721]" pageId="3" pageNumber="4">
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<emphasis id="9495EAF6FFD4CE5DFF51F958FD49F932" bold="true" box="[141,745,1702,1721]" pageId="3" pageNumber="4">
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<uri id="D2703AE6FFD4CE5DFF51F958FD49F932" box="[141,745,1702,1721]" pageId="3" pageNumber="4">
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urn:lsid:zoobank.org:act:
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<uuid id="D2470C31FFD4CE5DFEB9F958FD49F932" box="[357,745,1702,1721]" pageId="3" pageNumber="4">9D828BDE-E151-48FE-91E7-E176E693B382</uuid>
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</uri>
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||
</emphasis>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="EEFB656FFFD4CE5DFF51F91FFE25F972" box="[141,389,1761,1786]" pageId="3" pageNumber="4" type="description">
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<paragraph id="A65E36E4FFD4CE5DFF51F91FFE25F972" blockId="3.[141,389,1761,1786]" box="[141,389,1761,1786]" pageId="3" pageNumber="4">
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||
<figureCitation id="3EDA2A61FFD4CE5DFF51F91FFF4BF971" box="[141,235,1761,1786]" captionStart="Figure 3" captionStartId="4.[141,209,1896,1918]" captionTargetBox="[198,1353,144,1883]" captionTargetId="figure-23@4.[141,1394,141,1886]" captionTargetPageId="4" captionText="Figure 3. A) Anolis elcopeensis (female, El Copé, Coclé, Panama).B) A. cf elcopeensis (male, south of Gamboa, Panamá, Panama). C) A. cf maculiventris (female, Yaviza, Darién, Panama)." figureDoi="http://doi.org/10.5281/zenodo.11148381" httpUri="https://zenodo.org/record/11148381/files/figure.png" pageId="3" pageNumber="4">Figs. 3A</figureCitation>
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,
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||
<figureCitation id="3EDA2A61FFD4CE5DFF2BF91CFE99F971" box="[247,313,1762,1786]" captionStart="Figure 4" captionStartId="6.[141,209,1831,1853]" captionTargetBox="[204,1360,125,1824]" captionTargetId="figure-23@6.[141,1394,122,1827]" captionTargetPageId="6" captionText="Figure 4. A) Anolis elcopeensis (male dewlap, El Valle de Antón, Coclé, Panama). B) Anolis elcopeensis (female dewlap, El Copé, Coclé, Panama).C) A. cf.elcopeensis (male dewlap, south of Gamboa, Panamá, Panama).D) A. cf.elcopeensis (male dewlap, Cerro Azul, Panamá, Panama). E) A. cf. elcopeensis (male dewlap, Lake Bayano, Panama). F) A. cf. elcopeensis (male dewlap, Metetí, Darién, Panama). G) A. cf. maculiventris (male dewlap, Yaviza, Panama). H) A. maculiventris (male dewlap, near Buenaventura, Colombia)." figureDoi="http://doi.org/10.5281/zenodo.11148391" httpUri="https://zenodo.org/record/11148391/files/figure.png" pageId="3" pageNumber="4">4A–B</figureCitation>
|
||
,
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||
<figureCitation id="3EDA2A61FFD4CE5DFE98F91FFEC6F972" box="[324,358,1761,1785]" captionStart="Figure 5" captionStartId="7.[141,209,1213,1235]" captionTargetBox="[188,1348,144,1201]" captionTargetId="figure-19@7.[141,1394,141,1205]" captionTargetPageId="7" captionText="Figure 5. A) Anolis elcopeensis (male, El Copé, Coclé, Panama). B) A. gruuo (male, Hato Chami, Chiriquí, Panama). Note bulging tail base in A. gruuo." figureDoi="http://doi.org/10.5281/zenodo.11148395" httpUri="https://zenodo.org/record/11148395/files/figure.png" pageId="3" pageNumber="4">5A</figureCitation>
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||
,
|
||
<figureCitation id="3EDA2A61FFD4CE5DFEADF91FFE21F972" box="[369,385,1761,1785]" captionStart="Figure 6" captionStartId="8.[142,210,1004,1026]" captionTargetBox="[166,1340,144,990]" captionTargetId="figure-237@8.[141,1394,141,993]" captionTargetPageId="8" captionText="Figure 6. A) Ventral and B) dorsal head scales of holotype of A. elcopeensis." figureDoi="http://doi.org/10.5281/zenodo.11148399" httpUri="https://zenodo.org/record/11148399/files/figure.png" pageId="3" pageNumber="4">6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EEFB656FFFD4CE5DFF51F8DEFB4FF910" pageId="3" pageNumber="4" type="materials_examined">
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||
<paragraph id="A65E36E4FFD4CE5DFF51F8DEFEAAF8B7" blockId="3.[141,266,1824,1852]" box="[141,266,1824,1852]" pageId="3" pageNumber="4">
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||
<emphasis id="9495EAF6FFD4CE5DFF51F8DEFEAAF8B7" bold="true" box="[141,266,1824,1852]" pageId="3" pageNumber="4">
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||
<typeStatus id="795A8846FFD4CE5DFF51F8DEFEAAF8B7" box="[141,266,1824,1852]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
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||
<paragraph id="A65E36E4FFD4CE5DFF51F89DFA93FC73" blockId="3.[141,746,1890,1947]" lastBlockId="3.[791,1395,960,1016]" pageId="3" pageNumber="4">
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||
<materialsCitation id="16893CB9FFD4CE5DFF51F89DFA93FC73" collectionCode="MSB" collectorName="Parque Nacional G. D. Omar Torrijos H." country="Panama" latitude="8.66815" location="Cocle Province" longitude="-80.59267" pageId="3" pageNumber="4" specimenCode="MSB 95571" specimenCount="1" stateProvince="Cocle" typeStatus="holotype">
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||
<specimenCode id="F6479E9FFFD4CE5DFF51F89DFEB8F8F1" box="[141,280,1890,1915]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95571</specimenCode>
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||
, adult male from
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<collectorName id="0B145332FFD4CE5DFE2CF89DFEF3F811" pageId="3" pageNumber="4">Parque Nacional G.D. Omar Torrijos H.</collectorName>
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||
,
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||
<collectingRegion id="6425F806FFD4CE5DFEB9F87CFDB2F811" box="[357,530,1922,1946]" country="Panama" name="Cocle" pageId="3" pageNumber="4">Coclé Province</collectingRegion>
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||
,
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<collectingCountry id="DEF67674FFD4CE5DFDFEF87DFDD6F811" box="[546,630,1923,1946]" name="Panama" pageId="3" pageNumber="4">Panama</collectingCountry>
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(
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<geoCoordinate id="C3D55023FFD4CE5DFD57F87CFD46F811" box="[651,742,1922,1946]" degrees="8.66815" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="8.66815">8.66815</geoCoordinate>
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,
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||
<geoCoordinate id="C3D55023FFD4CE5DFCCBFC3EFC27FC53" box="[791,903,960,984]" degrees="80.59267" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="-80.59267">-80.59267</geoCoordinate>
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,
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||
<quantity id="61199B01FFD4CE5DFC4FFC3EFC7BFC53" box="[915,987,960,984]" metricMagnitude="2" metricUnit="m" metricValue="8.01" pageId="3" pageNumber="4" unit="m" value="801.0">801 m</quantity>
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), collected by Eric Schaad on
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<date id="D25F1024FFD4CE5DFAF4FC3EFC01FC73" pageId="3" pageNumber="4" value="2008-12-13">13 December 2008</date>
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from the trails near the visitor center.
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</materialsCitation>
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</paragraph>
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<paragraph id="A65E36E4FFD4CE5DFCCBFBE1FB2BFBB0" blockId="3.[791,1163,1055,1083]" box="[791,1163,1055,1083]" pageId="3" pageNumber="4">
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<emphasis id="9495EAF6FFD4CE5DFCCBFBE1FB2BFBB0" bold="true" box="[791,1163,1055,1083]" pageId="3" pageNumber="4">
|
||
<typeStatus id="795A8846FFD4CE5DFCCBFBE1FC03FBB0" box="[791,931,1055,1083]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
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||
scored for traits
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||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD4CE5DFCCBFB9CFCF3FA71" blockId="3.[790,1396,1121,1530]" pageId="3" pageNumber="4">
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<materialsCitation id="16893CB9FFD4CE5DFCCBFB9CFA8EFBF2" box="[791,1326,1121,1146]" collectionCode="MSB" collectorName="Parque Nacional G. D. Omar Torrijos H." country="Panama" latitude="8.66815" location="Cocle Province" pageId="3" pageNumber="4" specimenCode="MSB 95570" specimenCount="1" stateProvince="Cocle" typeStatus="paratype">
|
||
<specimenCode id="F6479E9FFFD4CE5DFCCBFB9CFC3BFBF2" box="[791,923,1121,1146]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95570</specimenCode>
|
||
bears the same data as the holotype
|
||
</materialsCitation>
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||
.
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<materialsCitation id="16893CB9FFD4CE5DFAE6FB9CFCF0FB51" collectingDate="2003-12" collectionCode="MSB" collectorName="Steven Poe & Caleb Hickman" country="Panama" latitude="8.66815" location="Cocle Province" pageId="3" pageNumber="4" specimenCode="MSB 95550-2" specimenCount="1" stateProvince="Cocle" typeStatus="paratype">
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||
<specimenCode id="F6479E9FFFD4CE5DFAE6FB9CFCD0FB12" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95550-2</specimenCode>
|
||
bear the same locality data as the
|
||
<typeStatus id="795A8846FFD4CE5DFB02FB7FFA9EFB12" box="[1246,1342,1153,1177]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
, collected by
|
||
<collectorName id="0B145332FFD4CE5DFC5FFB5CFC5DFB32" box="[899,1021,1186,1210]" pageId="3" pageNumber="4">Steven Poe</collectorName>
|
||
and
|
||
<collectorName id="0B145332FFD4CE5DFBE9FB5FFB7FFB32" box="[1077,1247,1185,1209]" pageId="3" pageNumber="4">Caleb Hickman</collectorName>
|
||
in
|
||
<date id="D25F1024FFD4CE5DFAD9FB5FFCF0FB51" pageId="3" pageNumber="4" value="2003-12">
|
||
<collectingDate id="C21BE9CCFFD4CE5DFAD9FB5FFCF0FB51" pageId="3" pageNumber="4" value="2003-12">December 2003</collectingDate>
|
||
</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="16893CB9FFD4CE5DFC82FB3CFBF6FAB1" collectingDate="2003-12" collectionCode="MSB" collectorName="Poe & Hickman" country="Panama" elevation="570" latitude="8.859476" location="El Valle de Anton" longLatPrecision="1" longitude="-80.14686" municipality="Chorro Las Mozas" pageId="3" pageNumber="4" specimenCode="MSB 95543-9" specimenCount="1" stateProvince="Cocle" typeStatus="paratype">
|
||
<specimenCode id="F6479E9FFFD4CE5DFC82FB3CFC5DFB52" box="[862,1021,1217,1242]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95543-9</specimenCode>
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||
, west of
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<location id="A33E603FFFD4CE5DFBB4FB3FFA94FB52" LSID="urn:lsid:plazi:treatment:2E4887F2FFD4CE57FF51F99EFD06F8E8:A33E603FFFD4CE5DFBB4FB3FFA94FB52" box="[1128,1332,1217,1241]" country="Panama" latitude="8.859476" longLatPrecision="1" longitude="-80.14686" municipality="Chorro Las Mozas" name="El Valle de Anton" pageId="3" pageNumber="4" stateProvince="Cocle">El Valle de Antón</location>
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||
, road ending at
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||
<collectingMunicipality id="463AAC9EFFD4CE5DFC5BFB1FFBF8FB72" box="[903,1112,1249,1273]" pageId="3" pageNumber="4">Chorro Las Mozas</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="6425F806FFD4CE5DFBBAFB1FFAB0FB72" box="[1126,1296,1249,1273]" country="Panama" name="Cocle" pageId="3" pageNumber="4">Coclé Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="DEF67674FFD4CE5DFAC2FB1CFAD2FB72" box="[1310,1394,1250,1273]" name="Panama" pageId="3" pageNumber="4">Panama</collectingCountry>
|
||
(
|
||
<geoCoordinate id="C3D55023FFD4CE5DFCC2FAFFFC26FA92" box="[798,902,1281,1305]" degrees="8.859476" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="1" value="8.859476">8.859476</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C3D55023FFD4CE5DFC44FAFFFBA8FA92" box="[920,1032,1281,1305]" degrees="80.14686" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="1" value="-80.14686">-80.14686</geoCoordinate>
|
||
,
|
||
<quantity id="61199B01FFD4CE5DFBC5FAFFFBC7FA91" box="[1049,1127,1281,1306]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="4" unit="m" value="570.0">
|
||
<elevation id="2DCCD1D7FFD4CE5DFBC5FAFFFBC7FA91" box="[1049,1127,1281,1306]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="4" unit="m" value="570.0">570 m</elevation>
|
||
</quantity>
|
||
), collected by
|
||
<collectorName id="0B145332FFD4CE5DFACBFAFCFA9FFA92" box="[1303,1343,1282,1305]" pageId="3" pageNumber="4">Poe</collectorName>
|
||
and
|
||
<collectorName id="0B145332FFD4CE5DFCCBFADFFCD9FAB2" box="[791,889,1313,1337]" pageId="3" pageNumber="4">Hickman</collectorName>
|
||
in
|
||
<date id="D25F1024FFD4CE5DFC78FADFFBF6FAB1" box="[932,1110,1313,1338]" pageId="3" pageNumber="4" value="2003-12">
|
||
<collectingDate id="C21BE9CCFFD4CE5DFC78FADFFBF6FAB1" box="[932,1110,1313,1338]" pageId="3" pageNumber="4" value="2003-12">December 2003</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="16893CB9FFD4CE5DFBB8FADCFC03FA11" collectingDate="2004-07-30" collectingDateMax="2004-08-04" collectingDateMin="2004-07-30" collectionCode="MSB" collectorName="Poe & Erik Hulebak & Heather MacInnes" country="Panama" location="El Valle de Anton" pageId="3" pageNumber="4" specimenCode="MSB 95554-7, 95560-1" specimenCount="1" stateProvince="Cocle" typeStatus="paratype">
|
||
<specimenCode id="F6479E9FFFD4CE5DFBB8FADCFAA4FAB2" box="[1124,1284,1313,1338]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95554-7</specimenCode>
|
||
,
|
||
<specimenCode id="F6479E9FFFD4CE5DFACFFADFFACFFAB2" box="[1299,1391,1313,1337]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">95560-1</specimenCode>
|
||
, same west of
|
||
<location id="A33E603FFFD4CE5DFC79FABFFBFEFAD1" LSID="urn:lsid:plazi:treatment:2E4887F2FFD4CE57FF51F99EFD06F8E8:A33E603FFFD4CE5DFC79FABFFBFEFAD1" box="[933,1118,1345,1370]" country="Panama" name="El Valle de Anton" pageId="3" pageNumber="4" stateProvince="Cocle">El Valle de Anton</location>
|
||
locality, collected by
|
||
<collectorName id="0B145332FFD4CE5DFA98FABCFACFFAD2" box="[1348,1391,1346,1369]" pageId="3" pageNumber="4">Poe</collectorName>
|
||
,
|
||
<collectorName id="0B145332FFD4CE5DFCCAFA9FFC0AFAF2" box="[790,938,1377,1401]" pageId="3" pageNumber="4">Erik Hulebak</collectorName>
|
||
, and
|
||
<collectorName id="0B145332FFD4CE5DFC3BFA9FFB0EFAF2" box="[999,1198,1377,1401]" pageId="3" pageNumber="4">Heather MacInnes</collectorName>
|
||
during
|
||
<date id="D25F1024FFD4CE5DFADAFA9CFC03FA11" pageId="3" pageNumber="4" value="2004-07-30" valueMax="2004-08-04" valueMin="2004-07-30">
|
||
<collectingDate id="C21BE9CCFFD4CE5DFADAFA9CFC03FA11" pageId="3" pageNumber="4" value="2004-07-30" valueMax="2004-08-04" valueMin="2004-07-30">30 July–4 August 2004</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="16893CB9FFD4CE5DFC71FA7CFCF0FA71" collectingDate="2004-07-31" collectionCode="MSB" collectorName="Poe & Hulebak & MacInnes" country="Panama" elevation="617" latitude="8.6129" location="Hotel Campestre" longLatPrecision="7" longitude="-80.1251" municipality="El Valle de Anton" pageId="3" pageNumber="4" specimenCode="MSB 95559" specimenCount="1" stateProvince="Cocle" typeStatus="paratype">
|
||
<specimenCode id="F6479E9FFFD4CE5DFC71FA7CFB91FA12" box="[941,1073,1409,1434]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95559</specimenCode>
|
||
,
|
||
<location id="A33E603FFFD4CE5DFBE0FA7FFB53FA11" LSID="urn:lsid:plazi:treatment:2E4887F2FFD4CE57FF51F99EFD06F8E8:A33E603FFFD4CE5DFBE0FA7FFB53FA11" box="[1084,1267,1409,1434]" country="Panama" latitude="8.6129" longLatPrecision="7" longitude="-80.1251" municipality="El Valle de Anton" name="Hotel Campestre" pageId="3" pageNumber="4" stateProvince="Cocle">Hotel Campestre</location>
|
||
,
|
||
<collectingMunicipality id="463AAC9EFFD4CE5DFB22FA7FFCFCFA32" pageId="3" pageNumber="4">El Valle de Antón</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="6425F806FFD4CE5DFCB5FA5FFBB1FA32" box="[873,1041,1441,1465]" country="Panama" name="Cocle" pageId="3" pageNumber="4">Coclé Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="DEF67674FFD4CE5DFBC2FA5CFBD2FA32" box="[1054,1138,1442,1465]" name="Panama" pageId="3" pageNumber="4">Panama</collectingCountry>
|
||
(
|
||
<geoCoordinate id="C3D55023FFD4CE5DFB5EFA5FFB6FFA32" box="[1154,1231,1441,1465]" degrees="8.6129" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="5" value="8.6129">8.6129</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C3D55023FFD4CE5DFB00FA5FFA9EFA32" box="[1244,1342,1441,1465]" degrees="80.1251" direction="west" orientation="longitude" pageId="3" pageNumber="4" precision="5" value="-80.1251">-80.1251</geoCoordinate>
|
||
,
|
||
<quantity id="61199B01FFD4CE5DFA97FA5FFC91FA51" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="3" pageNumber="4" unit="m" value="617.0">
|
||
<elevation id="2DCCD1D7FFD4CE5DFA97FA5FFC91FA51" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="3" pageNumber="4" unit="m" value="617.0">617 m</elevation>
|
||
</quantity>
|
||
), collected by
|
||
<collectorName id="0B145332FFD4CE5DFC10FA3CFC57FA52" box="[972,1015,1474,1497]" pageId="3" pageNumber="4">Poe</collectorName>
|
||
,
|
||
<collectorName id="0B145332FFD4CE5DFBDDFA3FFBFEFA52" box="[1025,1118,1473,1497]" pageId="3" pageNumber="4">Hulebak</collectorName>
|
||
, and
|
||
<collectorName id="0B145332FFD4CE5DFB4AFA3CFB5FFA52" box="[1174,1279,1474,1497]" pageId="3" pageNumber="4">MacInnes</collectorName>
|
||
on
|
||
<date id="D25F1024FFD4CE5DFAFBFA3CFCF0FA71" pageId="3" pageNumber="4" value="2004-07-31">
|
||
<collectingDate id="C21BE9CCFFD4CE5DFAFBFA3CFCF0FA71" pageId="3" pageNumber="4" value="2004-07-31">31 July 2004</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD4CE5DFCCBF9DEFB7EF9B7" blockId="3.[791,1246,1568,1596]" box="[791,1246,1568,1596]" pageId="3" pageNumber="4">
|
||
<emphasis id="9495EAF6FFD4CE5DFCCBF9DEFB7EF9B7" bold="true" box="[791,1246,1568,1596]" pageId="3" pageNumber="4">
|
||
Additional topotypical
|
||
<typeStatus id="795A8846FFD4CE5DFB8FF9DEFB7EF9B7" box="[1107,1246,1568,1596]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD4CE5DFCCBF99DFB4FF910" blockId="3.[790,1395,1634,1691]" pageId="3" pageNumber="4">
|
||
<materialsCitation id="16893CB9FFD4CE5DFCCBF99DFB4BF910" collectingDate="2008-12" collectionCode="MSB" collectorName="Schaad" country="Panama" latitude="8.66815" location="Cocle Province" pageId="3" pageNumber="4" specimenCode="MSB 95569, 95572-9" specimenCount="1" stateProvince="Cocle">
|
||
<specimenCode id="F6479E9FFFD4CE5DFCCBF99DFC3AF9F1" box="[791,922,1634,1659]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">MSB 95569</specimenCode>
|
||
,
|
||
<specimenCode id="F6479E9FFFD4CE5DFC78F99CFC5DF9F1" box="[932,1021,1634,1658]" collectionCode="MSB" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="3" pageNumber="4" type="Museum">95572-9</specimenCode>
|
||
bear the same data as the holotype, collected by
|
||
<collectorName id="0B145332FFD4CE5DFC42F97CFC4BF911" box="[926,1003,1666,1690]" pageId="3" pageNumber="4">Schaad</collectorName>
|
||
during
|
||
<date id="D25F1024FFD4CE5DFBE2F97CFB4BF910" box="[1086,1259,1666,1691]" pageId="3" pageNumber="4" value="2008-12">
|
||
<collectingDate id="C21BE9CCFFD4CE5DFBE2F97CFB4BF910" box="[1086,1259,1666,1691]" pageId="3" pageNumber="4" value="2008-12">December 2008</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EEFB656FFFD4CE5BFCCBF93FFB19FB63" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
|
||
<paragraph id="A65E36E4FFD4CE5DFCCBF93FFC03F956" blockId="3.[791,931,1729,1757]" box="[791,931,1729,1757]" pageId="3" pageNumber="4">
|
||
<emphasis id="9495EAF6FFD4CE5DFCCBF93FFC03F956" bold="true" box="[791,931,1729,1757]" pageId="3" pageNumber="4">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD4CE5BFCCBF8FDFECFFBA3" blockId="3.[791,1395,1795,1948]" lastBlockId="5.[140,748,784,1960]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
|
||
<taxonomicName id="61E14D67FFD4CE5DFCCBF8FDFC7FF890" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[791,991,1795,1819]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD4CE5DFCCBF8FDFC7FF890" box="[791,991,1795,1819]" italics="true" pageId="3" pageNumber="4">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is a small grayish-brown anole with smooth ventral scales and short limbs. We diagnose this species relative to its 11 closest relatives (
|
||
<figureCitation id="3EDA2A61FFD4CE5DFB08F8BDFAB3F8D7" box="[1236,1299,1859,1884]" captionStart="Fig" captionStartId="1.[141,176,1885,1907]" captionTargetBox="[177,1330,814,1871]" captionTargetId="figure-44@1.[141,1394,802,1878]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic estimate of placement of A. elcopeensis sp. nov. based on Bayesian analysis of morphological and molecular data. Numbers are clade credibility values." figureDoi="http://doi.org/10.5281/zenodo.11148377" httpUri="https://zenodo.org/record/11148377/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
).
|
||
<taxonomicName id="61E14D67FFD4CE5DFAF2F8BDFC32F8F0" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD4CE5DFAF2F8BDFC32F8F0" italics="true" pageId="3" pageNumber="4">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is unlikely to be confused with
|
||
<taxonomicName id="61E14D67FFD4CE5DFB2BF89AFCEEF810" authorityName="D'Orbigny" authorityYear="1837" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="9495EAF6FFD4CE5DFB2BF89AFCEEF810" italics="true" pageId="3" pageNumber="4">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
(Amazon basin; solid pink male dewlap),
|
||
<taxonomicName id="61E14D67FFD4CE5BFAF2F87AFF6DFCA3" authorityName="Schwartz & Thomas" authorityYear="1975" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="bocourti">
|
||
<emphasis id="9495EAF6FFD4CE5BFAF2F87AFF6DFCA3" italics="true" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">A. bocourti</emphasis>
|
||
</taxonomicName>
|
||
(Amazon basin; white male dewlap),
|
||
<taxonomicName id="61E14D67FFD2CE5BFDBCFCEFFD49FCA3" authorityName="Werner" authorityYear="1916" box="[608,745,784,808]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFDBCFCEFFD49FCA3" box="[608,745,784,808]" italics="true" pageId="5" pageNumber="6">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
(northeastern Andes; pink and orangish-red male dewlap),
|
||
<taxonomicName id="61E14D67FFD2CE5BFF15FCAFFE9EFCE3" authorityName="Ayala & Williams" authorityYear="1988" box="[201,318,848,872]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="medemi">
|
||
<emphasis id="9495EAF6FFD2CE5BFF15FCAFFE9EFCE3" box="[201,318,848,872]" italics="true" pageId="5" pageNumber="6">A. medemi</emphasis>
|
||
</taxonomicName>
|
||
(Gorgona Island,
|
||
<collectingCountry id="DEF67674FFD2CE5BFDD1FCAEFDDAFCE3" box="[525,634,848,872]" name="Colombia" pageId="5" pageNumber="6">Colombia</collectingCountry>
|
||
; pink and orange male dewlap),
|
||
<taxonomicName id="61E14D67FFD2CE5BFE5BFC8FFE55FC03" authorityName="Boulenger" authorityYear="1908" box="[391,501,881,904]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="9495EAF6FFD2CE5BFE5BFC8FFE55FC03" box="[391,501,881,904]" italics="true" pageId="5" pageNumber="6">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
(northwestern Andes; pink and reddish-orange dewlap), or
|
||
<taxonomicName id="61E14D67FFD2CE5BFDF5FC6FFD15FC23" authorityName="Barbour" authorityYear="1932" box="[553,693,913,936]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="9495EAF6FFD2CE5BFDF5FC6FFD15FC23" box="[553,693,913,936]" italics="true" pageId="5" pageNumber="6">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
(extreme northwestern Andes; orange-red and yellow dewlap) based on geography. Nevertheless,
|
||
<taxonomicName id="61E14D67FFD2CE5BFD90FC2FFD49FC63" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[588,745,976,1000]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFD90FC2FFD49FC63" box="[588,745,976,1000]" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
differs from each of these species in its solid orange male dewlap color pattern.
|
||
</paragraph>
|
||
<caption id="F29E666CFFD3CE5AFF51F896FD13F810" ID-DOI="http://doi.org/10.5281/zenodo.11148381" ID-Zenodo-Dep="11148381" httpUri="https://zenodo.org/record/11148381/files/figure.png" pageId="4" pageNumber="5" startId="4.[141,209,1896,1918]" targetBox="[198,1353,144,1883]" targetPageId="4" targetType="figure">
|
||
<paragraph id="A65E36E4FFD3CE5AFF51F896FD13F810" blockId="4.[140,1396,1896,1947]" pageId="4" pageNumber="5">
|
||
<emphasis id="9495EAF6FFD3CE5AFF51F896FEA5F8F5" bold="true" box="[141,261,1896,1918]" pageId="4" pageNumber="5">Figure 3. A)</emphasis>
|
||
<taxonomicName id="61E14D67FFD3CE5AFED6F897FE1CF8F5" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[266,444,1897,1918]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD3CE5AFED6F897FE1CF8F5" box="[266,444,1897,1918]" italics="true" pageId="4" pageNumber="5">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(female, El Copé, Coclé, Panama).
|
||
<emphasis id="9495EAF6FFD3CE5AFCD2F896FC86F8F5" bold="true" box="[782,806,1896,1918]" pageId="4" pageNumber="5">B)</emphasis>
|
||
<taxonomicName id="61E14D67FFD3CE5AFCF0F897FC6CF8F5" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[812,972,1897,1919]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD3CE5AFCF0F897FCE2F8F5" box="[812,834,1897,1918]" italics="true" pageId="4" pageNumber="5">A.</emphasis>
|
||
cf
|
||
<emphasis id="9495EAF6FFD3CE5AFC81F897FC6CF8F5" box="[861,972,1897,1918]" italics="true" pageId="4" pageNumber="5">elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male, south of Gamboa, Panamá, Panama).
|
||
<emphasis id="9495EAF6FFD3CE5AFF50F87BFF05F810" bold="true" box="[140,165,1925,1947]" pageId="4" pageNumber="5">C)</emphasis>
|
||
<taxonomicName id="61E14D67FFD3CE5AFF71F878FEC3F811" authorityName="Boulenger" authorityYear="1898" box="[173,355,1925,1947]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="9495EAF6FFD3CE5AFF71F878FF62F810" box="[173,194,1926,1947]" italics="true" pageId="4" pageNumber="5">A.</emphasis>
|
||
cf
|
||
<emphasis id="9495EAF6FFD3CE5AFF3CF87BFEC3F811" box="[224,355,1925,1946]" italics="true" pageId="4" pageNumber="5">maculiventris</emphasis>
|
||
</taxonomicName>
|
||
(female, Yaviza, Darién, Panama).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F29E666CFFD2CE5BFF51FD1EFC04FD7D" ID-DOI="http://doi.org/10.5281/zenodo.11148389" ID-Zenodo-Dep="11148389" box="[141,932,736,758]" httpUri="https://zenodo.org/record/11148389/files/figure.png" pageId="5" pageNumber="6" startId="5.[141,209,736,758]" targetBox="[198,1353,146,725]" targetPageId="5" targetType="figure">
|
||
<paragraph id="A65E36E4FFD2CE5BFF51FD1EFC04FD7D" blockId="5.[141,932,736,758]" box="[141,932,736,758]" pageId="5" pageNumber="6">
|
||
<emphasis id="9495EAF6FFD2CE5BFF51FD1EFE2AFD7D" bold="true" box="[141,394,736,758]" pageId="5" pageNumber="6">Figure 3 (continued). D)</emphasis>
|
||
<taxonomicName id="61E14D67FFD2CE5BFE4CFD1FFD8DFD7E" authorityName="Boulenger" authorityYear="1898" box="[400,557,736,758]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="9495EAF6FFD2CE5BFE4CFD1FFD8DFD7E" box="[400,557,736,758]" italics="true" pageId="5" pageNumber="6">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
(male, near Buenaventura, Colombia).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A65E36E4FFD2CE5BFF71FBCEFD03FAE3" blockId="5.[140,748,784,1960]" pageId="5" pageNumber="6">
|
||
<taxonomicName id="61E14D67FFD2CE5BFF71FBCEFED6FBC3" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[173,374,1072,1096]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFF71FBCEFED6FBC3" box="[173,374,1072,1096]" italics="true" pageId="5" pageNumber="6">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is most easily distinguished from the Central American members of its clade by male dewlap color (
|
||
<figureCitation id="3EDA2A61FFD2CE5BFEF0FB8EFED1FB03" box="[300,369,1136,1160]" captionStart="Figure 4" captionStartId="6.[141,209,1831,1853]" captionTargetBox="[204,1360,125,1824]" captionTargetId="figure-23@6.[141,1394,122,1827]" captionTargetPageId="6" captionText="Figure 4. A) Anolis elcopeensis (male dewlap, El Valle de Antón, Coclé, Panama). B) Anolis elcopeensis (female dewlap, El Copé, Coclé, Panama).C) A. cf.elcopeensis (male dewlap, south of Gamboa, Panamá, Panama).D) A. cf.elcopeensis (male dewlap, Cerro Azul, Panamá, Panama). E) A. cf. elcopeensis (male dewlap, Lake Bayano, Panama). F) A. cf. elcopeensis (male dewlap, Metetí, Darién, Panama). G) A. cf. maculiventris (male dewlap, Yaviza, Panama). H) A. maculiventris (male dewlap, near Buenaventura, Colombia)." figureDoi="http://doi.org/10.5281/zenodo.11148391" httpUri="https://zenodo.org/record/11148391/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
|
||
; except for
|
||
<taxonomicName id="61E14D67FFD2CE5BFE24FB8FFDF8FB03" authorityName="POE" authorityYear="1628" box="[504,600,1137,1160]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="9495EAF6FFD2CE5BFE24FB8FFDF8FB03" box="[504,600,1137,1160]" italics="true" pageId="5" pageNumber="6">A. gruuo</emphasis>
|
||
</taxonomicName>
|
||
) and smaller body size (maximum
|
||
<collectionCode id="C0F0AE21FFD2CE5BFEA7FB6EFE0CFB23" box="[379,428,1168,1192]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
=
|
||
<quantity id="61199B01FFD2CE5BFE11FB6EFDBCFB23" box="[461,540,1168,1192]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="6" unit="mm" value="45.0">45 mm</quantity>
|
||
,
|
||
<emphasis id="9495EAF6FFD2CE5BFDF7FB6FFD98FB23" box="[555,568,1169,1192]" italics="true" pageId="5" pageNumber="6">n</emphasis>
|
||
= 35):
|
||
<taxonomicName id="61E14D67FFD2CE5BFD50FB6FFD4CFB23" authorityName="POE" authorityYear="1628" box="[652,748,1169,1192]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="9495EAF6FFD2CE5BFD50FB6FFD4CFB23" box="[652,748,1169,1192]" italics="true" pageId="5" pageNumber="6">A. gruuo</emphasis>
|
||
</taxonomicName>
|
||
(solid orange male dewlap; maximum
|
||
<collectionCode id="C0F0AE21FFD2CE5BFDEBFB4EFDC8FB43" box="[567,616,1200,1224]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
=
|
||
<quantity id="61199B01FFD2CE5BFD57FB4EFD7EFB43" box="[651,734,1200,1224]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="6" unit="mm" value="52.0">52 mm</quantity>
|
||
);
|
||
<taxonomicName id="61E14D67FFD2CE5BFF51FB2FFEFCFB63" authorityName="Kohler, Ponce, Sunyer & Batista" authorityYear="2007" box="[141,348,1232,1256]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="pseudokemptoni">
|
||
<emphasis id="9495EAF6FFD2CE5BFF51FB2FFEFCFB63" box="[141,348,1232,1256]" italics="true" pageId="5" pageNumber="6">A. pseudokemptoni</emphasis>
|
||
</taxonomicName>
|
||
(red-orange anterior, pink posterior male dewlap; maximum
|
||
<collectionCode id="C0F0AE21FFD2CE5BFE7FFB0EFE74FA83" box="[419,468,1264,1288]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
=
|
||
<quantity id="61199B01FFD2CE5BFE26FB0EFDEEFA83" box="[506,590,1264,1288]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="6" unit="mm" value="55.0">55 mm</quantity>
|
||
);
|
||
<taxonomicName id="61E14D67FFD2CE5BFDB9FB0FFD4AFA83" authorityName="Dunn" authorityYear="1940" box="[613,746,1264,1288]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="kemptoni">
|
||
<emphasis id="9495EAF6FFD2CE5BFDB9FB0FFD4AFA83" box="[613,746,1264,1288]" italics="true" pageId="5" pageNumber="6">A. kemptoni</emphasis>
|
||
</taxonomicName>
|
||
(red-orange anterior, pink posterior male dewlap; maximum
|
||
<collectionCode id="C0F0AE21FFD2CE5BFF16FACEFF5BFAC3" box="[202,251,1328,1352]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
=
|
||
<quantity id="61199B01FFD2CE5BFECBFACEFEC6FAC3" box="[279,358,1328,1352]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="6" unit="mm" value="53.0">53 mm</quantity>
|
||
);
|
||
<taxonomicName id="61E14D67FFD2CE5BFEABFACFFDABFAC3" authorityName="Arosemena & Ibanez" authorityYear="1993" box="[375,523,1328,1352]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fortunensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFEABFACFFDABFAC3" box="[375,523,1328,1352]" italics="true" pageId="5" pageNumber="6">A. fortunensis</emphasis>
|
||
</taxonomicName>
|
||
(red anterior, orange posterior male dewlap; maximum
|
||
<collectionCode id="C0F0AE21FFD2CE5BFE27FAAEFD8CFAE3" box="[507,556,1360,1384]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
=
|
||
<quantity id="61199B01FFD2CE5BFD94FAAEFD38FAE3" box="[584,664,1360,1384]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="6" unit="mm" value="49.0">49 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD2CE5BFF71FA8FFB19FB63" blockId="5.[140,748,784,1960]" lastBlockId="5.[791,1397,784,1256]" pageId="5" pageNumber="6">
|
||
In the field
|
||
<taxonomicName id="61E14D67FFD2CE5BFEFFFA8FFE1AFA03" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[291,442,1392,1416]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFEFFFA8FFE1AFA03" box="[291,442,1392,1416]" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is most likely to be confused with
|
||
<taxonomicName id="61E14D67FFD2CE5BFF1EFA6FFEBFFA23" authorityName="POE" authorityYear="1628" box="[194,287,1425,1448]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="9495EAF6FFD2CE5BFF1EFA6FFEBFFA23" box="[194,287,1425,1448]" italics="true" pageId="5" pageNumber="6">A. gruuo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="61E14D67FFD2CE5BFE93FA6FFE7AFA23" authorityName="Echelle, Echelle & Fitch" authorityYear="1971" box="[335,474,1425,1448]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="9495EAF6FFD2CE5BFE93FA6FFE7AFA23" box="[335,474,1425,1448]" italics="true" pageId="5" pageNumber="6">A. carpenteri</emphasis>
|
||
</taxonomicName>
|
||
, which have similar solid orange male dewlaps (
|
||
<taxonomicName id="61E14D67FFD2CE5BFEA5FA4FFE6AFA43" authorityName="Dunn" authorityYear="1930" box="[377,458,1456,1480]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="altae">
|
||
<emphasis id="9495EAF6FFD2CE5BFEA5FA4FFE6AFA43" box="[377,458,1456,1480]" italics="true" pageId="5" pageNumber="6">A. altae</emphasis>
|
||
</taxonomicName>
|
||
, which also is similar, does not occur in
|
||
<collectingCountry id="DEF67674FFD2CE5BFEC2FA2EFED7FA63" box="[286,375,1488,1512]" name="Panama" pageId="5" pageNumber="6">Panama</collectingCountry>
|
||
); and
|
||
<taxonomicName id="61E14D67FFD2CE5BFE1DFA2FFDE2FA63" authorityName="Cope" authorityYear="1862" box="[449,578,1488,1512]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="limifrons">
|
||
<emphasis id="9495EAF6FFD2CE5BFE1DFA2FFDE2FA63" box="[449,578,1488,1512]" italics="true" pageId="5" pageNumber="6">A. limifrons</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="61E14D67FFD2CE5BFDA2FA2FFD43FA63" authorityName="Ruthven" authorityYear="1916" box="[638,739,1489,1512]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gaigei">
|
||
<emphasis id="9495EAF6FFD2CE5BFDA2FA2FFD43FA63" box="[638,739,1489,1512]" italics="true" pageId="5" pageNumber="6">A. gaigei</emphasis>
|
||
</taxonomicName>
|
||
, with which it is frequently sympatric (
|
||
<taxonomicName id="61E14D67FFD2CE5BFDF8FA0EFD49F983" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[548,745,1520,1544]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFDF8FA0EFD49F983" box="[548,745,1520,1544]" italics="true" pageId="5" pageNumber="6">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is amply genetically distinct from each of these species;
|
||
<figureCitation id="3EDA2A61FFD2CE5BFF52F9CEFF70F9C3" box="[142,208,1584,1608]" captionStart="Fig" captionStartId="1.[141,176,1885,1907]" captionTargetBox="[177,1330,814,1871]" captionTargetId="figure-44@1.[141,1394,802,1878]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic estimate of placement of A. elcopeensis sp. nov. based on Bayesian analysis of morphological and molecular data. Numbers are clade credibility values." figureDoi="http://doi.org/10.5281/zenodo.11148377" httpUri="https://zenodo.org/record/11148377/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="3EDA2A61FFD2CE5BFF02F9CEFF4EF9C3" box="[222,238,1584,1608]" captionStart="Fig" captionStartId="2.[141,176,1917,1939]" captionTargetBox="[180,1186,1037,1894]" captionTargetId="figure-455@2.[141,1394,1018,1915]" captionTargetPageId="2" captionText="Fig. 2. Phylogenetic estimate of the mitochondrial COI gene for samples of A. elcopeensis and close relatives based on Bayesian analysis. Numbers are clade credibility values." figureDoi="http://doi.org/10.5281/zenodo.11148379" httpUri="https://zenodo.org/record/11148379/files/figure.png" pageId="5" pageNumber="6">2</figureCitation>
|
||
;
|
||
<taxonomicName id="61E14D67FFD2CE5BFF25F9CFFEFAF9C3" authorityName="Ruthven" authorityYear="1916" box="[249,346,1585,1608]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gaigei">
|
||
<emphasis id="9495EAF6FFD2CE5BFF25F9CFFEFAF9C3" box="[249,346,1585,1608]" italics="true" pageId="5" pageNumber="6">A. gaigei</emphasis>
|
||
</taxonomicName>
|
||
is phylogenetically very distant from the
|
||
<taxonomicName id="61E14D67FFD2CE5BFF6BF9AEFEE1F9E3" authorityName="D'Orbigny" authorityYear="1837" box="[183,321,1616,1640]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="9495EAF6FFD2CE5BFF6BF9AEFEE1F9E3" box="[183,321,1616,1640]" italics="true" pageId="5" pageNumber="6">fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group, data not shown). In addition to being larger, male
|
||
<taxonomicName id="61E14D67FFD2CE5BFE87F98FFE1CF903" authorityName="POE" authorityYear="1628" box="[347,444,1649,1672]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="9495EAF6FFD2CE5BFE87F98FFE1CF903" box="[347,444,1649,1672]" italics="true" pageId="5" pageNumber="6">A. gruuo</emphasis>
|
||
</taxonomicName>
|
||
display an externally bulging tailbase in our photos, presumably indicating larger hemipenes, which we did not observe in male
|
||
<taxonomicName id="61E14D67FFD2CE5BFD4FF94FFF7CF963" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFD4FF94FFF7CF963" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="3EDA2A61FFD2CE5BFF32F92EFE93F963" box="[238,307,1744,1768]" captionStart="Figure 5" captionStartId="7.[141,209,1213,1235]" captionTargetBox="[188,1348,144,1201]" captionTargetId="figure-19@7.[141,1394,141,1205]" captionTargetPageId="7" captionText="Figure 5. A) Anolis elcopeensis (male, El Copé, Coclé, Panama). B) A. gruuo (male, Hato Chami, Chiriquí, Panama). Note bulging tail base in A. gruuo." figureDoi="http://doi.org/10.5281/zenodo.11148395" httpUri="https://zenodo.org/record/11148395/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
|
||
).
|
||
<taxonomicName id="61E14D67FFD2CE5BFE97F92EFE7AF963" authorityName="POE" authorityYear="1628" box="[331,474,1744,1768]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="9495EAF6FFD2CE5BFE97F92EFE7AF963" box="[331,474,1744,1768]" italics="true" pageId="5" pageNumber="6">Anolis gruuo</emphasis>
|
||
</taxonomicName>
|
||
is found at mid to high elevations (
|
||
<quantity id="61199B01FFD2CE5BFED5F90EFE39F883" box="[265,409,1776,1800]" metricMagnitude="3" metricUnit="m" metricValue="1.1949999999999998" metricValueMax="1.53" metricValueMin="0.86" pageId="5" pageNumber="6" unit="m" value="1195.0" valueMax="1530.0" valueMin="860.0">860–1,530 m</quantity>
|
||
) of the Serrania Tabasara from Santa Fe west
|
||
<quantity id="61199B01FFD2CE5BFEEDF8EEFED7F8A3" box="[305,375,1808,1832]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="6" unit="km" value="80.0">80 km</quantity>
|
||
to just past Hato Chami (
|
||
<bibRefCitation id="C2704B15FFD2CE5BFD4BF8EEFEA5F8C3" author="Lotzkat S & Stadler L & Batista A & Hertz A & Ponce M & Hamad N & Kohler G." pageId="5" pageNumber="6" pagination="620 - 625" refId="ref5687" refString="Lotzkat S, Stadler L, Batista A, Hertz A, Ponce M, Hamad N, Kohler G. 2012. Distribution extension for Anolis gruuo Kohler, Ponce, Sunyer and Batista, 2007 (Reptilia: Squamata: Dactyloidae) in the Comarca Ngobe-Bugle of western Panama, and first records from Veraguas province. Check List 8: 620 - 625." type="journal article" year="2012">Lotzkat et al. 2012</bibRefCitation>
|
||
). We found
|
||
<taxonomicName id="61E14D67FFD2CE5BFE56F8CEFDF0F8C3" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[394,592,1840,1864]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFE56F8CEFDF0F8C3" box="[394,592,1840,1864]" italics="true" pageId="5" pageNumber="6">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
at mid to low elevations (
|
||
<quantity id="61199B01FFD2CE5BFED0F8AEFE2BF8E3" box="[268,395,1872,1896]" metricMagnitude="2" metricUnit="m" metricValue="5.2299999999999995" metricValueMax="8.01" metricValueMin="2.45" pageId="5" pageNumber="6" unit="m" value="523.0" valueMax="801.0" valueMin="245.0">245–801 m</quantity>
|
||
) from El Copé east to Altos de Campana and possibly further (see below).
|
||
<taxonomicName id="61E14D67FFD2CE5BFDACF88EFF79F823" authorityName="Echelle, Echelle & Fitch" authorityYear="1971" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="9495EAF6FFD2CE5BFDACF88EFF79F823" italics="true" pageId="5" pageNumber="6">Anolis carpenteri</emphasis>
|
||
</taxonomicName>
|
||
has a dorsal greenish tint and we have observed it to become patterned only when stressed.
|
||
<taxonomicName id="61E14D67FFD2CE5BFB2FFCEEFCC6FCC3" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFB2FFCEEFCC6FCC3" italics="true" pageId="5" pageNumber="6">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is never green, and usually displays banding on the tail and a dark interorbital bar regardless of mood.
|
||
<taxonomicName id="61E14D67FFD2CE5BFCCBFC8EFC70FC03" authorityName="Echelle, Echelle & Fitch" authorityYear="1971" box="[791,976,880,904]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="9495EAF6FFD2CE5BFCCBFC8EFC70FC03" box="[791,976,880,904]" italics="true" pageId="5" pageNumber="6">Anolis carpenteri</emphasis>
|
||
</taxonomicName>
|
||
is found on the Caribbean slope at mid to low elevations. All of our collections of
|
||
<taxonomicName id="61E14D67FFD2CE5BFB06FC6FFAD2FC23" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1242,1394,912,936]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFB06FC6FFAD2FC23" box="[1242,1394,912,936]" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
are on the Pacific slope.
|
||
<taxonomicName id="61E14D67FFD2CE5BFBFFFC4EFB4AFC43" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1059,1258,944,968]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFBFFFC4EFB4AFC43" box="[1059,1258,944,968]" italics="true" pageId="5" pageNumber="6">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="61E14D67FFD2CE5BFAFFFC4FFCEDFC63" authorityName="Cope" authorityYear="1862" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="limifrons">
|
||
<emphasis id="9495EAF6FFD2CE5BFAFFFC4FFCEDFC63" italics="true" pageId="5" pageNumber="6">A. limifrons</emphasis>
|
||
</taxonomicName>
|
||
differ in male dewlap color (solid orange in
|
||
<taxonomicName id="61E14D67FFD2CE5BFAFCFC2FFCC6FB83" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFAFCFC2FFCC6FB83" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
, dirty white with or without basal yellow spot in
|
||
<taxonomicName id="61E14D67FFD2CE5BFCCBFBEFFC34FBA3" authorityName="Cope" authorityYear="1862" box="[791,916,1040,1064]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="limifrons">
|
||
<emphasis id="9495EAF6FFD2CE5BFCCBFBEFFC34FBA3" box="[791,916,1040,1064]" italics="true" pageId="5" pageNumber="6">A. limifrons</emphasis>
|
||
</taxonomicName>
|
||
). Females of these frequently sympatric species may be distinguished by limb length. The adpressed hindlimb of
|
||
<taxonomicName id="61E14D67FFD2CE5BFC78FBAFFBE1FBE3" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[932,1089,1104,1128]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFC78FBAFFBE1FBE3" box="[932,1089,1104,1128]" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
usually reaches to the ear, whereas in
|
||
<taxonomicName id="61E14D67FFD2CE5BFC49FB8FFBB3FB03" authorityName="Cope" authorityYear="1862" box="[917,1043,1136,1160]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="limifrons">
|
||
<emphasis id="9495EAF6FFD2CE5BFC49FB8FFBB3FB03" box="[917,1043,1136,1160]" italics="true" pageId="5" pageNumber="6">A. limifrons</emphasis>
|
||
</taxonomicName>
|
||
the hindlimb is longer, reaching anterior to the eye.
|
||
<taxonomicName id="61E14D67FFD2CE5BFC35FB6EFB0FFB23" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1001,1199,1168,1192]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFC35FB6EFB0FFB23" box="[1001,1199,1168,1192]" italics="true" pageId="5" pageNumber="6">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="61E14D67FFD2CE5BFB39FB6FFAE6FB23" authorityName="Ruthven" authorityYear="1916" box="[1253,1350,1169,1192]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gaigei">
|
||
<emphasis id="9495EAF6FFD2CE5BFB39FB6FFAE6FB23" box="[1253,1350,1169,1192]" italics="true" pageId="5" pageNumber="6">A. gaigei</emphasis>
|
||
</taxonomicName>
|
||
differ in the condition of the ventral scales (strongly keeled in
|
||
<taxonomicName id="61E14D67FFD2CE5BFCEEFB2FFC33FB63" authorityName="Ruthven" authorityYear="1916" box="[818,915,1233,1256]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gaigei">
|
||
<emphasis id="9495EAF6FFD2CE5BFCEEFB2FFC33FB63" box="[818,915,1233,1256]" italics="true" pageId="5" pageNumber="6">A. gaigei</emphasis>
|
||
</taxonomicName>
|
||
; smooth in
|
||
<taxonomicName id="61E14D67FFD2CE5BFBCDFB2FFB0AFB63" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1041,1194,1232,1256]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFBCDFB2FFB0AFB63" box="[1041,1194,1232,1256]" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EEFB656FFFD2CE56FCCBFAD4FAC7F95C" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="A65E36E4FFD2CE5BFCCBFAD4FC0DFACD" blockId="5.[791,941,1322,1350]" box="[791,941,1322,1350]" pageId="5" pageNumber="6">
|
||
<emphasis id="9495EAF6FFD2CE5BFCCBFAD4FC0DFACD" bold="true" box="[791,941,1322,1350]" pageId="5" pageNumber="6">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD2CE5BFCCBFA92FB39F9AF" blockId="5.[791,1395,1388,1572]" pageId="5" pageNumber="6">
|
||
The name honors the
|
||
<typeStatus id="795A8846FFD2CE5BFBDBFA93FB95FA0E" box="[1031,1077,1389,1413]" pageId="5" pageNumber="6">type</typeStatus>
|
||
locality, Parque Nacional G. D. Omar Torrijos H., and the people who have worked to maintain this wonderful forest. The park previously was named P. N. El Copé. Several
|
||
<taxonomicNameLabel id="8FA6578DFFD2CE5BFB56FA33FAABFA6F" box="[1162,1291,1484,1508]" pageId="5" pageNumber="6" rank="species">new species</taxonomicNameLabel>
|
||
of
|
||
<taxonomicName id="61E14D67FFD2CE5BFAF3FA32FAD3FA6F" authorityName="Daudin" authorityYear="1802" box="[1327,1395,1484,1508]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="9495EAF6FFD2CE5BFAF3FA32FAD3FA6F" box="[1327,1395,1484,1508]" italics="true" pageId="5" pageNumber="6">Anolis</emphasis>
|
||
</taxonomicName>
|
||
recently have been described from the park (i.e.,
|
||
<taxonomicName id="61E14D67FFD2CE5BFAEFFA12FCCBF9AF" authorityName="Hulebak, Poe, Ibanez & Williams" authorityYear="2007" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="kunayalae">
|
||
<emphasis id="9495EAF6FFD2CE5BFAEFFA12FCCBF9AF" italics="true" pageId="5" pageNumber="6">A. kunayalae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="61E14D67FFD2CE5BFCA4F9F2FC43F9AF" authorityName="Poe, Latella, Ryan & Schaad" authorityYear="2009" box="[888,995,1548,1572]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ibanezi">
|
||
<emphasis id="9495EAF6FFD2CE5BFCA4F9F2FC43F9AF" box="[888,995,1548,1572]" italics="true" pageId="5" pageNumber="6">A. ibanezi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="61E14D67FFD2CE5BFC2CF9F2FB29F9AF" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1008,1161,1548,1572]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD2CE5BFC2CF9F2FB29F9AF" box="[1008,1161,1548,1572]" italics="true" pageId="5" pageNumber="6">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD2CE5BFCCBF998FCF7F943" blockId="5.[791,1395,1638,1736]" pageId="5" pageNumber="6">
|
||
<emphasis id="9495EAF6FFD2CE5BFCCBF998FCF7F943" bold="true" pageId="5" pageNumber="6">
|
||
External description of
|
||
<typeStatus id="795A8846FFD2CE5BFBB4F998FB42F909" box="[1128,1250,1638,1666]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
(
|
||
<typeStatus id="795A8846FFD2CE5BFB2BF998FAD3F909" box="[1271,1395,1638,1666]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
|
||
variation in parentheses, measurements in mm)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFD2CE5BFCCBF910FBABF82D" blockId="5.[789,1395,1774,1958]" pageId="5" pageNumber="6">
|
||
Snout to vent length 43.5 (males to 44.6; females to 44.4); head length 0.23
|
||
<collectionCode id="C0F0AE21FFD2CE5BFC10F8F0FC5DF8AD" box="[972,1021,1806,1830]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
(0.22–0.26
|
||
<collectionCode id="C0F0AE21FFD2CE5BFBA1F8F0FB14F8AD" box="[1149,1204,1806,1830]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
); head width 0.14
|
||
<collectionCode id="C0F0AE21FFD2CE5BFCCAF8D0FCE7F8CD" box="[790,839,1838,1862]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
(0.13–0.15
|
||
<collectionCode id="C0F0AE21FFD2CE5BFC10F8D0FBA3F8CD" box="[972,1027,1838,1862]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
); femoral length 0.25
|
||
<collectionCode id="C0F0AE21FFD2CE5BFB28F8D0FA85F8CD" box="[1268,1317,1838,1862]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
(0.25– 0.28
|
||
<collectionCode id="C0F0AE21FFD2CE5BFC93F8B0FC26F8ED" box="[847,902,1870,1894]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
); ear height 0.03
|
||
<collectionCode id="C0F0AE21FFD2CE5BFB97F8B0FBDCF8ED" box="[1099,1148,1870,1894]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
(0.02–0.03
|
||
<collectionCode id="C0F0AE21FFD2CE5BFAD8F8B0FA9BF8ED" box="[1284,1339,1870,1894]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
); tail length 1.7
|
||
<collectionCode id="C0F0AE21FFD2CE5BFC4DF890FC62F80D" box="[913,962,1902,1926]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
(1.5–1.7
|
||
<collectionCode id="C0F0AE21FFD2CE5BFBEDF890FBC8F80D" box="[1073,1128,1902,1926]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
); fourth toe length 0.17
|
||
<collectionCode id="C0F0AE21FFD2CE5BFCCBF870FCE8F82D" box="[791,840,1934,1958]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
(0.13–0.17
|
||
<collectionCode id="C0F0AE21FFD2CE5BFC15F870FBA0F82D" box="[969,1024,1934,1958]" pageId="5" pageNumber="6">SVL</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<caption id="F29E666CFFD1CE58FF51F8D9FF5CF83A" ID-DOI="http://doi.org/10.5281/zenodo.11148391" ID-Zenodo-Dep="11148391" httpUri="https://zenodo.org/record/11148391/files/figure.png" pageId="6" pageNumber="7" startId="6.[141,209,1831,1853]" targetBox="[204,1360,125,1824]" targetPageId="6" targetType="figure">
|
||
<paragraph id="A65E36E4FFD1CE58FF51F8D9FF5CF83A" blockId="6.[140,1395,1831,1969]" pageId="6" pageNumber="7">
|
||
<emphasis id="9495EAF6FFD1CE58FF51F8D9FEA6F8B6" bold="true" box="[141,262,1831,1853]" pageId="6" pageNumber="7">Figure 4. A)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FED0F8D9FE1EF8B7" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[268,446,1831,1852]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD1CE58FED0F8D9FE1EF8B7" box="[268,446,1831,1852]" italics="true" pageId="6" pageNumber="7">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, El Valle de Antón, Coclé, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FC75F8D9FC61F8B6" bold="true" box="[937,961,1831,1853]" pageId="6" pageNumber="7">B)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FC1BF8D9FBD9F8B7" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[967,1145,1831,1852]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD1CE58FC1BF8D9FBD9F8B7" box="[967,1145,1831,1852]" italics="true" pageId="6" pageNumber="7">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(female dewlap, El Copé, Coclé, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FEF2F8BAFEE7F8D1" bold="true" box="[302,327,1860,1882]" pageId="6" pageNumber="7">C)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FE91F8BBFE52F8D2" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[333,498,1860,1882]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD1CE58FE91F8BBFEC3F8D1" box="[333,355,1861,1882]" italics="true" pageId="6" pageNumber="7">A.</emphasis>
|
||
cf.
|
||
<emphasis id="9495EAF6FFD1CE58FE5FF8BAFE52F8D2" box="[387,498,1860,1881]" italics="true" pageId="6" pageNumber="7">elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, south of Gamboa, Panamá, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FC3AF8BAFBA0F8D1" bold="true" box="[998,1024,1860,1882]" pageId="6" pageNumber="7">D)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FBDAF8BBFB0BF8D2" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1030,1195,1860,1882]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD1CE58FBDAF8BBFBBBF8D1" box="[1030,1051,1861,1882]" italics="true" pageId="6" pageNumber="7">A.</emphasis>
|
||
cf.
|
||
<emphasis id="9495EAF6FFD1CE58FBE0F8BAFB0BF8D2" box="[1084,1195,1860,1881]" italics="true" pageId="6" pageNumber="7">elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, Cerro Azul, Panamá, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FE58F89FFE3CF8FC" bold="true" box="[388,412,1889,1911]" pageId="6" pageNumber="7">E)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FE7EF89FFDEDF8FD" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[418,589,1889,1911]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD1CE58FE7EF89FFE17F8FD" box="[418,439,1889,1910]" italics="true" pageId="6" pageNumber="7">A.</emphasis>
|
||
cf.
|
||
<emphasis id="9495EAF6FFD1CE58FE02F89FFDEDF8FD" box="[478,589,1889,1910]" italics="true" pageId="6" pageNumber="7">elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, Lake Bayano, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FC0DF89FFC47F8FC" bold="true" box="[977,999,1889,1911]" pageId="6" pageNumber="7">F)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FC32F89FFB39F8FD" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[1006,1177,1889,1911]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD1CE58FC32F89FFC5DF8FD" box="[1006,1021,1889,1910]" italics="true" pageId="6" pageNumber="7">A</emphasis>
|
||
. cf.
|
||
<emphasis id="9495EAF6FFD1CE58FBF6F89FFB39F8FD" box="[1066,1177,1889,1910]" italics="true" pageId="6" pageNumber="7">elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, Metetí, Darién, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FEE1F880FEF8F81F" bold="true" box="[317,344,1918,1940]" pageId="6" pageNumber="7">G)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FE82F880FDBCF818" authorityName="Boulenger" authorityYear="1898" box="[350,540,1918,1940]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="9495EAF6FFD1CE58FE82F880FED4F818" box="[350,372,1918,1939]" italics="true" pageId="6" pageNumber="7">A.</emphasis>
|
||
cf.
|
||
<emphasis id="9495EAF6FFD1CE58FE45F880FDBCF818" box="[409,540,1918,1939]" italics="true" pageId="6" pageNumber="7">maculiventris</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, Yaviza, Panama).
|
||
<emphasis id="9495EAF6FFD1CE58FCBEF880FCDCF81F" bold="true" box="[866,892,1918,1940]" pageId="6" pageNumber="7">H)</emphasis>
|
||
<taxonomicName id="61E14D67FFD1CE58FC5EF880FB81F818" authorityName="Boulenger" authorityYear="1898" box="[898,1057,1918,1939]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="9495EAF6FFD1CE58FC5EF880FB81F818" box="[898,1057,1918,1939]" italics="true" pageId="6" pageNumber="7">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
(male dewlap, near Buenaventura, Colombia).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F29E666CFFD0CE59FF51FB43FEF6FB64" ID-DOI="http://doi.org/10.5281/zenodo.11148395" ID-Zenodo-Dep="11148395" httpUri="https://zenodo.org/record/11148395/files/figure.png" pageId="7" pageNumber="8" startId="7.[141,209,1213,1235]" targetBox="[188,1348,144,1201]" targetPageId="7" targetType="figure">
|
||
<paragraph id="A65E36E4FFD0CE59FF51FB43FEF6FB64" blockId="7.[141,1394,1213,1264]" pageId="7" pageNumber="8">
|
||
<emphasis id="9495EAF6FFD0CE59FF51FB43FEA6FB58" bold="true" box="[141,262,1213,1235]" pageId="7" pageNumber="8">Figure 5. A)</emphasis>
|
||
<taxonomicName id="61E14D67FFD0CE59FED0FB43FE1EFB59" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[268,446,1213,1234]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFD0CE59FED0FB43FE1EFB59" box="[268,446,1213,1234]" italics="true" pageId="7" pageNumber="8">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(male, El Copé, Coclé, Panama).
|
||
<emphasis id="9495EAF6FFD0CE59FCDDFB43FCB9FB58" bold="true" box="[769,793,1213,1235]" pageId="7" pageNumber="8">B)</emphasis>
|
||
<taxonomicName id="61E14D67FFD0CE59FCC3FB43FCD2FB58" authorityName="POE" authorityYear="1628" box="[799,882,1213,1235]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="9495EAF6FFD0CE59FCC3FB43FCD2FB58" box="[799,882,1213,1235]" italics="true" pageId="7" pageNumber="8">A. gruuo</emphasis>
|
||
</taxonomicName>
|
||
(male, Hato Chami, Chiriquí, Panama). Note bulging tail base in
|
||
<taxonomicName id="61E14D67FFD0CE59FF27FB24FEF2FB64" authorityName="POE" authorityYear="1628" box="[251,338,1242,1263]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="gruuo">A. gruuo</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A65E36E4FFD0CE59FF51FAF5FAAFF8E8" blockId="7.[140,747,1291,1923]" lastBlockId="7.[790,1395,1291,1891]" pageId="7" pageNumber="8">
|
||
Dorsal head scales multicarinate on frontal area and unito multicarinate on nasal area, mostly unicarinate in supraocular disc, some smooth scales between supraorbital semicircles and posterior to orbits (or frontal and supraocular areas nearly smooth); frontal depression present; dorsal surface of rostral scale smooth, not notched; thirteen (10–14) scales across the snout between second canthals; supraorbital semicircles distinct, separated by three (1–3) scales; three (2–5) scales separate interparietal and supraorbital semicircles; supraocular disk with some enlarged scales, bordered by a complete row of small scales; one elongate supraciliary scale followed by a series of small scales; seven (4–7) loreal rows; greater than 35 total loreals; elongate anterior nasal scale contacts sulcus between rostral and first supralabial (or nasal not greatly elongate); preoccipital absent; seven (6–9) supralabials to center of eye; six (4–7) postrostrals; 6 (4– 8) postmentals; gradually enlarged scales in supraocular disc; mental completely (or partially) divided posteriorly, extends posterolaterally beyond rostral (or nearly even with sulcus), with posterior border in concave arc (or straight line transverse to head); sublabials weakly enlarged, not much larger than medial scales; dewlap large, reaching posterior to axillae (proportionately smaller in small males; variable in females: absent or small, to axillae); six-seven rows of single scales on male dewlap; tubelike axillary pocket absent; enlarged postcloacal scales present (or absent in all females and some males); nuchal, dorsal, and caudal crests absent; dorsal scales keeled; approximately two (0–3) enlarged middorsal rows; twelve (9–15) longitudinal scale rows in 5% of
|
||
<collectionCode id="C0F0AE21FFD0CE59FCCBF995FCEDF908" box="[791,845,1643,1667]" pageId="7" pageNumber="8">SVL</collectionCode>
|
||
; ventral scales smooth, in transverse rows; eleven (8–11) longitudinal scale rows in 5% of
|
||
<collectionCode id="C0F0AE21FFD0CE59FB00F975FAB2F928" box="[1244,1298,1675,1699]" pageId="7" pageNumber="8">SVL</collectionCode>
|
||
; anterior thigh scales large, keeled, becoming smaller and smooth posteriorly; supradigitals multicarinate; toepads expanded and overlap first phalanx; fifteen (13–16) expanded lamellae under third and fourth phalanges of fourth toe (counted using the approach of Williams et al., [1995]); tail with single row of keeled middorsal scales.
|
||
</paragraph>
|
||
<caption id="F29E666CFFDFCE56FF52FC12FCD0FB8A" ID-DOI="http://doi.org/10.5281/zenodo.11148399" ID-Zenodo-Dep="11148399" box="[142,880,1004,1026]" httpUri="https://zenodo.org/record/11148399/files/figure.png" pageId="8" pageNumber="9" startId="8.[142,210,1004,1026]" targetBox="[166,1340,144,990]" targetPageId="8" targetType="figure">
|
||
<paragraph id="A65E36E4FFDFCE56FF52FC12FCD0FB8A" blockId="8.[142,880,1004,1026]" box="[142,880,1004,1026]" pageId="8" pageNumber="9">
|
||
<emphasis id="9495EAF6FFDFCE56FF52FC12FEA7FB89" bold="true" box="[142,263,1004,1026]" pageId="8" pageNumber="9">Figure 6. A)</emphasis>
|
||
Ventral and
|
||
<emphasis id="9495EAF6FFDFCE56FE5CFC12FE38FB89" bold="true" box="[384,408,1004,1026]" pageId="8" pageNumber="9">B)</emphasis>
|
||
dorsal head scales of holotype of
|
||
<taxonomicName id="61E14D67FFDFCE56FD3CFC13FCCCFB8A" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[736,876,1004,1026]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="elcopeensis">A. elcopeensis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A65E36E4FFDFCE56FF51FBDEFE32FBD4" blockId="8.[141,745,1056,1119]" pageId="8" pageNumber="9">
|
||
<emphasis id="9495EAF6FFDFCE56FF51FBDEFE32FBD4" bold="true" pageId="8" pageNumber="9">
|
||
Skeletal description (based on dry skeletons
|
||
<collectionCode id="C0F0AE21FFDFCE56FF0FFBBDFEB4FBD4" box="[211,276,1091,1119]" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" pageId="8" pageNumber="9" type="Museum">MSB</collectionCode>
|
||
95560-1)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFDFCE56FF51FB71FAB8FAB3" blockId="8.[140,747,1167,1959]" lastBlockId="8.[791,1396,1056,1336]" pageId="8" pageNumber="9">Parietal roof slightly convex, with Y-shaped parietal crests with a short stem, with no casquing, lacking crenulation on edges, with anterolateral corners flush with posterolateral edges of frontal; posterior roof of parietal slopes inferiorly; supraoccipital crests completely visible dorsally (no “half-funnel”); pineal foramen extends posteriorly into parietal, forming a U that opens at the parietal-frontal suture; dorsal skull bones smooth; postfrontal present, appears partially fused; prefrontal separated from nasal by anterior extension of frontal; frontal sutures anteriorly with nasals; no parallel crests on nasals; external nares bordered posteriorly by nasals; premaxilla dorsally nonoverlapping, laterally flush with nasals; dorsal aspect of jugal terminates on lateral or posterior surface of postorbital; posterior aspect of jugal slightly convex; epipterygoid contacts parietal dorsally; pterygoid and palatine teeth absent; lateral edge of vomer is smooth, without posteriorly directed lateral processes; maxilla extends posteriorly beyond ectopterygoid on ventral surface of skull; crest between basipterygoid processes of basisphenoid absent; lateral shelf of quadrate absent; black pigment is present on parietal only; posteriormost mandibular tooth is posterior to anterior mylohyoid foramen; splenial absent; ventral aspect of anteromedial process of coronoid juts posteriorly or slopes smoothly anteriorly; external opening of surangular foramen bordered by both dentary and surangular; posterior suture of dentary blunt or pronged; anteriormost aspect of posterior border of dentary is well within mandibular fossa; labial process of coronoid present; coronoid does not extend posterolaterally beyond surangular foramen; jaw sculpturing absent; angular absent; teeth unicuspid anteriorly, tricuspid posteriorly; angular process of articular present; ten premaxillary tooth positions.</paragraph>
|
||
<paragraph id="A65E36E4FFDFCE56FCCBFA84FB8BFA32" blockId="8.[791,1395,1402,1465]" pageId="8" pageNumber="9">
|
||
<emphasis id="9495EAF6FFDFCE56FCCBFA84FB8BFA32" bold="true" pageId="8" pageNumber="9">Color in life (adapted from field notes and color photos by SP)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFDFCE56FCCBFA21FAC7F95C" blockId="8.[791,1395,1503,1751]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="61E14D67FFDFCE56FCCBFA21FC40FA7C" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[791,992,1503,1527]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDFCE56FCCBFA21FC40FA7C" box="[791,992,1503,1527]" italics="true" pageId="8" pageNumber="9">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is a brown or brownish-gray lizard. Its dorsum may be marked with faint lateral banding, rows of spotting, or striations (
|
||
<figureCitation id="3EDA2A61FFDFCE56FBBCF9E1FB13F9BC" box="[1120,1203,1567,1591]" captionStart="Figure 3" captionStartId="4.[141,209,1896,1918]" captionTargetBox="[198,1353,144,1883]" captionTargetId="figure-23@4.[141,1394,141,1886]" captionTargetPageId="4" captionText="Figure 3. A) Anolis elcopeensis (female, El Copé, Coclé, Panama).B) A. cf elcopeensis (male, south of Gamboa, Panamá, Panama). C) A. cf maculiventris (female, Yaviza, Darién, Panama)." figureDoi="http://doi.org/10.5281/zenodo.11148381" httpUri="https://zenodo.org/record/11148381/files/figure.png" pageId="8" pageNumber="9">Fig. 3A</figureCitation>
|
||
) or appear lightly speckled (
|
||
<figureCitation id="3EDA2A61FFDFCE56FC54F9C1FC42F9DC" box="[904,994,1599,1623]" captionStart="Figure 5" captionStartId="7.[141,209,1213,1235]" captionTargetBox="[188,1348,144,1201]" captionTargetId="figure-19@7.[141,1394,141,1205]" captionTargetPageId="7" captionText="Figure 5. A) Anolis elcopeensis (male, El Copé, Coclé, Panama). B) A. gruuo (male, Hato Chami, Chiriquí, Panama). Note bulging tail base in A. gruuo." figureDoi="http://doi.org/10.5281/zenodo.11148395" httpUri="https://zenodo.org/record/11148395/files/figure.png" pageId="8" pageNumber="9">Fig. 5A</figureCitation>
|
||
), or may be nearly patternless pale gray or dark brown. The tail usually is banded, and there usually is a dark dorsal interorbital bar (
|
||
<figureCitation id="3EDA2A61FFDFCE56FB06F981FA93F91C" box="[1242,1331,1663,1687]" captionStart="Fig" captionStartId="1.[141,176,1885,1907]" captionTargetBox="[177,1330,814,1871]" captionTargetId="figure-44@1.[141,1394,802,1878]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic estimate of placement of A. elcopeensis sp. nov. based on Bayesian analysis of morphological and molecular data. Numbers are clade credibility values." figureDoi="http://doi.org/10.5281/zenodo.11148377" httpUri="https://zenodo.org/record/11148377/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
|
||
). The ventral body usually displays dark brown markings on white background. The iris is brown. The throat is pale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EEFB656FFFDFCE57FCCBF8F2FD06F8E8" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="A65E36E4FFDFCE56FCCBF8F2FB6CF8A3" blockId="8.[791,1228,1804,1832]" box="[791,1228,1804,1832]" pageId="8" pageNumber="9">
|
||
<emphasis id="9495EAF6FFDFCE56FCCBF8F2FB6CF8A3" bold="true" box="[791,1228,1804,1832]" pageId="8" pageNumber="9">Distribution and natural history</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A65E36E4FFDFCE57FCCBF8B0FE96FA08" blockId="8.[791,1395,1870,1958]" lastBlockId="9.[141,747,1067,1891]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<taxonomicName id="61E14D67FFDFCE56FCCBF8B0FC43F8ED" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[791,995,1870,1894]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDFCE56FCCBF8B0FC43F8ED" box="[791,995,1870,1894]" italics="true" pageId="8" pageNumber="9">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
is known from
|
||
<collectingRegion id="6425F806FFDFCE56FB7AF8B0FAF1F8ED" box="[1190,1361,1870,1894]" country="Panama" name="Cocle" pageId="8" pageNumber="9">Coclé Province</collectingRegion>
|
||
in
|
||
<collectingCountry id="DEF67674FFDFCE56FCCBF891FCCBF80D" box="[791,875,1903,1926]" name="Panama" pageId="8" pageNumber="9">Panama</collectingCountry>
|
||
and potentially from
|
||
<collectingCountry id="DEF67674FFDFCE56FB86F890FB0EF80D" box="[1114,1198,1902,1926]" name="Panama" pageId="8" pageNumber="9">Panamá</collectingCountry>
|
||
and
|
||
<collectingRegion id="6425F806FFDFCE56FB3AF890FA8FF80D" box="[1254,1327,1902,1926]" country="Panama" name="Darien" pageId="8" pageNumber="9">Darién</collectingRegion>
|
||
Provinces (
|
||
<figureCitation id="3EDA2A61FFDFCE56FCBDF870FC08F82D" box="[865,936,1934,1958]" captionStart="Figure 7" captionStartId="9.[141,209,899,921]" captionTargetBox="[141,1394,144,888]" captionTargetId="figure-249@9.[139,1398,141,891]" captionTargetPageId="9" captionText="Figure 7. Map of eastern Panama showing type locality of Anolis elcopeensis (star; sampled for both morphology and COI), localities sampled for COI (white circles), and localities for additional specimens referenced in text (black circles). Closed circles and star indicate localities for confirmed A. elcopeensis according to morphological and molecular comparisons. Open circles indicate localities for specimens referred to as Anolis cf. elcopeensis that may represent A. elcopeensis or undescribed species. Square marks locality for A. maculiventris or a similar species (see text).Appendix 2 lists voucher specimens for localities." figureDoi="http://doi.org/10.5281/zenodo.11148401" httpUri="https://zenodo.org/record/11148401/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
|
||
; see below). These provinces have been well surveyed for herpetofauna, however much of
|
||
<collectingRegion id="6425F806FFDECE57FD77FBD5FD49FBC8" box="[683,745,1067,1091]" country="Panama" name="Cocle" pageId="9" pageNumber="10">Coclé</collectingRegion>
|
||
is difficult to access and remains unexplored. Habitat in this province ranges from lowland rainforest in the Canal Zone up to cool tropical premontane rainforest in Parqué Nacional Omar Torrijos. This range spans an elevation gradient from sea level up to over 1,000 meters. We have collected topotypical and paratopotypical
|
||
<taxonomicName id="61E14D67FFDECE57FD83FB15FD4AFA88" box="[607,746,1259,1283]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="copeensis">
|
||
<emphasis id="9495EAF6FFDECE57FD83FB15FD4AFA88" box="[607,746,1259,1283]" italics="true" pageId="9" pageNumber="10">A. copeensis</emphasis>
|
||
</taxonomicName>
|
||
from 245 to 801 meters. We have found
|
||
<taxonomicName id="61E14D67FFDECE57FDEAFAF5FD6EFAA8" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[566,718,1291,1315]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FDEAFAF5FD6EFAA8" box="[566,718,1291,1315]" italics="true" pageId="9" pageNumber="10">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
in near-pristine primary and old selectively logged forests (i.e., at the
|
||
<typeStatus id="795A8846FFDECE57FED7FAB5FE99FAE8" box="[267,313,1355,1379]" pageId="9" pageNumber="10">type</typeStatus>
|
||
locality) and in heavily disturbed roadside vegetation.
|
||
</paragraph>
|
||
<caption id="F29E666CFFDECE57FF51FC7DFB03FB87" ID-DOI="http://doi.org/10.5281/zenodo.11148401" ID-Zenodo-Dep="11148401" httpUri="https://zenodo.org/record/11148401/files/figure.png" pageId="9" pageNumber="10" startId="9.[141,209,899,921]" targetBox="[141,1394,144,888]" targetPageId="9" targetType="figure">
|
||
<paragraph id="A65E36E4FFDECE57FF51FC7DFB03FB87" blockId="9.[141,1395,899,1036]" pageId="9" pageNumber="10">
|
||
<emphasis id="9495EAF6FFDECE57FF51FC7DFF49FC13" bold="true" box="[141,233,899,921]" pageId="9" pageNumber="10">Figure 7.</emphasis>
|
||
Map of eastern Panama showing type locality of
|
||
<taxonomicName id="61E14D67FFDECE57FD1CFC7DFCD2FC13" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[704,882,899,920]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FD1CFC7DFCD2FC13" box="[704,882,899,920]" italics="true" pageId="9" pageNumber="10">Anolis elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
(star; sampled for both morphology and COI), localities sampled for COI (white circles), and localities for additional specimens referenced in text (black circles). Closed circles and star indicate localities for confirmed
|
||
<taxonomicName id="61E14D67FFDECE57FE2CFC43FDDBFC59" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[496,635,957,978]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FE2CFC43FDDBFC59" box="[496,635,957,978]" italics="true" pageId="9" pageNumber="10">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
according to morphological and molecular comparisons. Open circles indicate localities for specimens referred to as
|
||
<taxonomicName id="61E14D67FFDECE57FE29FC27FD65FC65" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[501,709,985,1007]" class="Squamata" family="Dactyloidae" genus="Anolis" isUncertain="true" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FE29FC27FD92FC65" box="[501,562,985,1006]" italics="true" pageId="9" pageNumber="10">Anolis</emphasis>
|
||
cf.
|
||
<emphasis id="9495EAF6FFDECE57FD8AFC27FD65FC65" box="[598,709,985,1006]" italics="true" pageId="9" pageNumber="10">elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
that may represent
|
||
<taxonomicName id="61E14D67FFDECE57FCA3FC24FBA8FC65" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[895,1032,985,1007]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FCA3FC24FBA8FC65" box="[895,1032,985,1007]" italics="true" pageId="9" pageNumber="10">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
or undescribed species. Square marks locality for
|
||
<taxonomicName id="61E14D67FFDECE57FF21FC08FE3BFB80" authorityName="Boulenger" authorityYear="1898" box="[253,411,1014,1035]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="9495EAF6FFDECE57FF21FC08FE3BFB80" box="[253,411,1014,1035]" italics="true" pageId="9" pageNumber="10">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
or a similar species (see text). Appendix 2 lists voucher specimens for localities.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A65E36E4FFDECE57FF71FA75FD06F8E8" blockId="9.[141,747,1067,1891]" pageId="9" pageNumber="10">
|
||
All of our collections of
|
||
<taxonomicName id="61E14D67FFDECE57FE1BFA75FDC4FA28" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" box="[455,612,1419,1443]" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FE1BFA75FDC4FA28" box="[455,612,1419,1443]" italics="true" pageId="9" pageNumber="10">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
occurred at night when anoles sleep. Among 35 recorded observations at El Copé, mean sleeping perch height was
|
||
<quantity id="61199B01FFDECE57FD67FA35FF02F989" metricMagnitude="0" metricUnit="m" metricValue="4.15" pageId="9" pageNumber="10" unit="m" value="4.15">4.15 m</quantity>
|
||
(standard deviation 2.35). Among 30 individuals for which sleeping perch
|
||
<typeStatus id="795A8846FFDECE57FE5BF9F5FE15F9A8" box="[391,437,1547,1571]" pageId="9" pageNumber="10">type</typeStatus>
|
||
was recorded, 24 were on twigs, three were on leaves, and three were on vines. Thus, among perches easily surveyable by humans (i.e., excluding high canopy, burrows, etc.), narrow perches that are relatively high seem to be preferred by
|
||
<taxonomicName id="61E14D67FFDECE57FD70F975FF55F948" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FD70F975FF55F948" italics="true" pageId="9" pageNumber="10">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
. Diurnally, the species has been observed to be active on the ground and on vegetation at heights up to
|
||
<quantity id="61199B01FFDECE57FF70F915FF4AF889" box="[172,234,1771,1795]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="m" value="1.5">1.5 m</quantity>
|
||
(Mason Ryan, pers. comm.). On
|
||
<date id="D25F1024FFDECE57FD8BF915FD47F888" box="[599,743,1771,1795]" pageId="9" pageNumber="10" value="2002-07-23">23 July 2002</date>
|
||
, Ryan observed a Cocoa Woodcreeper (
|
||
<taxonomicName id="61E14D67FFDECE57FD94F8F5FF56F8C8" baseAuthorityName="Jardine" baseAuthorityYear="1847" class="Aves" family="Furnariidae" genus="Xiphorhynchus" kingdom="Animalia" order="Passeriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="susurrans">
|
||
<emphasis id="9495EAF6FFDECE57FD94F8F5FF56F8C8" italics="true" pageId="9" pageNumber="10">Xiphorhynchus susurrans</emphasis>
|
||
</taxonomicName>
|
||
) catching and consuming an adult male
|
||
<taxonomicName id="61E14D67FFDECE57FD6CF8D5FF55F8E8" authorityName="Poe & Scarpetta & Schaad Abstract.-We & Central & Anolis & We" authorityYear="2015" class="Squamata" family="Dactyloidae" genus="Anolis" kingdom="Animalia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="9495EAF6FFDECE57FD6CF8D5FF55F8E8" italics="true" pageId="9" pageNumber="10">A. elcopeensis</emphasis>
|
||
</taxonomicName>
|
||
that was displaying from a tree buttress.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |