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<document id="2B1B9724A0C4B510DCF025E8BBBC7B4B" ID-CLB-Dataset="105312" ID-DOI="10.11646/zootaxa.5168.3.2" ID-GBIF-Dataset="53574cb3-f6fb-436c-bf7e-8cc59f163982" ID-ISSN="1175-5326" ID-Zenodo-Dep="6882742" ID-ZooBank="8B7C6BF7-0B3D-420F-BDAF-EB4F15E211AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658485582440" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2022" docId="E04B4F00FFE4FFCEFF4FFF3479C65280" docLanguage="en" docName="zootaxa.5168.3.2.pdf" docOrigin="Zootaxa 5168 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coleolissus deformipenis Kataev 2022, sp. n." docType="treatment" docVersion="6" lastPageNumber="294" masterDocId="1C723778FFE3FFC7FFD8FFA37C5B5029" masterDocTitle="Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China" masterLastPageNumber="305" masterPageNumber="285" pageNumber="292" updateTime="1699361575247" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73A37788EE77FC9DF0CE34C204522B36">Notes on carabid beetles of the genera Coleolissus and Siopelus (Coleoptera: Carabidae: Harpalini), with description of four new species with unusual aedeagi from India and China</mods:title>
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<mods:name id="A3F2FA53DB1A0CC45F89B15C7770F6D7" type="personal">
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<mods:namePart id="79682AAC92E46976756C3CE9602FB4C8">Kataev, Boris M.</mods:namePart>
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<mods:title id="5B339BD4BDCDF8C84D491350425AB2B0">Zootaxa</mods:title>
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<mods:date id="A41936D6369FE1B141CFBEDF0F14A424">2022</mods:date>
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<mods:number id="10921DD3FB51D6AE61AE47CFCA68184A">2022-07-22</mods:number>
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<mods:number id="B6016A0E84EB3C144E40B2AC202AACA1">5168</mods:number>
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<mods:classification id="950F16D8550CA44D9424C5A421165F35">journal article</mods:classification>
<mods:identifier id="9B20B05795B8037106A84215A61A239A" type="CLB-Dataset">105312</mods:identifier>
<mods:identifier id="FFF739AF84FAB0BE958C4731B95B85DE" type="DOI">10.11646/zootaxa.5168.3.2</mods:identifier>
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<treatment id="E04B4F00FFE4FFCEFF4FFF3479C65280" ID-DOI="http://doi.org/10.5281/zenodo.6882734" ID-GBIF-Taxon="197852160" ID-Zenodo-Dep="6882734" LSID="urn:lsid:plazi:treatment:E04B4F00FFE4FFCEFF4FFF3479C65280" httpUri="http://treatment.plazi.org/id/E04B4F00FFE4FFCEFF4FFF3479C65280" lastPageId="9" lastPageNumber="294" pageId="7" pageNumber="292">
<subSubSection id="20F8AD9DFFE4FFC0FF4FFF347E545098" box="[151,527,151,178]" pageId="7" pageNumber="292" type="nomenclature">
<paragraph id="685DFE16FFE4FFC0FF4FFF347E545098" blockId="7.[151,527,151,214]" box="[151,527,151,178]" pageId="7" pageNumber="292">
<heading id="3315497AFFE4FFC0FF4FFF347E545098" bold="true" box="[151,527,151,178]" fontSize="11" level="1" pageId="7" pageNumber="292" reason="1">
<emphasis id="5A962204FFE4FFC0FF4FFF347E545098" bold="true" box="[151,527,151,178]" pageId="7" pageNumber="292">
<taxonomicName id="AFE28595FFE4FFC0FF4FFF347D9E509B" authority="Kataev, 2022" authorityName="Kataev" authorityYear="2022" box="[151,453,151,178]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="292" phylum="Arthropoda" rank="species" species="deformipenis" status="sp. nov.">
<emphasis id="5A962204FFE4FFC0FF4FFF347D9E509B" bold="true" box="[151,453,151,178]" italics="true" pageId="7" pageNumber="292">Coleolissus deformipenis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41A59F7FFFE4FFC0FE15FF347E545098" box="[461,527,151,177]" pageId="7" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="20F8AD9DFFE4FFC0FF4FFF187D0350FF" box="[151,344,187,214]" pageId="7" pageNumber="292" type="description">
<paragraph id="685DFE16FFE4FFC0FF4FFF187D0350FF" blockId="7.[151,527,151,214]" box="[151,344,187,214]" pageId="7" pageNumber="292">
(
<figureCitation id="F0D9E293FFE4FFC0FF47FF187CA650FC" box="[159,253,187,213]" captionStart="FIGURES 9, 10" captionStartId="5.[151,264,1992,2016]" captionTargetBox="[230,1352,827,1957]" captionTargetId="figure-209@5.[223,1363,813,1968]" captionTargetPageId="5" captionText="FIGURES 9, 10. Coleolissus, habitus. 9, C. kejvali sp. n. (holotype); 10, C. deformipenis sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6882752" httpUri="https://zenodo.org/record/6882752/files/figure.png" pageId="7" pageNumber="292">Figs. 10</figureCitation>
,
<figureCitation id="F0D9E293FFE4FFC0FED1FF187D1450FF" box="[265,335,187,214]" captionStart="FIGURES 15–18" captionStartId="8.[151,264,1969,1993]" captionTargetBox="[162,1420,714,1933]" captionTargetId="figure-224@8.[151,1436,705,1945]" captionTargetPageId="8" captionText="FIGURES 15–18. Coleolissus deformipenis sp. n., male genitalia (holotype). 15, left paramere; 16, 17, median lobe of aedeagus; 18, right paramere. Right lateral (15), left lateral (16, 18) and dorsal (17) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882756" httpUri="https://zenodo.org/record/6882756/files/figure.png" pageId="7" pageNumber="292">15–18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="20F8AD9DFFE4FFC0FF4FFEA07D345168" pageId="7" pageNumber="292" type="materials_examined">
<paragraph id="685DFE16FFE4FFC0FF4FFEA07D345168" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="292">
<emphasis id="5A962204FFE4FFC0FF4FFEA07D115134" bold="true" box="[151,330,259,285]" pageId="7" pageNumber="292">Type material.</emphasis>
<materialsCitation id="D88AF44BFFE4FFC0FE8DFEA07D345168" ID-GBIF-Occurrence="3865867317" collectingDate="1997-04-12" collectingDateMax="1997-04-18" collectingDateMin="1997-04-12" collectionCode="ZIN" country="India" elevation="1740" latitude="10.15" location="Eravikulam" longLatPrecision="1298" longitude="77.066666" pageId="7" pageNumber="292" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="B75940B4FFE4FFC0FE8DFEA07D995134" box="[341,450,259,285]" pageId="7" pageNumber="292" type="holotype">Holotype</typeStatus>
: male, “
<collectingCountry id="10F5BE86FFE4FFC0FDFCFEA07E395134" box="[548,610,259,285]" name="India" pageId="7" pageNumber="292">India</collectingCountry>
,
<collectingRegion id="AA2630F4FFE4FFC0FDAAFEA07EE45134" box="[626,703,259,285]" country="India" name="Kerala" pageId="7" pageNumber="292">Kerala</collectingRegion>
,
<locationDeviation id="79EFE734FFE4FFC0FD17FEA77FC95137" box="[719,914,259,286]" location="Munnar" pageId="7" pageNumber="292">
<quantity id="AF1A53F3FFE4FFC0FD17FEA77F565134" box="[719,781,259,286]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="292" unit="km" value="7.0">7 km</quantity>
N Munnar
</locationDeviation>
,
<location id="6D3DA8CDFFE4FFC0FC7AFEA078735134" LSID="urn:lsid:plazi:treatment:E04B4F00FFE4FFCEFF4FFF3479C65280:6D3DA8CDFFE4FFC0FC7AFEA078735134" box="[930,1064,259,285]" country="India" latitude="10.15" longLatPrecision="1298" longitude="77.066666" name="Eravikulam" pageId="7" pageNumber="292" stateProvince="Kerala">Eravikulam</location>
,
<geoCoordinate id="0DD698D1FFE4FFC0FBEFFEA678CE5109" box="[1079,1173,261,288]" degrees="10" direction="north" minutes="09" orientation="latitude" pageId="7" pageNumber="292" precision="925" value="10.15">10°09′N</geoCoordinate>
<geoCoordinate id="0DD698D1FFE4FFC0FB78FEA678A05109" box="[1184,1275,261,288]" degrees="77" direction="east" minutes="04" orientation="longitude" pageId="7" pageNumber="292" precision="925" value="77.066666">77°04′E</geoCoordinate>
<quantity id="AF1A53F3FFE4FFC0FADEFEA079395137" box="[1286,1378,259,286]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="7" pageNumber="292" unit="m" value="1740.0">
<elevation id="E3CF1925FFE4FFC0FADEFEA079395137" box="[1286,1378,259,286]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="7" pageNumber="292" unit="m" value="1740.0">1740 m</elevation>
</quantity>
,
<date id="1C5CD8D6FFE4FFC0FAAAFEA07D4E5168" pageId="7" pageNumber="292" value="1997-04-12" valueMax="1997-04-18" valueMin="1997-04-12">
<collectingDate id="0C18213EFFE4FFC0FAAAFEA07D4E5168" pageId="7" pageNumber="292" value="1997-04-12" valueMax="1997-04-18" valueMin="1997-04-12">12– 18.IV.1997</collectingDate>
</date>
” (
<collectionCode id="0EF366D3FFE4FFC0FEF7FE847D385168" box="[303,355,295,321]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="7" pageNumber="292" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="20F8AD9DFFE4FFCEFF1FFEE87F255168" lastPageId="9" lastPageNumber="294" pageId="7" pageNumber="292" type="description">
<paragraph id="685DFE16FFE4FFC0FF1FFEE87CB551A0" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="292">
<emphasis id="5A962204FFE4FFC0FF1FFEE87D0A514C" bold="true" box="[199,337,331,357]" pageId="7" pageNumber="292">Description</emphasis>
(male). Body elongate, medium-sized for the genus, length
<quantity id="AF1A53F3FFE4FFC0FC20FEE8780A514F" box="[1016,1105,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="7" pageNumber="292" unit="mm" value="8.4">8.4 mm</quantity>
, width
<quantity id="AF1A53F3FFE4FFC0FB7CFEE878A5514F" box="[1188,1278,331,358]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="292" unit="mm" value="3.3">3.3 mm</quantity>
. Habitus as in
<figureCitation id="F0D9E293FFE4FFC0FF4FFECC7CB151A0" box="[151,234,367,393]" captionStart="FIGURES 9, 10" captionStartId="5.[151,264,1992,2016]" captionTargetBox="[230,1352,827,1957]" captionTargetId="figure-209@5.[223,1363,813,1968]" captionTargetPageId="5" captionText="FIGURES 9, 10. Coleolissus, habitus. 9, C. kejvali sp. n. (holotype); 10, C. deformipenis sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6882752" httpUri="https://zenodo.org/record/6882752/files/figure.png" pageId="7" pageNumber="292">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="685DFE16FFE4FFC0FF1FFE307E1C51DC" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="292">Body black, shiny and iridescent, with labrum, mandibles basally and very narrow lateral margins of pronotum reddish brown; elytral epipleura reddish black; palps, antennae and legs yellowish brown; femora slightly paler than tibiae and tarsi; coxae blackish brown.</paragraph>
<paragraph id="685DFE16FFE4FFC0FF1FFE5C78C65488" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="292">
Head medium-sized (HWmax/PWmax 0.69, HWmin/PWmax 0.53), impunctate. Eyes large, strongly convex (HWmax/HWmin 1.30),almost touching ventrally margins of buccal fissure. Genae glabrous.Tempora short, slightly convex, somewhat steeply sloping to neck. Labrum somewhat flat, very shallowly concave anteriorly. Clypeus very slightly convex mediobasally, very shallowly emarginated along anterior margin, with one setigerous pore at each outer angle and with very fine and short longitudinal wrinkles adjacent medially to each pore. Fronto-clypeal suture very fine, superficial, almost straight. Frontal foveae markedly deepened, without prolongation on clypeus. Frontoocular furrows fine, impressed at clypeus, vaguely reaching supraorbital furrows. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by distance less than width of antennomere 2 basally.
<collectingRegion id="AA2630F4FFE4FFC0FAE5FC8379C75313" box="[1341,1436,800,826]" country="Myanmar" name="Chin" pageId="7" pageNumber="292">Mentum</collectingRegion>
separated from submentum by complete transverse suture, with a prominent, rather wide median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite moderately widened distally, almost truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, narrow, markedly projecting beyond ligular sclerite and separated from it apically. Basal labial palpomere not carinate; penultimate labial palpomere about as long as apical one. Mandibles elongate, somewhat narrow, evenly curved apically, with very fine oblique wrinkles in apical half dorsally; left mandible acute, at most only slightly blunted at tip. Dorsal microsculpture visible throughout, very distinct at anterior margin of clypeus, finer on remaining surface, consisting of mixture of isodiametric and slightly transverse meshes. Antennae slender, surpassing pronotal basal edge by approximately two and a half apical antennomeres, pubescent from middle of antennomere 3, with antennomeres 4–8 about 2.5 times as long as wide and basal antennomere about as long as antennomere 3.
</paragraph>
<paragraph id="685DFE16FFE4FFC0FF1FFB0F78A356B0" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="292">Pronotum comparatively narrow (PWmax/PL 1.30), widest just before the middle, slightly more strongly narrowed apically than basally (PWmax/PWmin-ap 1.45, PWmax/PWmin-bas 1.36), with one lateral seta inserted in widest point. Sides rounded along entire length, more widely in basal half; lateral bead complete, throughout very narrow. Apical margin markedly, almost roundly emarginated, very narrowly bordered along entire length; apical border slightly obliterated at middle. Apical angles protruding anteriorly, narrowly rounded at apex. Basal margin almost straight medially, slightly rounded laterally, bordered along entire length, almost as long as apical margin (PWmin-bas/PWmin-ap 1.07) and slightly shorter than base of elytra between humeral angles; basal edge without fringe of short setae. Basal angles widely rounded. Disc convex. Lateral depressions beginning from apical angles as grooves, distinctly widened from lateral setae and fused basally with basal foveae forming flat and wide laterobasal depressions isolated from each other by convex area; basal foveae distinct, very narrow, situated at inner margins of laterobasal depressions. Median line fine, superficial, not reaching apical and basal margins. Anterior transverse depression very shallow, almost indistinct. Disc densely and finely punctate, slightly more coarsely in laterobasal depressions and with very small, indistinct punctures in central portion; punctures in laterobasal depressions confluent. Microsculpture highly obliterated, consisting of indistinct transverse meshes.</paragraph>
<paragraph id="685DFE16FFE4FFCFFF1FF9007D1550D0" blockId="7.[151,1437,259,2014]" lastBlockId="8.[151,1437,151,681]" lastPageId="8" lastPageNumber="293" pageId="7" pageNumber="292">Elytra convex, elongate oval (EL/EW 1.62, EL/PL 2.61, EW/PWmax 1.23), widest behind middle; sides rounded along entire length, very broadly medially; subapical sinuation shallow, but distinct. Humerus angularly rounded, with very small, almost indistinct denticle at apex faintly visible from behind. Sutural angle narrowly rounded at tip, not extended posteriorly. Basal edge markedly arched, forming an obtuse angle with lateral margin. Striae with granulate microsculpture on bottom, impressed along entire length, reaching anteriorly basal elytral edge, comparatively wide, widest apically. Intervals glabrous and impunctate, very strongly narrowed apically, moderately convex on disc and rather strongly convex before apex. Parascutellar (abbreviate) striole long, with a large setigerous pore basally isolated from basal elytral edge. Interval 3 with a series of 5 small discal setigerous pores adjoining stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series with very short gap at middle, consisting of 23 setigerous pores. Lateral groove flat, narrow basally, gradually widened apically, with granulate meshes. Microsculpture on intervals highly obliterated, not forming distinct meshes. Wings fully developed.</paragraph>
<paragraph id="685DFE16FFEBFFCFFF1FFEA07EEF5168" blockId="8.[151,1437,151,681]" pageId="8" pageNumber="293">Thoracic sternites punctate, with very fine and short setae on prosternum anteriorly. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</paragraph>
<paragraph id="685DFE16FFEBFFCFFF1FFEE87DC85248" blockId="8.[151,1437,151,681]" pageId="8" pageNumber="293">Metacoxa with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side distally. Protibia without longitudinal sulcus on dorsal side, with one preapical spine (in male) on outer margin. Metafemur ventrally with two long setae at posterior margin and with a few very short setae at anterior margin. Tarsi glabrous dorsally. Pro- and mesotarsomeres 1–4 (in male) markedly widened, with biseriate adhesive scales ventrally; mesotarsomere 1 slightly longer than mesotarsomere 2. Metatarsus slender, markedly longer than HWmax; metatarsomeres 1–3 only slightly, metatarsomere 4 markedly widened distally; metatarsomere 1 elongate, very slightly longer than metatarsomeres 2 and 3 combined. Tarsomere 5 with two pairs of ventro-lateral setae.</paragraph>
<paragraph id="685DFE16FFEBFFCFFF1FFDC87FCA5280" blockId="8.[151,1437,151,681]" pageId="8" pageNumber="293">Abdominal sternites glabrous, with only obligatory setae; apex of last visible sternite (VII) concave (in male), with one pair of marginal setae; these setae not distant from margin.</paragraph>
<caption id="3C9DAE9EFFEBFFCFFF4FF81278C957C5" ID-DOI="http://doi.org/10.5281/zenodo.6882756" ID-Zenodo-Dep="6882756" httpUri="https://zenodo.org/record/6882756/files/figure.png" pageId="8" pageNumber="293" startId="8.[151,264,1969,1993]" targetBox="[162,1420,714,1933]" targetPageId="8">
<paragraph id="685DFE16FFEBFFCFFF4FF81278C957C5" blockId="8.[151,1436,1969,2029]" pageId="8" pageNumber="293">
<emphasis id="5A962204FFEBFFCFFF4FF8127D0957E0" bold="true" box="[151,338,1969,1993]" pageId="8" pageNumber="293">FIGURES 15–18.</emphasis>
<taxonomicName id="AFE28595FFEBFFCFFE83F8127E0357E0" authority="Kataev, 2022" authorityName="Kataev" authorityYear="2022" box="[347,600,1969,1993]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="293" phylum="Arthropoda" rank="species" species="deformipenis" status="sp. nov.">
<emphasis id="5A962204FFEBFFCFFE83F8127E0357E0" box="[347,600,1969,1993]" italics="true" pageId="8" pageNumber="293">Coleolissus deformipenis</emphasis>
</taxonomicName>
<emphasis id="5A962204FFEBFFCFFD85F8127ECF57E0" bold="true" box="[605,660,1969,1993]" pageId="8" pageNumber="293">
<taxonomicNameLabel id="41A59F7FFFEBFFCFFD85F8127ECF57E0" box="[605,660,1969,1993]" pageId="8" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male genitalia (holotype). 15, left paramere; 16, 17, median lobe of aedeagus; 18, right paramere. Right lateral (15), left lateral (16, 18) and dorsal (17) views. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="685DFE16FFEAFFCEFF1FFF347F255168" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="294">
Aedeagus (
<figureCitation id="F0D9E293FFEAFFCEFE90FF347D8A5098" box="[328,465,150,177]" captionStart="FIGURES 15–18" captionStartId="8.[151,264,1969,1993]" captionTargetBox="[162,1420,714,1933]" captionTargetId="figure-224@8.[151,1436,705,1945]" captionTargetPageId="8" captionText="FIGURES 15–18. Coleolissus deformipenis sp. n., male genitalia (holotype). 15, left paramere; 16, 17, median lobe of aedeagus; 18, right paramere. Right lateral (15), left lateral (16, 18) and dorsal (17) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882756" httpUri="https://zenodo.org/record/6882756/files/figure.png" pageId="9" pageNumber="294">Figs. 15–18</figureCitation>
) with basal bulb and parameres comparatively large (
<figureCitation id="F0D9E293FFEAFFCEFB9CFF3478885098" box="[1092,1235,150,177]" captionStart="FIGURES 15–18" captionStartId="8.[151,264,1969,1993]" captionTargetBox="[162,1420,714,1933]" captionTargetId="figure-224@8.[151,1436,705,1945]" captionTargetPageId="8" captionText="FIGURES 15–18. Coleolissus deformipenis sp. n., male genitalia (holotype). 15, left paramere; 16, 17, median lobe of aedeagus; 18, right paramere. Right lateral (15), left lateral (16, 18) and dorsal (17) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882756" httpUri="https://zenodo.org/record/6882756/files/figure.png" pageId="9" pageNumber="294">Figs. 15, 18</figureCitation>
). Median lobe of aedeagus strongly asymmetrical, sinuously curved, in lateral view (
<figureCitation id="F0D9E293FFEAFFCEFC41FF187FB750FC" box="[921,1004,187,213]" captionStart="FIGURES 15–18" captionStartId="8.[151,264,1969,1993]" captionTargetBox="[162,1420,714,1933]" captionTargetId="figure-224@8.[151,1436,705,1945]" captionTargetPageId="8" captionText="FIGURES 15–18. Coleolissus deformipenis sp. n., male genitalia (holotype). 15, left paramere; 16, 17, median lobe of aedeagus; 18, right paramere. Right lateral (15), left lateral (16, 18) and dorsal (17) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882756" httpUri="https://zenodo.org/record/6882756/files/figure.png" pageId="9" pageNumber="294">Fig. 16</figureCitation>
) arcuate, with ventral margin slightly convex in middle portion; almost transverse apical capitulum slightly hooked dorsally; in dorsal view apical portion (
<figureCitation id="F0D9E293FFEAFFCEFF47FEA07CAA5134" box="[159,241,259,285]" captionStart="FIGURES 15–18" captionStartId="8.[151,264,1969,1993]" captionTargetBox="[162,1420,714,1933]" captionTargetId="figure-224@8.[151,1436,705,1945]" captionTargetPageId="8" captionText="FIGURES 15–18. Coleolissus deformipenis sp. n., male genitalia (holotype). 15, left paramere; 16, 17, median lobe of aedeagus; 18, right paramere. Right lateral (15), left lateral (16, 18) and dorsal (17) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6882756" httpUri="https://zenodo.org/record/6882756/files/figure.png" pageId="9" pageNumber="294">Fig. 17</figureCitation>
) sharply bent to the right; terminal lamella narrow, much longer than wide. Apical orifice vast and strongly curved along median lobe. Internal sac without sclerotic elements.
</paragraph>
</subSubSection>
<subSubSection id="20F8AD9DFFEAFFCEFF1FFEE87F9B52AC" pageId="9" pageNumber="294" type="etymology">
<paragraph id="685DFE16FFEAFFCEFF1FFEE87D8551A0" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="294">
<emphasis id="5A962204FFEAFFCEFF1FFEE87D15514C" bold="true" box="[199,334,331,357]" pageId="9" pageNumber="294">Etymology.</emphasis>
The specific name is a Latin noun at apposition, referring to the unusual (deformed) form of the median lobe of the aedeagus.
</paragraph>
<paragraph id="685DFE16FFEAFFCEFF1FFE317F9B52AC" blockId="9.[151,1437,150,681]" pageId="9" pageNumber="294">
<emphasis id="5A962204FFEAFFCEFF1FFE317D395185" bold="true" box="[199,354,402,428]" pageId="9" pageNumber="294">Comparison.</emphasis>
This new species is somewhat similar in habitus to
<taxonomicName id="AFE28595FFEAFFCEFC6CFE3078185184" authorityName="Bates" authorityYear="1878" box="[948,1091,403,429]" class="Insecta" family="Elateridae" genus="Hypolithus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="294" phylum="Arthropoda" rank="species" species="perlucens">
<emphasis id="5A962204FFEAFFCEFC6CFE3078185184" box="[948,1091,403,429]" italics="true" pageId="9" pageNumber="294">C. perlucens</emphasis>
</taxonomicName>
and
<taxonomicName id="AFE28595FFEAFFCEFBA4FE3078B25184" authority="Kataev, 2022" authorityName="Kataev" authorityYear="2022" box="[1148,1257,403,429]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="294" phylum="Arthropoda" rank="species" species="kejvali" status="sp. nov.">
<emphasis id="5A962204FFEAFFCEFBA4FE3078B25184" box="[1148,1257,403,429]" italics="true" pageId="9" pageNumber="294">C. kejvali</emphasis>
</taxonomicName>
<emphasis id="5A962204FFEAFFCEFB28FE30796A5184" bold="true" box="[1264,1329,403,429]" pageId="9" pageNumber="294">
<taxonomicNameLabel id="41A59F7FFFEAFFCEFB28FE30796A5184" box="[1264,1329,403,429]" pageId="9" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but well differs from them in having body more elongate, fronto-ocular furrows vaguely reaching supraorbital furrows, elytra with wider striae and more convex intervals, apex of last visible abdominal sternite in male concave and median lobe of aedeagus strongly asymmetrical. In addition,
<taxonomicName id="AFE28595FFEAFFCEFC88FE5C785F5230" authority="Kataev, 2022" authorityName="Kataev" authorityYear="2022" box="[848,1028,511,537]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="294" phylum="Arthropoda" rank="species" species="deformipenis" status="sp. nov.">
<emphasis id="5A962204FFEAFFCEFC88FE5C785F5230" box="[848,1028,511,537]" italics="true" pageId="9" pageNumber="294">C. deformipenis</emphasis>
</taxonomicName>
<emphasis id="5A962204FFEAFFCEFBCDFE5C780C5230" bold="true" box="[1045,1111,511,537]" pageId="9" pageNumber="294">
<taxonomicNameLabel id="41A59F7FFFEAFFCEFBCDFE5C780C5230" box="[1045,1111,511,537]" pageId="9" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="AFE28595FFEAFFCEFB37FE5C79255230" authorityName="Bates" authorityYear="1878" box="[1263,1406,511,537]" class="Insecta" family="Elateridae" genus="Hypolithus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="294" phylum="Arthropoda" rank="species" species="perlucens">
<emphasis id="5A962204FFEAFFCEFB37FE5C79255230" box="[1263,1406,511,537]" italics="true" pageId="9" pageNumber="294">C. perlucens</emphasis>
</taxonomicName>
in elytral intervals impunctate, abdominal sternites not pubescent, last visible abdominal sternite of male with one pair of setae and protibia not sulcate, and from
<taxonomicName id="AFE28595FFEAFFCEFD7EFDE47F4A5248" authority="Kataev, 2022" authorityName="Kataev" authorityYear="2022" box="[678,785,583,609]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="294" phylum="Arthropoda" rank="species" species="kejvali" status="sp. nov.">
<emphasis id="5A962204FFEAFFCEFD7EFDE47F4A5248" box="[678,785,583,609]" italics="true" pageId="9" pageNumber="294">C. kejvali</emphasis>
</taxonomicName>
<emphasis id="5A962204FFEAFFCEFCC0FDE47F0C5248" bold="true" box="[792,855,583,609]" pageId="9" pageNumber="294">
<taxonomicNameLabel id="41A59F7FFFEAFFCEFCC0FDE47F0C5248" box="[792,855,583,609]" pageId="9" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in having more distinct microsculpture on head and visible throughout, and male pro- and mesotarsi more strongly widened.
</paragraph>
</subSubSection>
<subSubSection id="20F8AD9DFFEAFFCEFF1FFD2D79C65280" box="[199,1437,654,681]" pageId="9" pageNumber="294" type="distribution">
<paragraph id="685DFE16FFEAFFCEFF1FFD2D79C65280" blockId="9.[151,1437,150,681]" box="[199,1437,654,681]" pageId="9" pageNumber="294">
<emphasis id="5A962204FFEAFFCEFF1FFD2D7D3A5281" bold="true" box="[199,353,654,680]" pageId="9" pageNumber="294">Distribution.</emphasis>
Known only from the
<typeStatus id="B75940B4FFEAFFCEFD84FD2C7ED75280" box="[604,652,655,681]" pageId="9" pageNumber="294">type</typeStatus>
locality (
<quantity id="AF1A53F3FFEAFFCEFD2DFD2C7F765280" box="[757,813,655,681]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="294" unit="km" value="7.0">7 km</quantity>
N of Munnar) in the Western Ghats,
<collectingRegion id="AA2630F4FFEAFFCEFB1EFD2C79095280" box="[1222,1362,655,681]" country="India" name="Kerala" pageId="9" pageNumber="294">Kerala State</collectingRegion>
,
<collectingCountry id="10F5BE86FFEAFFCEFA84FD2C79C25280" box="[1372,1433,655,681]" name="India" pageId="9" pageNumber="294">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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