treatments-xml/data/DF/72/B5/DF72B52DDE0CFF84FF541A7FFA12086C.xml

<document id="D5D072E04ACB4D2AA667AEFB2C7C3432" ID-CLB-Dataset="24859" ID-DOI="10.11646/zootaxa.4700.3.1" ID-GBIF-Dataset="1b0654ea-eecb-4242-ad9a-850602f7d185" ID-ISSN="1175-5326" ID-Zenodo-Dep="3548509" ID-ZooBank="B0712990-7A3B-4042-9A4B-5CE96AFAECF8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1574256400520" checkinUser="plazi" docAuthor="Grogan, William L., Díaz, Florentina, Spinelli, Gustavo R. &amp; Ronderos, Maria M." docDate="2019" docId="DF72B52DDE0CFF84FF541A7FFA12086C" docLanguage="en" docName="zootaxa.4700.3.1.pdf" docOrigin="Zootaxa 4700 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea latiala Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos 2019, n. sp." docType="treatment" docVersion="10" lastPageNumber="314" masterDocId="234BCD55DE06FF89FFC31D26FF870D63" masterDocTitle="The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer" masterLastPageNumber="325" masterPageNumber="301" pageNumber="311" updateTime="1698701345973" updateUser="plazi">
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<mods:title id="3A446E78EDEFD04F3902024BCDE82142">The Biting Midges of the Caribbean island Curaçao (Diptera: Ceratopogonidae) I. Species in the genus Dasyhelea Kieffer</mods:title>
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<mods:namePart id="24C6D86AB22B0C8D25F893F520702273">Díaz, Florentina</mods:namePart>
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<paragraph id="5764043BDE0CFF83FF541A7FFE350A10" blockId="10.[151,434,1881,1943]" box="[151,434,1881,1908]" pageId="10" pageNumber="311">
<heading id="0C2CB357DE0CFF83FF541A7FFE350A10" bold="true" box="[151,434,1881,1908]" fontSize="11" level="1" pageId="10" pageNumber="311" reason="1">
<emphasis id="65AFD829DE0CFF83FF541A7FFE350A10" bold="true" box="[151,434,1881,1908]" pageId="10" pageNumber="311">
<taxonomicName id="90DB7FB8DE0CFF83FF541A7FFEEE0A17" ID-CoL="8BQVJ" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[151,361,1881,1908]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="latiala" status="sp. nov.">
<emphasis id="65AFD829DE0CFF83FF541A7FFEEE0A17" bold="true" box="[151,361,1881,1908]" italics="true" pageId="10" pageNumber="311">Dasyhelea latiala</emphasis>
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<taxonomicNameLabel id="7E9C6552DE0CFF83FEB31A7FFE350A10" box="[368,434,1881,1907]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="5764043BDE0CFF83FF541A5BFE060AF4" blockId="10.[151,434,1881,1943]" box="[151,385,1916,1943]" pageId="10" pageNumber="311">
(
<figureCitation id="CFE018BEDE0CFF83FF5C1A5BFEA10AF4" box="[159,294,1917,1943]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="10" pageNumber="311">Figs. 11–21</figureCitation>
, 
<figureCitation id="CFE018BEDE0CFF83FEF11A5AFEFD0AF4" box="[306,378,1916,1943]" captionStart="FIGURES 52–69" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 52–69. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 52–54 D. aliciae n. sp., 55–57 D. recurva n. sp., 58–60 D. latiala n. sp., 61–63 D. cyrtostyla n. sp., 64–66 D. curacaoensis n. sp., 67–69 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="10" pageNumber="311">58–60</figureCitation>
)
</paragraph>
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<subSubSection id="1FC157B0DE0CFF82FF541AE2FDDD0A48" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311" type="diagnosis">
<paragraph id="5764043BDE0CFF82FF541AE2FAF40CCC" blockId="10.[151,1436,1988,2015]" lastBlockId="11.[151,1437,152,1835]" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311">
<emphasis id="65AFD829DE0CFF83FF541AE2FE950ABC" bold="true" box="[151,274,1988,2015]" pageId="10" pageNumber="311">Diagnosis.</emphasis>
The only Neotropical species of the mutabilis group with the following combination of characters. Males 3 
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palpal segment slender, palpal ratio 4.00; gonocoxal apodemes slightly asymmetrical; paramere, slender with bifid apex; aedeagus very complex, composed of two sections, a heavily sclerotized basal portion and a hyaline distal portion; and tergite 9 with rounded apical region that extends considerably beyond gonocoxites and elongate, stout, broadly divergent widely spaced apicolateral processes. Females with moderately long antennal flagellum, flagellomeres 2–8 moniliform to vasiform, 9–13 elongate, much longer than 2–8 and antennal ratio 0.78–0.97; palpus moderately long, segment 5 extending beyond tip of proboscis; wing very broad, length/width ratio 2.03–2.16; subgenital plate with a sub-circular basal portion with an ovoid lumen and a very broad trapezoidal apical portion; spermatheca moderately large (42.5 x 
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), ovoid, heavily sclerotized with a short, slightly recurved neck.
</paragraph>
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<emphasis id="65AFD829DE0DFF82FF041C9FFE830CB0" bold="true" box="[199,260,441,467]" pageId="11" pageNumber="312">Male</emphasis>
. Head (
<figureCitation id="CFE018BEDE0DFF82FE9D1C9FFE360CB0" box="[350,433,441,467]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 11</figureCitation>
). Dark brown. Eyes contiguous for a distance equal to the width of 2 ommatidia. Antennal flagellum with flagellomeres 2–9 rhomboid, 10–13 elongate, 13 broader than 10–12; 2–10 with single whorl of long sensillae chaetica comprising the dense plume; antennal ratio 0.97; flagellum length 
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. Frontal sclerite rhomboidal, with slender elongate ventral extension. Clypeus (
<figureCitation id="CFE018BEDE0DFF82FC8D1F03FC190F5C" box="[846,926,549,575]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 12</figureCitation>
) with 4 pairs of setae. Palpus (
<figureCitation id="CFE018BEDE0DFF82FB2C1F03FAB90F5C" box="[1263,1342,549,575]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 13</figureCitation>
) brown, elongate, apex extending considerably beyond tip of proboscis; segment 3 slender, as long as 4+5 combined, apex extending near apex of proboscis, with scattered capitate sensillae on inner surface; palpal ratio 4.00. Thorax. Scutum dark brown; scutellum pale brown with 6 large, 2 smaller setae. Wing (
<figureCitation id="CFE018BEDE0DFF82FC211FB7FBB40FC8" box="[994,1075,657,683]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 14</figureCitation>
) moderately slender; membrane hyaline with moderately dense macrotrichiae; veins brown; 2 
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radial cell very short, open apically; apices of costa, radius perpendicular with wing margin; fork of CuA 
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at level of base of 2 
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radial cell; anal lobe moderately developed; wing length 
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, width 
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; costal ratio 0.46. Abdomen. Dark brown. Genitalia (
<figureCitation id="CFE018BEDE0DFF82FB1F1FDBFABD0E74" box="[1244,1338,764,791]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Figs. 15</figureCitation>
, 
<figureCitation id="CFE018BEDE0DFF82FA851FDAFA160E74" box="[1350,1425,764,791]" captionStart="FIGURES 52–69" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 52–69. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 52–54 D. aliciae n. sp., 55–57 D. recurva n. sp., 58–60 D. latiala n. sp., 61–63 D. cyrtostyla n. sp., 64–66 D. curacaoensis n. sp., 67–69 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="11" pageNumber="312">58–60</figureCitation>
). Tergite 9 base broad, tapering progressively distally to rounded apex that extends considerably beyond gonocoxites; apicolateral processes elongate, stout, broadly divergent with subapical seta; cercus conspicuous, located at base of apicolateral process with 3–4 setae. Sternite 9 (
<figureCitation id="CFE018BEDE0DFF82FD641E4FFC830EE0" box="[679,772,872,899]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Figs. 15</figureCitation>
, 
<figureCitation id="CFE018BEDE0DFF82FCD31E4EFCAA0EE0" box="[784,813,872,899]" captionStart="FIGURES 52–69" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 52–69. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 52–54 D. aliciae n. sp., 55–57 D. recurva n. sp., 58–60 D. latiala n. sp., 61–63 D. cyrtostyla n. sp., 64–66 D. curacaoensis n. sp., 67–69 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="11" pageNumber="312">58</figureCitation>
) 0.45 length of width; base broadly curved; posteromedian portion broad, tapering slightly distally with notched apex. Gonocoxite stout, 1.7 x longer than broad, distal margin straight, apicomesal surface with group of dense, moderately long setae; gonostylus nearly as long as gonocoxite, proximal 1/2 broad, tapering abruptly at mid-length, slightly curved with pointed tip. Paramere and gonocoxal apodemes (
<figureCitation id="CFE018BEDE0DFF82FE9D1EDFFE3A0970" box="[350,445,1016,1043]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Figs. 15</figureCitation>
, 
<figureCitation id="CFE018BEDE0DFF82FE081EDEFE6F0970" box="[459,488,1016,1043]" captionStart="FIGURES 52–69" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 52–69. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 52–54 D. aliciae n. sp., 55–57 D. recurva n. sp., 58–60 D. latiala n. sp., 61–63 D. cyrtostyla n. sp., 64–66 D. curacaoensis n. sp., 67–69 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="11" pageNumber="312">59</figureCitation>
) nearly symmetrical; gonocoxal apodemes slender on proximal 1/2, distal portions broader with small basal spur, apices truncate; paramere slender, nearly straight, apex bifurcate.Aedeagus (
<figureCitation id="CFE018BEDE0DFF82FAF8193BFA100954" box="[1339,1431,1052,1079]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Figs. 15</figureCitation>
, 
<figureCitation id="CFE018BEDE0DFF82FF541967FF33093F" box="[151,180,1089,1116]" captionStart="FIGURES 52–69" captionStartId="20.[151,264,1944,1969]" captionTargetBox="[187,1405,192,1908]" captionTargetId="figure@20.[170,1417,181,1919]" captionTargetPageId="20" captionText="FIGURES 52–69. Male Dasyhelea genitalia structures: sternites 9 (left column), gonocoxal apodemes and parameres (middle column), and aedeagi (right column). 52–54 D. aliciae n. sp., 55–57 D. recurva n. sp., 58–60 D. latiala n. sp., 61–63 D. cyrtostyla n. sp., 64–66 D. curacaoensis n. sp., 67–69 D. rhopaloparamera n. sp. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548521" httpUri="https://zenodo.org/record/3548521/files/figure.png" pageId="11" pageNumber="312">60</figureCitation>
) 0.60 length of greatest width, very complex, composed of two sections; basal arch stout, slightly curved, extending nearly 0.50 of total length, distal margin notched; basal arm stout, anterolaterally directed with mid-posterior outer tooth; posterior projection short, stout, with slightly expanded rounded tip; distal portion covered by a hyaline envelope that reaches tips of posteromedian projection and borne on pair of slender, heavily sclerotized processes directed anterolaterally, contiguous to inner margin of basal arms.
</paragraph>
<paragraph id="5764043BDE0DFF82FF0419D3FDDD0A48" blockId="11.[151,1437,152,1835]" pageId="11" pageNumber="312">
<emphasis id="65AFD829DE0DFF82FF0419D3FE99086C" bold="true" box="[199,286,1269,1295]" pageId="11" pageNumber="312">Female</emphasis>
. Head (
<figureCitation id="CFE018BEDE0DFF82FEB519D3FE4E0873" box="[374,457,1269,1296]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 16</figureCitation>
). Dark brown. Eyes contiguous for a distance equal to the width of 2–3 ommatidia. Antennal flagellum with flagellomeres 2–8 moniliform to vasiform, 9–13 longer than 2–8, 13 longest; antennal ratio 0.78–0.97 (0.87, n=2); flagellum length 0.37–0.40 (n=2) mm. Frontal sclerite rhomboidal, with slender elongate ventral extension. Clypeus (
<figureCitation id="CFE018BEDE0DFF82FE161847FDAF0818" box="[469,552,1377,1403]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 17</figureCitation>
) with 7 pairs of setae; anterior margin concave. Palpus (
<figureCitation id="CFE018BEDE0DFF82FB6E1847FA870818" box="[1197,1280,1377,1403]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 18</figureCitation>
) light brown; segment 3 nearly as long as 4+5 combined, with scattered capitate sensillae on mesal surface; palpal ratio 3.66 (n=2). Thorax (
<figureCitation id="CFE018BEDE0DFF82FE86188FFE1E08A0" box="[325,409,1449,1475]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 19</figureCitation>
). Scutum dark brown, paler on humeral angles and prescutellar depressions; scutellum yellowish, with 6 large, 2 smaller setae. Legs light brown, tarsomeres 5 more darkly infuscated; hind tibial comb with 6 spines; prothoracic tarsal ratio 2.14–2.25 (2.19, n=2), mesothoracic tarsal ratio 2.00, metathoracic tarsal ratio 2.18–2.22 (2.20, n=2). Wing (
<figureCitation id="CFE018BEDE0DFF82FE211B33FDB40B4C" box="[482,563,1557,1583]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 20</figureCitation>
) very broad, length/width ratio 2.03–2.16; membrane hyaline with dense macrotrichiae; veins brown; 2 
<superScript id="A0AEA973DE0DFF82FE7A1B1EFE4E0B25" attach="left" box="[441,457,1592,1606]" fontSize="6" pageId="11" pageNumber="312">nd</superScript>
radial cell very short, open distally; apices of costa, radius at slight oblique angle with wing margin; fork of CuA 
<subScript id="CB5F067EDE0DFF82FE7C1B48FE400B1F" attach="left" box="[447,455,1646,1660]" fontSize="6" pageId="11" pageNumber="312">1</subScript>
just anterior to base of 2 
<superScript id="A0AEA973DE0DFF82FD231B7AFD770B09" attach="left" box="[736,752,1628,1642]" fontSize="6" pageId="11" pageNumber="312">nd</superScript>
radial cell; anal lobe well developed; wing length 0.65–0.80 (0.72, n=2) mm, width 0.32–0.37 (0.35, n=2) mm; costal ratio 0.50 (n=2). Halter yellowish. Abdomen (
<figureCitation id="CFE018BEDE0DFF82FAFA1BA7FA080BF8" box="[1337,1423,1665,1691]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 21</figureCitation>
). Dark brown. Subgenital plate (
<figureCitation id="CFE018BEDE0DFF82FE2C1B83FDC70BDC" box="[495,576,1701,1727]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 21</figureCitation>
) with a sub-circular basal portion with an ovoid lumen and broad trapezoidal apical portion; posteromedian margin rounded with a small bifurcate posteromedian projection; posterolateral arms slender, recurved 30˚ at mid-length. Spermatheca (
<figureCitation id="CFE018BEDE0DFF82FD0F1BCBFC9A0A64" box="[716,797,1773,1799]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="11" pageNumber="312">Fig. 21</figureCitation>
) small (42.5 x 
<quantity id="9023A9DEDE0DFF82FC061BCAFBA20A64" box="[965,1061,1772,1799]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="11" pageNumber="312" unit="mm" value="37.5">37.5 µm</quantity>
), ovoid, heavily sclerotized; neck short, slightly recurved, length 
<quantity id="9023A9DEDE0DFF82FE361A36FDD10A48" box="[501,598,1808,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="312" unit="mm" value="12.5">12.5 µm</quantity>
.
</paragraph>
</subSubSection>
<caption id="03A454B3DE0AFF85FF541A1AFB4D0A1B" ID-DOI="http://doi.org/10.5281/zenodo.3548513" ID-Zenodo-Dep="3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="12" pageNumber="313" startId="12.[151,264,1852,1877]" targetBox="[174,1407,195,1793]" targetPageId="12">
<paragraph id="5764043BDE0AFF85FF541A1AFB4D0A1B" blockId="12.[151,1436,1852,1913]" pageId="12" pageNumber="313">
<emphasis id="65AFD829DE0AFF85FF541A1AFECA0A36" bold="true" box="[151,333,1852,1877]" pageId="12" pageNumber="313">FIGURES 11–21</emphasis>
. 
<taxonomicName id="90DB7FB8DE0AFF85FE9A1A1BFD8A0A36" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[345,525,1853,1877]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="latiala" status="sp. nov.">
<emphasis id="65AFD829DE0AFF85FE9A1A1BFD8A0A36" box="[345,525,1853,1877]" italics="true" pageId="12" pageNumber="313">Dasyhelea latiala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0AFF85FDD71A1BFDCA0A37" box="[532,589,1853,1877]" pageId="12" pageNumber="313" rank="species">
<emphasis id="65AFD829DE0AFF85FDD71A1BFDA50A36" bold="true" box="[532,546,1853,1877]" pageId="12" pageNumber="313">n</emphasis>
. 
<emphasis id="65AFD829DE0AFF85FDEC1A1BFDC00A36" bold="true" box="[559,583,1853,1877]" pageId="12" pageNumber="313">sp</emphasis>
.
</taxonomicNameLabel>
Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="1FC157B0DE0BFF84FF041DBEFE4B0DD0" box="[199,460,152,179]" pageId="13" pageNumber="314" type="distribution">
<paragraph id="5764043BDE0BFF84FF041DBEFE4B0DD0" blockId="13.[151,1437,152,1295]" box="[199,460,152,179]" pageId="13" pageNumber="314">
<emphasis id="65AFD829DE0BFF84FF041DBEFEDE0DD0" bold="true" box="[199,345,152,179]" pageId="13" pageNumber="314">Distribution</emphasis>
. 
<collectingCountry id="2FCC44ABDE0BFF84FEA41DBFFE4F0DD0" box="[359,456,153,179]" name="Curaçao" pageId="13" pageNumber="314">Curaçao</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0BFF84FF041D9BFBE40C04" pageId="13" pageNumber="314" type="materials_examined">
<paragraph id="5764043BDE0BFF84FF041D9BFB600C7C" blockId="13.[151,1437,152,1295]" pageId="13" pageNumber="314">
<emphasis id="65AFD829DE0BFF84FF041D9BFEF60DB4" bold="true" box="[199,369,188,215]" pageId="13" pageNumber="314">Type Material</emphasis>
. 
<materialsCitation id="E7B30E66DE0BFF84FEBC1D9BFB600C7C" ID-GBIF-Occurrence="2454903105" collectingDate="2014-11-06" collectingDateMax="2015-12-05" collectingDateMin="2014-11-06" collectionCode="FSCA, MLPA" collectorName="M. C. Thomas" country="Curaçao" latitude="12.126911" location="Cambi" longLatPrecision="1" longitude="-69.96699" pageId="13" pageNumber="314" specimenCount="4" typeStatus="holotype">
<typeStatus id="8860BA99DE0BFF84FEBC1D9BFE600DB4" box="[383,487,189,215]" pageId="13" pageNumber="314" type="holotype">Holotype</typeStatus>
male, labeled “ 
<collectingCountry id="2FCC44ABDE0BFF84FD561D9BFD7F0DB4" box="[661,760,189,215]" name="Curaçao" pageId="13" pageNumber="314">Curaçao</collectingCountry>
, Piscadera Baai, 
<date id="236522FBDE0BFF84FC741D9AFBBE0DB4" box="[951,1081,188,215]" pageId="13" pageNumber="314" value="2015-12-05">
<collectingDate id="3321DB13DE0BFF84FC741D9AFBBE0DB4" box="[951,1081,188,215]" pageId="13" pageNumber="314" value="2015-12-05">5-XII-2015</collectingDate>
</date>
, 
<collectorName id="FA2E61EDDE0BFF84FB861D9BFB6E0DB4" box="[1093,1257,189,215]" pageId="13" pageNumber="314">M. C. Thomas</collectorName>
, 
<collectingMethod id="8E9A7C2CDE0BFF84FB361D9BFA1A0DB4" box="[1269,1437,189,215]" pageId="13" pageNumber="314">Blacklight trap</collectingMethod>
in mangroves”; 
<typeStatus id="8860BA99DE0BFF84FE881DC7FE220D98" box="[331,421,225,251]" pageId="13" pageNumber="314" type="allotype">allotype</typeStatus>
female with same data except 
<quantity id="9023A9DEDE0BFF84FCC11DC6FCD70D98" box="[770,848,224,251]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="314" unit="km" value="0.5">0.5 km</quantity>
N of 
<location id="520452E0DE0BFF84FC511DC7FC670D98" LSID="urn:lsid:plazi:treatment:DF72B52DDE0CFF84FF541A7FFA12086C:520452E0DE0BFF84FC511DC7FC670D98" box="[914,992,225,251]" country="Curaçao" latitude="12.126911" longLatPrecision="1" longitude="-69.96699" name="Cambi" pageId="13" pageNumber="314">Cambi</location>
, 
<geoCoordinate id="32EF62FCDE0BFF84FC281DC6FB190D98" box="[1003,1182,224,251]" degrees="12" direction="north" minutes="07" orientation="latitude" pageId="13" pageNumber="314" precision="1" seconds="36.88" value="12.126911">12°07ʹ36.88ʹʹ N</geoCoordinate>
, 
<geoCoordinate id="32EF62FCDE0BFF84FB691DC6FAD40D98" box="[1194,1363,224,251]" degrees="69" direction="west" minutes="58" orientation="longitude" pageId="13" pageNumber="314" precision="1" seconds="1.14" value="-69.96699">69°58ʹ1.14ʹʹ W</geoCoordinate>
, 
<date id="236522FBDE0BFF84FA9D1DC7FF480C7C" pageId="13" pageNumber="314" value="2014-11-06">
<collectingDate id="3321DB13DE0BFF84FA9D1DC7FF480C7C" pageId="13" pageNumber="314" value="2014-11-06">6-XI- 2014</collectingDate>
</date>
(deposited in 
<collectionCode id="31CA9CFEDE0BFF84FEAC1C23FE3D0C7C" box="[367,442,261,287]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="13" pageNumber="314" type="Museum">FSCA</collectionCode>
). One female 
<typeStatus id="8860BA99DE0BFF84FD961C23FD320C7C" box="[597,693,261,287]" pageId="13" pageNumber="314" type="paratype">paratype</typeStatus>
with same data as 
<typeStatus id="8860BA99DE0BFF84FC491C23FC6B0C7C" box="[906,1004,261,287]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
(deposited in 
<collectionCode id="31CA9CFEDE0BFF84FB481C23FB5B0C7C" box="[1163,1244,261,287]" pageId="13" pageNumber="314">MLPA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5764043BDE0BFF84FF041C0EFBE40C04" blockId="13.[151,1437,152,1295]" pageId="13" pageNumber="314">
<emphasis id="65AFD829DE0BFF84FF041C0EFDA60C20" bold="true" box="[199,545,296,323]" pageId="13" pageNumber="314">Derivation of specific epithet</emphasis>
. The specific epithet was derived from two Latin terms, 
<emphasis id="65AFD829DE0BFF84FB641C0FFB4B0C20" box="[1191,1228,297,323]" italics="true" pageId="13" pageNumber="314">lati</emphasis>
(broad, wide) and 
<emphasis id="65AFD829DE0BFF84FF541C6BFF3C0C04" box="[151,187,333,359]" italics="true" pageId="13" pageNumber="314">ala</emphasis>
(wing), a reference to the very broad wing (
<figureCitation id="CFE018BEDE0BFF84FD691C6BFD7C0C04" box="[682,763,333,359]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="13" pageNumber="314">Fig. 20</figureCitation>
) of females of this new species.
</paragraph>
</subSubSection>
<subSubSection id="1FC157B0DE0BFF84FF041C56FA12086C" pageId="13" pageNumber="314" type="discussion">
<paragraph id="5764043BDE0BFF84FF041C56FF530EC4" blockId="13.[151,1437,152,1295]" pageId="13" pageNumber="314">
<emphasis id="65AFD829DE0BFF84FF041C56FEC20CE8" bold="true" box="[199,325,368,395]" pageId="13" pageNumber="314">Discussion</emphasis>
. The paramere of the 
<typeStatus id="8860BA99DE0BFF84FD871C57FD210CE8" box="[580,678,369,395]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="90DB7FB8DE0BFF84FCCD1C54FCFC0CE8" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[782,891,369,395]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala" status="sp. nov.">
<emphasis id="65AFD829DE0BFF84FCCD1C54FCA50CE8" box="[782,802,370,395]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FCF11C57FCFC0CE8" box="[818,891,369,395]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0BFF84FC401C57FC460CE8" box="[899,961,369,395]" pageId="13" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
is similar to males of 
<taxonomicName id="90DB7FB8DE0BFF84FB071C54FF690CCC" authority="Diaz &amp; Spinelli" authorityName="Diaz &amp; Spinelli" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="yamana">
<emphasis id="65AFD829DE0BFF84FB071C54FB5F0CE8" box="[1220,1240,370,395]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FB2B1C54FAC60CE8" box="[1256,1345,370,395]" italics="true" pageId="13" pageNumber="314">yamana</emphasis>
Díaz &amp; Spinelli
</taxonomicName>
(
<emphasis id="65AFD829DE0BFF84FEC21CB0FE900CCC" box="[257,279,406,431]" italics="true" pageId="13" pageNumber="314">in</emphasis>
<bibRefCitation id="334A79CADE0BFF84FEDC1CB3FE5B0CCC" author="Diaz, F. &amp; Spinelli, G. R. &amp; Donato, M. &amp; Ronderos, M. M." box="[287,476,405,431]" pageId="13" pageNumber="314" pagination="2117 - 2175" refId="ref19101" refString="Diaz, F., Spinelli, G. R., Donato, M. &amp; Ronderos, M. M. (2014) A taxonomic revision of the Patagonian species of the Dasyhelea mutabilis group with a phylogenetic analysis of New World species (Diptera: Ceratopogonidae). Journal of Natural History, 48, 2117 - 2175. https: // doi. org / 10.1080 / 00222933.2014.909062" type="journal article" year="2014">
Díaz 
<emphasis id="65AFD829DE0BFF84FE9E1CB0FE170CCC" box="[349,400,405,431]" italics="true" pageId="13" pageNumber="314">et al</emphasis>
. 2014
</bibRefCitation>
: 
<figureCitation id="CFE018BEDE0BFF84FE2A1CB3FDCB0CCC" box="[489,588,405,431]" captionStart="FIGURES 11–21" captionStartId="12.[151,264,1852,1877]" captionTargetBox="[174,1407,195,1793]" captionTargetId="figure@12.[164,1423,181,1827]" captionTargetPageId="12" captionText="FIGURES 11–21. Dasyhelea latiala n. sp. Holotype male, 11–15, allotype female, 16–21. 11, 16 head. 12, 17 clypeus. 13, 18 palpus. 14, 20 wing. 15 genitalia. 19 thorax. 21 apex of abdomen and spermatheca. Scale bars = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3548513" httpUri="https://zenodo.org/record/3548513/files/figure.png" pageId="13" pageNumber="314">Fig. 17E</figureCitation>
) from 
<collectingRegion id="951FCAD9DE0BFF84FD5F1CB3FC400CCC" box="[668,967,405,431]" country="Argentina" name="Tierra del Fuego - Antartida e Islas del Atlantico Sur" pageId="13" pageNumber="314">Tierra del Fuego province</collectingRegion>
, 
<collectingCountry id="2FCC44ABDE0BFF84FC101CB3FBC50CCC" box="[979,1090,405,431]" name="Argentina" pageId="13" pageNumber="314">Argentina</collectingCountry>
which have a slightly shorter, narrower paramere with a more deeply bifid apex. In addition, the gonocoxal apodemes of 
<taxonomicName id="90DB7FB8DE0BFF84FB781C9CFABE0CB0" box="[1211,1337,441,467]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="yamana">
<emphasis id="65AFD829DE0BFF84FB781C9CFB480CB0" box="[1211,1231,442,467]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FB231C9CFABE0CB0" box="[1248,1337,442,467]" italics="true" pageId="13" pageNumber="314">yamana</emphasis>
</taxonomicName>
are also symmetrical with a short basolateral spur that is similar to the more elongate basolateral spurs on the larger, nearly symmetrical gonocoxal apodemes of 
<taxonomicName id="90DB7FB8DE0BFF84FDF51F24FD270F78" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[566,672,513,539]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FDF51F24FDCD0F78" box="[566,586,514,539]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FD941F27FD270F78" box="[599,672,513,539]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
. However, males of 
<taxonomicName id="90DB7FB8DE0BFF84FC471F24FC790F78" box="[900,1022,513,539]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="yamana">
<emphasis id="65AFD829DE0BFF84FC471F24FC1F0F78" box="[900,920,514,539]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FC661F24FC790F78" box="[933,1022,514,539]" italics="true" pageId="13" pageNumber="314">yamana</emphasis>
</taxonomicName>
differ from the 
<typeStatus id="8860BA99DE0BFF84FB6E1F27FA880F78" box="[1197,1295,513,539]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="90DB7FB8DE0BFF84FAF11F24FA1B0F78" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[1330,1436,513,539]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FAF11F24FAC10F78" box="[1330,1350,514,539]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FA901F27FA1B0F78" box="[1363,1436,513,539]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
in having a much longer wing (wing length 
<quantity id="9023A9DEDE0BFF84FD481F03FCA80F5C" box="[651,815,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="1.185" metricValueMax="1.23" metricValueMin="1.14" pageId="13" pageNumber="314" unit="mm" value="1.185" valueMax="1.23" valueMin="1.14">1.14–1.23 mm</quantity>
vs. 
<quantity id="9023A9DEDE0BFF84FC9C1F03FC430F5C" box="[863,964,549,575]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="314" unit="mm" value="0.8">0.80 mm</quantity>
in 
<taxonomicName id="90DB7FB8DE0BFF84FC2A1F00FBD20F5C" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[1001,1109,549,575]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FC2A1F00FC7A0F5C" box="[1001,1021,550,575]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FBCF1F03FBD20F5C" box="[1036,1109,549,575]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
) that is not as narrow with a more poorly developed anal lobe; a more typical aedeagus with a large, stout posteromedian projection, a broad nearly straight basal arch and slender posterolateral arms with recurved tips; sternite 9 has a longer, more narrow distal extension; stouter gonostyli; and tergite 9 has shorter, less widely divergent apicolateral processes. Females of 
<taxonomicName id="90DB7FB8DE0BFF84FF761F90FEB70FAC" box="[181,304,693,719]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="yamana">
<emphasis id="65AFD829DE0BFF84FF761F90FF4E0FAC" box="[181,201,694,719]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FF141F90FEB70FAC" box="[215,304,694,719]" italics="true" pageId="13" pageNumber="314">yamana</emphasis>
</taxonomicName>
differ from females of 
<taxonomicName id="90DB7FB8DE0BFF84FDF01F90FD1A0FAC" authority="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos, 2019" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[563,669,693,719]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala" status="sp. nov.">
<emphasis id="65AFD829DE0BFF84FDF01F90FDC00FAC" box="[563,583,694,719]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FD971F93FD1A0FAC" box="[596,669,693,719]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7E9C6552DE0BFF84FD671F93FD580FAC" box="[676,735,693,719]" pageId="13" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
in having a shorter antennal flagellum with only flagellomeres 9–13 slightly longer than broad; a more elongate 3 
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palpal segment with a larger palpal ratio (4.00–4.80 vs. 
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<taxonomicName id="90DB7FB8DE0BFF84FF541FD8FE840E74" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[151,259,765,791]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FF541FD8FF2C0E74" box="[151,171,766,791]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FF791FDBFE840E74" box="[186,259,765,791]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
); a longer wing (wing length 
<quantity id="9023A9DEDE0BFF84FD951FDBFD7E0E74" box="[598,761,765,791]" metricMagnitude="-3" metricUnit="m" metricValue="1.005" metricValueMax="1.08" metricValueMin="0.93" pageId="13" pageNumber="314" unit="mm" value="1.005" valueMax="1.08" valueMin="0.93">0.93–1.08 mm</quantity>
vs. 
<quantity id="9023A9DEDE0BFF84FCEB1FDAFC4B0E74" box="[808,972,764,791]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="13" pageNumber="314" unit="mm" value="0.725" valueMax="0.8" valueMin="0.65">0.65–0.80 mm</quantity>
in 
<taxonomicName id="90DB7FB8DE0BFF84FC331FD8FBDB0E74" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[1008,1116,765,791]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FC331FD8FB830E74" box="[1008,1028,766,791]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FBD01FDBFBDB0E74" box="[1043,1116,765,791]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
) with a shorter costa (costal ratio 0.41–0.45 vs. 
<quantity id="9023A9DEDE0BFF84FEAE1E06FE3D0E58" box="[365,442,800,827]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="13" pageNumber="314" unit="in" value="0.5">0.50 in</quantity>
<taxonomicName id="90DB7FB8DE0BFF84FE7C1E04FDAD0E58" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[447,554,801,827]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FE7C1E04FE540E58" box="[447,467,802,827]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FE221E07FDAD0E58" box="[481,554,801,827]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
) with a more poorly developed anal lobe; a broader frontal sclerite in which the dorsal portion is fused with the narrower ventral portion; a more typical semi-circular subgenital plate but without a broad trapezoidal anterior portion and posteromedian projection; and a spherical spermatheca with a short straight neck.
</paragraph>
<paragraph id="5764043BDE0BFF84FF041E97FCDF091C" blockId="13.[151,1437,152,1295]" pageId="13" pageNumber="314">
Males of 
<taxonomicName id="90DB7FB8DE0BFF84FEF01E94FD4A0EAF" authority="Wirth &amp; Waugh (1976)" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[307,717,945,972]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="65AFD829DE0BFF84FEF01E94FEC00EA8" box="[307,327,946,971]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FE941E97FE390EA8" box="[343,446,945,971]" italics="true" pageId="13" pageNumber="314">williamsi</emphasis>
Wirth &amp; 
<bibRefCitation id="334A79CADE0BFF84FDE81E97FD4A0EAF" author="Waugh, W. T. &amp; Wirth, W. W." box="[555,717,945,972]" pageId="13" pageNumber="314" pagination="219 - 247" refId="ref20962" refString="Waugh, W. T. &amp; Wirth, W. W. (1976) A revision of the genus Dasyhelea Kieffer of the eastern United States north of Florida (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 69, 219 - 247. https: // doi. org / 10.1093 / aesa / 69.2.219" type="journal article" year="1976">Waugh (1976)</bibRefCitation>
</taxonomicName>
have a very short broader paramere but the distal 2/3 is deeply bifid and their apices are broad and widely separated; the shape of the aedeagus is much different with long, stout lateral arms that are widely spaced basally, curved distally with broad nearly convergent tips; the distal 1/2 of the gonostyli are much broader; tergite 9 is much shorter with a straight margin and very short apicolateral processes; and sternite 9 is more elongate and extends just below the apex of the aedeagus. In addition, the thorax and much of the abdomen of both sexes of 
<taxonomicName id="90DB7FB8DE0BFF84FE241940FDF7091C" authorityName="Wirth &amp; Waugh" authorityYear="1976" box="[487,624,1125,1151]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="65AFD829DE0BFF84FE241940FE7C091C" box="[487,507,1126,1151]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FDCA1943FDF7091C" box="[521,624,1125,1151]" italics="true" pageId="13" pageNumber="314">williamsi</emphasis>
</taxonomicName>
is mostly yellowish.
</paragraph>
<paragraph id="5764043BDE0BFF84FF0419AFFA12086C" blockId="13.[151,1437,152,1295]" pageId="13" pageNumber="314">
Finally, the very unusual, greatly complex aedeagus of the 
<typeStatus id="8860BA99DE0BFF84FCA419AFFC4E09C0" box="[871,969,1161,1187]" pageId="13" pageNumber="314" type="holotype">holotype</typeStatus>
male of 
<taxonomicName id="90DB7FB8DE0BFF84FBF319ACFB1B09C0" authorityName="Grogan &amp; Díaz &amp; Spinelli &amp; Ronderos" authorityYear="2019" box="[1072,1180,1161,1187]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="latiala">
<emphasis id="65AFD829DE0BFF84FBF319ACFBC309C0" box="[1072,1092,1162,1187]" italics="true" pageId="13" pageNumber="314">D</emphasis>
. 
<emphasis id="65AFD829DE0BFF84FB9019AFFB1B09C0" box="[1107,1180,1161,1187]" italics="true" pageId="13" pageNumber="314">latiala</emphasis>
</taxonomicName>
does not resemble the aedeagi of other males in the mutabilis group including those in the fasciigera complex. Accordingly, it is possible that we have not correctly associated both sexes of this species. However, we are fairly confident of their association as all three specimens in the type series were collected in the same region and their overall colorations are similar.
</paragraph>
</subSubSection>
</treatment>
</document>