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<document id="0F85E00EA47629ACC05EAD2CE154AE00" ID-DOI="10.5281/zenodo.278201" ID-GBIF-Dataset="878ed78c-199d-465c-a2dc-e6fbdc65608a" ID-ISSN="1175-5326" ID-Zenodo-Dep="278201" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460319145060" checkinUser="plazi" docAuthor="Gutiérrez-Fonseca, Pablo E. &amp; Springer, Monika" docDate="2011" docId="EE0EC308163E2477FF35FD98FC2EFD78" docLanguage="en" docName="zt02965p038.pdf" docOrigin="Zootaxa 2965" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anacroneuria marca Stark 1998" docType="treatment" docVersion="7" lastPageNumber="30" masterDocId="1237BB7016332479FFA2FFF4FFD5FFFC" masterDocTitle="Description of the final instar nymphs of seven species from Anacroneuria Klapálek (Plecoptera: Perlidae) in Costa Rica, and first record for an additional genus in Central America" masterLastPageNumber="38" masterPageNumber="16" pageNumber="29" updateTime="1698255736077" updateUser="plazi">
<mods:mods id="A9B4C57B36A70C2CFF68BB695195B4A9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="77E37D457DA5AB929A266ECDE146B3F0">
<mods:title id="50830C2B2058CD425E3F4C56A9C223B9">Description of the final instar nymphs of seven species from Anacroneuria Klapálek (Plecoptera: Perlidae) in Costa Rica, and first record for an additional genus in Central America</mods:title>
</mods:titleInfo>
<mods:name id="9861677007F295726E389B4E4EB613B7" type="personal">
<mods:role id="ACFBB5B8BEB93921198304E6EF509ABB">
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</mods:role>
<mods:namePart id="6EFFBBA8D1009A6F46482B4D420AA49F">Gutiérrez-Fonseca, Pablo E.</mods:namePart>
</mods:name>
<mods:name id="D2E5965EEC57221849B48326306B61F3" type="personal">
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<mods:namePart id="84878F1BAD2FE685E0439DEF07FA7961">Springer, Monika</mods:namePart>
</mods:name>
<mods:typeOfResource id="D0B9E5E7654E774E959D05EA8D21DEF7">text</mods:typeOfResource>
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<mods:title id="8CF9D600780AA1ADD4C6C9E83333F8C6">Zootaxa</mods:title>
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<mods:date id="B29318577604C001E78A6648344E71A0">2011</mods:date>
<mods:detail id="A4118D5D87E874D1230AABE434267E8B" type="volume">
<mods:number id="0B96F8F8DB22186FFE65DA7E00BE5C22">2965</mods:number>
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<mods:classification id="FE1857E0201ABD0F346E6DC41D5A7CD6">journal article</mods:classification>
<mods:identifier id="F373B003CACC6117DB5589F70BCE739F" type="DOI">10.5281/zenodo.278201</mods:identifier>
<mods:identifier id="C4799CF08380742E5B7C431E44A5B705" type="GBIF-Dataset">878ed78c-199d-465c-a2dc-e6fbdc65608a</mods:identifier>
<mods:identifier id="4FD81A9FC4AE7EC087679F3FDDB0C4A5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="64D598DF075949F6FAA782D0FCC47AD1" type="Zenodo-Dep">278201</mods:identifier>
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<treatment id="EE0EC308163E2477FF35FD98FC2EFD78" ID-DOI="http://doi.org/10.5281/zenodo.5658115" ID-GBIF-Taxon="119402066" ID-Zenodo-Dep="5658115" LSID="urn:lsid:plazi:treatment:EE0EC308163E2477FF35FD98FC2EFD78" httpUri="http://treatment.plazi.org/id/EE0EC308163E2477FF35FD98FC2EFD78" lastPageId="14" lastPageNumber="30" pageId="13" pageNumber="29">
<subSubSection id="2EBD2195163E2474FF35FD98FEC9FD5A" pageId="13" pageNumber="29" type="nomenclature">
<paragraph id="6618721E163E2474FF35FD98FDFCFD79" blockId="13.[151,553,619,678]" box="[151,553,619,646]" pageId="13" pageNumber="29">
<heading id="3D50C572163E2474FF35FD98FDFCFD79" bold="true" box="[151,553,619,646]" fontSize="11" level="1" pageId="13" pageNumber="29" reason="1">
<taxonomicName id="A1A7099D163E2474FF35FD98FDFCFD79" ID-CoL="DC2D" authority="Stark, 1998" authorityName="Stark" authorityYear="1998" box="[151,553,619,646]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="29" phylum="Arthropoda" rank="species" species="marca">
<emphasis id="54D3AE0C163E2474FF35FD98FDFCFD79" bold="true" box="[151,553,619,646]" pageId="13" pageNumber="29">
<emphasis id="54D3AE0C163E2474FF35FD98FE41FD7A" bold="true" box="[151,404,620,646]" italics="true" pageId="13" pageNumber="29">Anacroneuria marca</emphasis>
<bibRefCitation id="02360FEF163E2474FE39FD9FFDFCFD79" author="Stark" box="[411,553,619,645]" pageId="13" pageNumber="29" refString="Stark, B. P. (1998) The Anacroneuria of Costa Rica and Panama (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 111, 551 - 603." type="journal article" year="1998">Stark, 1998</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="6618721E163E2474FF35FD7AFEC9FD5A" blockId="13.[151,553,619,678]" box="[151,284,654,678]" pageId="13" pageNumber="29">
<figureCitation id="FE9C6E9B163E2474FF35FD7AFEC9FD5A" box="[151,284,654,678]" captionStart="FIGURES 16 − 17" captionStartId="14.[151,264,1875,1898]" captionTargetBox="[247,1348,752,1784]" captionTargetId="figure@14.[151,1436,700,1855]" captionTargetPageId="14" captionText="FIGURES 16 − 17. Anacroneuria marca, 16. Head and thorax (pro, meso and meta notum); 17. Foreleg." httpUri="https://zenodo.org/record/278211/files/figure.png" pageId="13" pageNumber="29">Figs. 16–17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2EBD2195163E2474FF35FD22FBBFFBCE" pageId="13" pageNumber="29" type="description">
<paragraph id="6618721E163E2474FF35FD22FBBFFBCE" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="29">
<emphasis id="54D3AE0C163E2474FF35FD22FEFCFD13" bold="true" box="[151,297,726,751]" pageId="13" pageNumber="29">Description.</emphasis>
<emphasis id="54D3AE0C163E2474FE92FD22FE62FD12" box="[304,439,726,750]" italics="true" pageId="13" pageNumber="29">Body length</emphasis>
:
<quantity id="A15FDFFB163E2474FE64FD22FD96FD12" box="[454,579,726,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="13" pageNumber="29" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
.
<emphasis id="54D3AE0C163E2474FDECFD22FD5EFD12" box="[590,651,726,750]" italics="true" pageId="13" pageNumber="29">Head</emphasis>
(
<figureCitation id="FE9C6E9B163E2474FD39FD22FD25FD12" box="[667,752,726,750]" captionStart="FIGURES 16 − 17" captionStartId="14.[151,264,1875,1898]" captionTargetBox="[247,1348,752,1784]" captionTargetId="figure@14.[151,1436,700,1855]" captionTargetPageId="14" captionText="FIGURES 16 − 17. Anacroneuria marca, 16. Head and thorax (pro, meso and meta notum); 17. Foreleg." httpUri="https://zenodo.org/record/278211/files/figure.png" pageId="13" pageNumber="29">Fig. 16</figureCitation>
): brown forward of ocelli, with exception of pale m-line and pale anterior frontclypeus; often with clear spots on the outer side of the ocelli.
<emphasis id="54D3AE0C163E2474FB99FD08FB7FFCEF" box="[1083,1194,764,787]" italics="true" pageId="13" pageNumber="29">Pronotum</emphasis>
: oval, narrower than mesonotum; brown with pale, indistinct longitudinal areas forming a median stripe. Surface covered with fine short setae, lacking bristles. Margins completely fringed with thick bristles, somewhat variable in length.
<emphasis id="54D3AE0C163E2474FA80FCB0FEC2FC83" italics="true" pageId="13" pageNumber="29">Meso- and Metanotum</emphasis>
: pale irregular spots around midline; wing pads brown. Dorsal surfaces covered with fine short setae, and scattered thick bristles; well-defined anterior and lateral lines of thick bristles. Margins completely fringed with thick bristles, somewhat variable in length.
<emphasis id="54D3AE0C163E2474FD60FC5AFCCCFC3A" box="[706,793,942,966]" italics="true" pageId="13" pageNumber="29">Foreleg</emphasis>
(
<figureCitation id="FE9C6E9B163E2474FC88FC5AFCAAFC3A" box="[810,895,942,966]" captionStart="FIGURES 16 − 17" captionStartId="14.[151,264,1875,1898]" captionTargetBox="[247,1348,752,1784]" captionTargetId="figure@14.[151,1436,700,1855]" captionTargetPageId="14" captionText="FIGURES 16 − 17. Anacroneuria marca, 16. Head and thorax (pro, meso and meta notum); 17. Foreleg." httpUri="https://zenodo.org/record/278211/files/figure.png" pageId="13" pageNumber="29">Fig. 17</figureCitation>
): dorsal surfaces covered with fine, short setae, except for hairless median bar. Posterior margin with very dense fringe of long setae. Femur dorsally without welldefined transverse row of thick bristles; short and medium-sized spines on dorsal surface, forming an arch that extends to 2/3 length of femur; basal posterior patch of very small spines along margin.
</paragraph>
</subSubSection>
<subSubSection id="2EBD2195163E2474FF67FBCAFBFFFB3E" pageId="13" pageNumber="29" type="distribution">
<paragraph id="6618721E163E2474FF67FBCAFE32FBAA" blockId="13.[151,1437,726,2011]" box="[197,487,1086,1111]" pageId="13" pageNumber="29">
<emphasis id="54D3AE0C163E2474FF67FBCAFE8BFBAB" bold="true" box="[197,350,1086,1111]" pageId="13" pageNumber="29">Distribution.</emphasis>
<collectingCountry id="1EB0328E163E2474FEC7FBCAFE36FBAA" box="[357,483,1086,1110]" name="Costa Rica" pageId="13" pageNumber="29">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="6618721E163E2474FF67FB97FBFFFB3E" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="29">
<emphasis id="54D3AE0C163E2474FF67FB97FE75FB87" bold="true" box="[197,416,1122,1147]" pageId="13" pageNumber="29">Local distribution</emphasis>
(
<figureCitation id="FE9C6E9B163E2474FE13FB96FDDDFB86" box="[433,520,1122,1146]" captionStart="FIGURE 18" captionStartId="15.[151,250,1513,1536]" captionTargetBox="[203,1410,223,1476]" captionTargetId="figure@15.[151,1436,193,1491]" captionTargetPageId="15" captionText="FIGURE 18. Map of Costa Rica showing distribution of Anacroneuria marca." httpUri="https://zenodo.org/record/278212/files/figure.png" pageId="13" pageNumber="29">Fig. 18</figureCitation>
): Lowland and middle elevation streams on both Pacific and Caribbean slopes. Inhabits streams across a wide range of altitudes, reaching up to 2700 masl. Together with
<taxonomicName id="A1A7099D163E2474FB31FB72FAE5FB62" box="[1171,1328,1158,1182]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="29" phylum="Arthropoda" rank="species" species="holzenthali">
<emphasis id="54D3AE0C163E2474FB31FB72FAE5FB62" box="[1171,1328,1158,1182]" italics="true" pageId="13" pageNumber="29">A. holzenthali</emphasis>
</taxonomicName>
, this species has one of the most widespread local distributions of all Costa Rican species.
</paragraph>
</subSubSection>
<subSubSection id="2EBD2195163E2477FF67FB3BFDBBFDEB" lastPageId="14" lastPageNumber="30" pageId="13" pageNumber="29" type="materials_examined">
<paragraph id="6618721E163E2474FF67FB3BFC33F9FA" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="29">
<emphasis id="54D3AE0C163E2474FF67FB3BFDF4FB1B" bold="true" box="[197,545,1230,1255]" pageId="13" pageNumber="29">Reared specimens. San José:</emphasis>
Río Savegre, San Gerardo de Dota,
<quantity id="A15FDFFB163E2474FC6AFB3AFBF4FB1A" box="[968,1057,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="29" unit="m" value="2700.0">2700 m</quantity>
,
<date id="121954DE163E2474FB8DFB3AFB75FB1A" box="[1071,1184,1230,1254]" pageId="13" pageNumber="29" value="2008-05-25">25.v.2008</date>
, Gutiérrez, 1 Ƥ; Quebrada sin nombre, Cerro de la Muerte, Santa María de Dota,
<quantity id="A15FDFFB163E2474FC9FFB06FC41FAF6" box="[829,916,1266,1290]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="29" unit="m" value="2200.0">2200 m</quantity>
,
<date id="121954DE163E2474FC02FB06FBDFFAF6" box="[928,1034,1266,1290]" pageId="13" pageNumber="29" value="2008-04-06">6.iv.2008</date>
, Gutiérrez, 1 3; Quebrada sin nombre, Santa María de Dota,
<quantity id="A15FDFFB163E2474FE68FAE2FDF1FAD2" box="[458,548,1302,1326]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="29" unit="m" value="2700.0">2700 m</quantity>
,
<date id="121954DE163E2474FD90FAE2FD48FAD2" box="[562,669,1302,1326]" pageId="13" pageNumber="29" value="2008-04-06">6.iv.2008</date>
, Gutiérrez, 1 3; Quebrada Cascada Bromelia, Cerro de la Muerte, Paraíso de los Quetzales,
<quantity id="A15FDFFB163E2474FE1BFACEFDC4FAAE" box="[441,529,1338,1362]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="29" unit="m" value="2700.0">2700 m</quantity>
,
<date id="121954DE163E2474FDBFFACEFD4AFAAE" box="[541,671,1338,1362]" pageId="13" pageNumber="29" value="2008-12-22">22.xii.2008</date>
, Gutiérrez, 3 3;
<emphasis id="54D3AE0C163E2474FCFCFACEFC18FAAF" bold="true" box="[862,973,1338,1363]" pageId="13" pageNumber="29">
<taxonomicName id="A1A7099D163E2474FCFCFACEFC12FAAF" box="[862,967,1338,1363]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="17" phylum="Arthropoda" rank="species" species="alajuela">Alajuela</taxonomicName>
:
</emphasis>
Río Niño, Upala,
<date id="121954DE163E2474FB3FFACEFADCFAAE" box="[1181,1289,1338,1362]" pageId="13" pageNumber="29" value="2008-11-09">9.xi.2008</date>
, Gutiérrez, 1 3, 1 Ƥ; Quebrada Rugama, Nuevo Arenal, Tilarán,
<quantity id="A15FDFFB163E2474FD41FAAAFCF9FA8A" box="[739,812,1374,1398]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="13" pageNumber="29" unit="m" value="550.0">550 m</quantity>
,
<date id="121954DE163E2474FC9BFAAAFC66FA8A" box="[825,947,1374,1398]" pageId="13" pageNumber="29" value="2008-09-20">20.ix.2008</date>
, Gutiérrez, 1 3; Río La Vieja,
<collectingCountry id="1EB0328E163E2474FAB1FAABFAE2FA8A" box="[1299,1335,1375,1398]" name="Philippines" pageId="13" pageNumber="29">PH</collectingCountry>
Chocosuela above dam, San Carlos,
<quantity id="A15FDFFB163E2474FE7CFA76FDF2FA66" box="[478,551,1410,1434]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="29" unit="m" value="730.0">730 m</quantity>
,
<date id="121954DE163E2474FD91FA76FD74FA66" box="[563,673,1410,1434]" pageId="13" pageNumber="29" value="2001-03-09">9.iii.2001</date>
, Gutiérrez, 8 3, 3 Ƥ; Río San Lorencito, Res. Biol. Alberto Manuel Brenes, San Ramón,
<quantity id="A15FDFFB163E2474FE2FFA52FE0CFA42" box="[397,473,1446,1470]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="13" pageNumber="29" unit="m" value="860.0">860 m</quantity>
,
<date id="121954DE163E2474FE4AFA52FDEDFA42" box="[488,568,1446,1470]" pageId="13" pageNumber="29" value="2008-10">x.2008</date>
, Gutiérrez, 4 Ƥ;
<emphasis id="54D3AE0C163E2474FCA0FA52FCBEFA43" bold="true" box="[770,875,1446,1471]" pageId="13" pageNumber="29">Heredia:</emphasis>
Río Sarapiquí, Res. Biol. La Tirimbina,
<quantity id="A15FDFFB163E2474FAE9FA52FA42FA42" box="[1355,1431,1446,1470]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="29" unit="m" value="180.0">180 m</quantity>
,
<date id="121954DE163E2474FF35FA3EFECDFA1E" box="[151,280,1482,1506]" pageId="13" pageNumber="29" value="2008-07-11">11.vii.2008</date>
, Gutiérrez, 3 3;
<emphasis id="54D3AE0C163E2474FE42FA3EFDECFA1F" bold="true" box="[480,569,1482,1507]" pageId="13" pageNumber="29">Limón:</emphasis>
Río Dos Novillos, Guápiles, several months 2006, Gutiérrez, 24 3, 25 Ƥ;
<emphasis id="54D3AE0C163E2474FF35FA1BFEFDF9FB" bold="true" box="[151,296,1519,1543]" pageId="13" pageNumber="29">Puntarenas:</emphasis>
Río Barranca, Esparza,
<quantity id="A15FDFFB163E2474FD9BFA1AFD54F9FA" box="[569,641,1518,1542]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="29" unit="m" value="450.0">450 m</quantity>
,
<date id="121954DE163E2474FD2FFA1AFD2CF9FA" box="[653,761,1518,1542]" pageId="13" pageNumber="29" value="2007-01-17">17.i.2007</date>
, Gutiérrez, 3 3,
<specimenCount id="70A1B997163E2474FC17FA1AFC16F9FA" box="[949,963,1518,1542]" pageId="13" pageNumber="29" type="generic">2</specimenCount>
Ƥ.
</paragraph>
<paragraph id="6618721E163E2477FF67F9E6FCBAFE33" blockId="13.[151,1437,726,2011]" lastBlockId="14.[151,1436,151,644]" lastPageId="14" lastPageNumber="30" pageId="13" pageNumber="29">
<emphasis id="54D3AE0C163E2474FF67F9E6FD7CF9D7" bold="true" box="[197,681,1554,1579]" pageId="13" pageNumber="29">Additional material examined. San José:</emphasis>
Quebrada Cascajal, primary forest, Coronado,
<quantity id="A15FDFFB163E2474FB19F9E6FAC4F9D6" box="[1211,1297,1554,1578]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="13" pageNumber="29" unit="m" value="1750.0">1750 m</quantity>
,
<date id="121954DE163E2474FABFF9E6FA4DF9D6" box="[1309,1432,1554,1578]" pageId="13" pageNumber="29" value="2003-09-28">28.ix.2003</date>
, S. Amador, 3 3, 4 Ƥ; Bajo la Hondura,
<quantity id="A15FDFFB163E2474FDC3F9C2FD6CF9B2" box="[609,697,1590,1614]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="29" unit="m" value="1200.0">1200 m</quantity>
,
<date id="121954DE163E2474FD64F9C2FC96F9B2" box="[710,835,1590,1614]" pageId="13" pageNumber="29" value="1990-03-30">30.iii.1990</date>
, W. Flowers, 3 Ƥ;
<emphasis id="54D3AE0C163E2474FBBEF9C2FB5FF9B3" bold="true" box="[1052,1162,1590,1615]" pageId="13" pageNumber="29">
<taxonomicName id="A1A7099D163E2474FBBEF9C2FB50F9B3" box="[1052,1157,1590,1615]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="17" phylum="Arthropoda" rank="species" species="alajuela">Alajuela</taxonomicName>
:
</emphasis>
Río San Lorencito, San Ramón, Res. Biol. Alberto Manuel Brenes,
<quantity id="A15FDFFB163E2474FD25F9AEFD04F98E" box="[647,721,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="29" unit="m" value="850.0">850 m</quantity>
,
<date id="121954DE163E2474FD7CF9AEFC84F98E" box="[734,849,1626,1650]" pageId="13" pageNumber="29" value="2003-10-18">18.x.2003</date>
, F. Ruiz, 1 3; same,
<date id="121954DE163E2474FBE3F9AEFB66F98E" box="[1089,1203,1626,1650]" pageId="13" pageNumber="29" value="2005-10-15">15.x.2005</date>
, M.J. Quesada, 1 Ƥ; same,
<date id="121954DE163E2474FF43F98AFE86F96A" box="[225,339,1662,1686]" pageId="13" pageNumber="29" value="1998-10-10">10.x.1998</date>
, UCR students, 1 Ƥ; same,
<date id="121954DE163E2474FD35F98AFCC4F96A" box="[663,785,1662,1686]" pageId="13" pageNumber="29" value="2005-09-24">24.ix.2005</date>
, M. Alfaro, 3 Ƥ; same,
<date id="121954DE163E2474FB8AF98AFB7EF96A" box="[1064,1195,1662,1686]" pageId="13" pageNumber="29" value="2000-12-30">30.xii.2000</date>
, PennState Univ students, 1 3; same,
<date id="121954DE163E2474FEC3F956FE08F946" box="[353,477,1698,1722]" pageId="13" pageNumber="29" value="1990-03-11">11.iii.1990</date>
, W. Flowers, 2 3, 1 Ƥ; Río Chachagua, San Carlos,
<quantity id="A15FDFFB163E2474FB8EF956FBA0F946" box="[1068,1141,1698,1722]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="29" unit="m" value="200.0">200 m</quantity>
,
<date id="121954DE163E2474FB23F956FB29F946" box="[1153,1276,1698,1722]" pageId="13" pageNumber="29" value="2003-09-19">19.ix.2003</date>
, R. Lara, 1 3; Río Peñas Blancas, San Carlos,
<quantity id="A15FDFFB163E2474FDA2F932FD9FF922" box="[512,586,1734,1758]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="29" unit="m" value="260.0">260 m</quantity>
,
<date id="121954DE163E2474FDF4F932FD1FF922" box="[598,714,1734,1758]" pageId="13" pageNumber="29" value="1998-12-08">8.xii.1998</date>
, Springer &amp; P. Paaby,
<date id="121954DE163E2474FC65F932FBC9F922" box="[967,1052,1734,1759]" pageId="13" pageNumber="29" value="2010-04-03">4 3, 10</date>
Ƥ; Río La Vieja,
<collectingCountry id="1EB0328E163E2474FB45F933FADEF922" box="[1255,1291,1735,1758]" name="Philippines" pageId="13" pageNumber="29">PH</collectingCountry>
Chocosuela, San Carlos,
<quantity id="A15FDFFB163E2474FE8AF91EFEA3F8FE" box="[296,374,1770,1794]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="13" pageNumber="29" unit="m" value="730.0">730 m</quantity>
,
<date id="121954DE163E2474FE24F91EFE2FF8FE" box="[390,506,1770,1794]" pageId="13" pageNumber="29" value="2001-12-04">4.xii.2001</date>
, Springer, 4 3, 4 Ƥ; same,
<date id="121954DE163E2474FCEEF91EFC13F8FE" box="[844,966,1770,1794]" pageId="13" pageNumber="29" value="2002-06-15">15.vi.2002</date>
, R. Acosta, 1 3, 2 Ƥ; same,
<date id="121954DE163E2474FA89F91EFA42F8FE" box="[1323,1431,1770,1794]" pageId="13" pageNumber="29" value="2002-11-09">9.xi.2002</date>
, Springer, 2 3, 7 Ƥ; same, downstream,
<quantity id="A15FDFFB163E2474FDF7F8FAFD4BF8DA" box="[597,670,1806,1830]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="13" pageNumber="29" unit="m" value="325.0">325 m</quantity>
,
<date id="121954DE163E2474FD09F8FAFCF3F8DA" box="[683,806,1806,1830]" pageId="13" pageNumber="29" value="2001-08-04">4.viii.2001</date>
, Springer, 1 Ƥ;
<emphasis id="54D3AE0C163E2474FC7AF8FAFB96F8DB" bold="true" box="[984,1091,1806,1831]" pageId="13" pageNumber="29">Cartago:</emphasis>
Río Grande de Orosí, Paraíso,
<quantity id="A15FDFFB163E2474FF35F8C6FF3AF8B6" box="[151,239,1842,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="29" unit="m" value="1200.0">1200 m</quantity>
,
<date id="121954DE163E2474FF59F8C6FE50F8B6" box="[251,389,1842,1866]" pageId="13" pageNumber="29" value="2005-08-28">28.viii.2005</date>
, A.M. Glavas, 1 Ƥ; Río Tausito, antes Trapiche, Pejibaye, Jiménez,
<date id="121954DE163E2474FB2BF8C6FAD6F8B6" box="[1161,1283,1842,1866]" pageId="13" pageNumber="29" value="2005-09-18">18.ix.2005</date>
, C. Lizano, 1 Ƥ; Río Purisil, Paraíso,
<quantity id="A15FDFFB163E2474FE10F8A2FDD9F892" box="[434,524,1878,1902]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="29" unit="m" value="1200.0">1200 m</quantity>
,
<date id="121954DE163E2474FDBEF8A2FD73F892" box="[540,678,1878,1902]" pageId="13" pageNumber="29" value="2005-08-28">28.viii.2005</date>
, A.M. Glavas, 1 3;
<emphasis id="54D3AE0C163E2474FC3EF8A2FBD0F893" bold="true" box="[924,1029,1878,1903]" pageId="13" pageNumber="29">Heredia:</emphasis>
Río Poza Azul, Sarapiquí,
<quantity id="A15FDFFB163E2474FAE8F8A2FA42F892" box="[1354,1431,1878,1902]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="29" unit="m" value="275.0">275 m</quantity>
,
<date id="121954DE163E2474FF35F88EFF25F86E" box="[151,240,1914,1938]" pageId="13" pageNumber="29" value="1998-03">iii.1998</date>
, P. Paaby, 1 3; Río Sarapiquí, Sarapiquí,
<quantity id="A15FDFFB163E2474FD76F88EFCCAF86E" box="[724,799,1914,1938]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="29" unit="m" value="275.0">275 m</quantity>
,
<date id="121954DE163E2474FC8FF88EFC52F86E" box="[813,903,1914,1938]" pageId="13" pageNumber="29" value="1998-03">iii.1998</date>
, P. Paaby, 3 3, 6 Ƥ; same,
<quantity id="A15FDFFB163E2474FB6AF88EFAC1F86E" box="[1224,1300,1914,1938]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="29" unit="m" value="200.0">200 m</quantity>
,
<date id="121954DE163E2474FA80F88EFAAFF86E" box="[1314,1402,1914,1938]" pageId="13" pageNumber="29" value="1998-03">iii.1998</date>
, P. Paaby, 2 3;
<emphasis id="54D3AE0C163E2474FE82F86AFEACF84B" bold="true" box="[288,377,1950,1975]" pageId="13" pageNumber="29">Limón:</emphasis>
Río Dos Novillos, Los Manantiales,
<quantity id="A15FDFFB163E2474FCB8F86AFCB6F84A" box="[794,867,1950,1974]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="29" unit="m" value="500.0">500 m</quantity>
,
<date id="121954DE163E2474FCCDF86AFC2FF84A" box="[879,1018,1950,1974]" pageId="13" pageNumber="29" value="2005-08-20">20.viii.2005</date>
, M. Montero-Guzman, 3 3, 2 Ƥ; Río
<collectingCountry id="1EB0328E163E2474FF35F836FECCF826" box="[151,281,1986,2010]" name="Costa Rica" pageId="13" pageNumber="29">Costa Rica</collectingCountry>
, Est.
<collectionCode id="00B6EADB163E2474FEC0F836FE7EF826" box="[354,427,1986,2010]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="13" pageNumber="29">INBio</collectionCode>
, Sarapiquí,
<date id="121954DE163E2474FD99F836FD7DF826" box="[571,680,1986,2010]" pageId="13" pageNumber="29" value="2002-11-05">5.xi.2002</date>
, A.
<collectingCountry id="1EB0328E163E2474FD70F837FD3DF827" box="[722,744,1987,2011]" name="Italy" pageId="13" pageNumber="29">Y</collectingCountry>
. Jimenéz, 1 3, 2 Ƥ; Río Bris,
<quantity id="A15FDFFB163E2474FBCBF836FB6DF826" box="[1129,1208,1986,2010]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="29" unit="m" value="500.0">500 m</quantity>
, arriba de la boca,
<date id="121954DE163D2477FF35FF63FEC7FF53" box="[151,274,151,175]" pageId="14" pageNumber="30" value="2000-03-27">27.iii.2000</date>
,
<collectionCode id="00B6EADB163D2477FE83FF6CFEBFFF4C" box="[289,362,152,176]" pageId="14" pageNumber="30">ANAI</collectionCode>
, 1 3, 1 Ƥ; Río Estrella, Boca cuen,
<quantity id="A15FDFFB163D2477FCB4FF63FCB7FF53" box="[790,866,151,175]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="30" unit="m" value="200.0">200 m</quantity>
,
<date id="121954DE163D2477FCD3FF63FC3CFF53" box="[881,1001,151,175]" pageId="14" pageNumber="30" value="2000-04-29">29.iv.2000</date>
,
<collectionCode id="00B6EADB163D2477FC55FF6CFB95FF4C" box="[1015,1088,152,176]" pageId="14" pageNumber="30">ANAI</collectionCode>
, 6 3, 5 Ƥ; Río Yorkin, arriba desembocadura Río Brai,
<quantity id="A15FDFFB163D2477FE1AFF48FDD5FF28" box="[440,512,188,212]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="30" unit="m" value="500.0">500 m</quantity>
,
<date id="121954DE163D2477FDA9FF48FD52FF28" box="[523,647,188,212]" pageId="14" pageNumber="30" value="2000-03-27">27.iii.2000</date>
, V Salgado, 2 3; Río Tsui,
<quantity id="A15FDFFB163D2477FC1AFF48FBD5FF28" box="[952,1024,188,212]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="30" unit="m" value="200.0">200 m</quantity>
,
<date id="121954DE163D2477FBA9FF48FB51FF28" box="[1035,1156,188,212]" pageId="14" pageNumber="30" value="2000-04-15">15.iv.2000</date>
,
<collectionCode id="00B6EADB163D2477FB2DFF48FB0DFF28" box="[1167,1240,188,212]" pageId="14" pageNumber="30">ANAI</collectionCode>
, 1 3; Río Yorkin, Puerto Yorkin,
<quantity id="A15FDFFB163D2477FEE6FF2BFE5AFF0B" box="[324,399,223,247]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="30" unit="m" value="100.0">100 m</quantity>
,
<date id="121954DE163D2477FE3EFF2BFDC0FF0B" box="[412,533,223,247]" pageId="14" pageNumber="30" value="2000-04-16">16.iv.2000</date>
,
<collectionCode id="00B6EADB163D2477FD80FF14FDBEFF04" box="[546,619,224,248]" pageId="14" pageNumber="30">ANAI</collectionCode>
, 1 3, 2 Ƥ;
<emphasis id="54D3AE0C163D2477FD4DFF14FC54FF04" bold="true" box="[751,897,224,248]" pageId="14" pageNumber="30">Puntarenas:</emphasis>
Río Convento, secondary forest, coffee plantation, Buenos Aires,
<quantity id="A15FDFFB163D2477FEDBFEF0FE11FEE0" box="[377,452,260,284]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="30" unit="m" value="950.0">950 m</quantity>
,
<date id="121954DE163D2477FE72FEF0FD87FEE0" box="[464,594,260,284]" pageId="14" pageNumber="30" value="1998-07-23">23.vii.1998</date>
, A. Nunn, 1 3; same,
<date id="121954DE163D2477FCF6FEF0FC11FEE0" box="[852,964,260,284]" pageId="14" pageNumber="30" value="1998-05-23">23.v.1998</date>
, A. Nunn, 2 3, 5 Ƥ; Río Sonador, Longo Mai, secondary forest, coffee plantation, Buenos Aires,
<quantity id="A15FDFFB163D2477FCAAFED3FC8BFEC3" box="[776,862,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="14" pageNumber="30" unit="m" value="1050.0">1050 m</quantity>
,
<date id="121954DE163D2477FCC8FED3FC31FEC3" box="[874,996,295,319]" pageId="14" pageNumber="30" value="1998-09-24">24.ix.1998</date>
, A. Nunn, 3 3; Quebr. Naranjal, below waterfall, Golfito,
<quantity id="A15FDFFB163D2477FECEFEB8FE7CFE98" box="[364,425,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="30" unit="m" value="10.0">10 m</quantity>
,
<date id="121954DE163D2477FE15FEB8FDFCFE98" box="[439,553,332,356]" pageId="14" pageNumber="30" value="2009-04" valueMax="2009-05-31" valueMin="2009-04-00">iv–v.2009</date>
, J. Hansen, 5 3, 5 Ƥ; Quebr. Naranjal, Golfito,
<quantity id="A15FDFFB163D2477FBF5FEB8FB41FE98" box="[1111,1172,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="30" unit="m" value="10.0">10 m</quantity>
,
<date id="121954DE163D2477FB00FEB8FADBFE98" box="[1186,1294,332,356]" pageId="14" pageNumber="30" value="2000-01-21">21.i.2000</date>
, Springer &amp; UCR students, 2 3, 1 Ƥ; Quebr. Cañaza, Golfito,
<quantity id="A15FDFFB163D2477FD1CFE9BFD2DFE7B" box="[702,760,367,391]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="30" unit="m" value="10.0">10 m</quantity>
,
<date id="121954DE163D2477FCA1FE9BFC85FE7B" box="[771,848,367,391]" pageId="14" pageNumber="30" value="1999-05">v.1999</date>
, T. Hermanson &amp; J. Steffen, 1 3; Río San Luis, zona abierta, San Luis Monteverde,
<quantity id="A15FDFFB163D2477FE55FE60FD85FE50" box="[503,592,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="30" unit="m" value="1200.0">1200 m</quantity>
,
<date id="121954DE163D2477FDFFFE60FD02FE50" box="[605,727,404,428]" pageId="14" pageNumber="30" value="1995-06-23">23.vi.1995</date>
, Springer &amp; OTS students, 1 Ƥ; Quebr. La Alondra, San Luis Monteverde,
<quantity id="A15FDFFB163D2477FE8EFE43FE56FE33" box="[300,387,439,463]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="30" unit="m" value="1200.0">1200 m</quantity>
,
<date id="121954DE163D2477FE2DFE43FDDCFE33" box="[399,521,439,463]" pageId="14" pageNumber="30" value="1995-06-23">23.vi.1995</date>
, Springer &amp; OTS students, 3 3.
</paragraph>
<paragraph id="6618721E163D2477FF67FE2FFDBBFDEB" blockId="14.[151,1436,151,644]" pageId="14" pageNumber="30">
<emphasis id="54D3AE0C163D2477FF67FE2FFE3EFE08" bold="true" box="[197,491,475,500]" pageId="14" pageNumber="30">Habitat and substratum.</emphasis>
Submerged leaf litter packs and roots of riverine plants in strong current, in medium sized rivers and streams with forest cover.
</paragraph>
</subSubSection>
<subSubSection id="2EBD2195163D2477FF67FDD7FC2EFD78" pageId="14" pageNumber="30" type="diagnosis">
<paragraph id="6618721E163D2477FF67FDD7FC2EFD78" blockId="14.[151,1436,151,644]" pageId="14" pageNumber="30">
<emphasis id="54D3AE0C163D2477FF67FDD7FEEAFDC0" bold="true" box="[197,319,547,572]" pageId="14" pageNumber="30">Diagnosis.</emphasis>
<taxonomicName id="A1A7099D163D2477FEEAFDD0FDE4FDC0" box="[328,561,548,572]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="14" pageNumber="30" phylum="Arthropoda" rank="species" species="marca">
<emphasis id="54D3AE0C163D2477FEEAFDD0FDE4FDC0" box="[328,561,548,572]" italics="true" pageId="14" pageNumber="30">Anacroneuria marca</emphasis>
</taxonomicName>
can be distinguished from all other associated species by the oval pronotum that is narrower than the mesonotum (
<figureCitation id="FE9C6E9B163D2477FD9FFDB3FD45FDA3" box="[573,656,583,607]" captionStart="FIGURES 16 − 17" captionStartId="14.[151,264,1875,1898]" captionTargetBox="[247,1348,752,1784]" captionTargetId="figure@14.[151,1436,700,1855]" captionTargetPageId="14" captionText="FIGURES 16 − 17. Anacroneuria marca, 16. Head and thorax (pro, meso and meta notum); 17. Foreleg." httpUri="https://zenodo.org/record/278211/files/figure.png" pageId="14" pageNumber="30">Fig. 16</figureCitation>
); also it is the largest species described within this study and lacks the well defined line of transverse, thick bristles dorsally on the foreleg (
<figureCitation id="FE9C6E9B163D2477FC39FD98FC3BFD78" box="[923,1006,620,644]" captionStart="FIGURES 16 − 17" captionStartId="14.[151,264,1875,1898]" captionTargetBox="[247,1348,752,1784]" captionTargetId="figure@14.[151,1436,700,1855]" captionTargetPageId="14" captionText="FIGURES 16 − 17. Anacroneuria marca, 16. Head and thorax (pro, meso and meta notum); 17. Foreleg." httpUri="https://zenodo.org/record/278211/files/figure.png" pageId="14" pageNumber="30">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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